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<mods:title id="B1C6CEF881DFA014975881DE2C6AEFD7">Taxonomic study of the genus Eloeophila Rondani, 1856 of Japan (Limoniidae Diptera)</mods:title>
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<paragraph id="8BDD368F121CFA12FF645DB0FE1EFE13" blockId="1.[151,472,438,465]" box="[151,472,438,465]" pageId="1" pageNumber="426">
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<taxonomicName id="4C624D0C121CFA12FF645DB0FE1EFE13" ID-CoL="87H3T" authority="Rondani, 1856" authorityName="Rondani" authorityYear="1856" box="[151,472,438,465]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D121CFA12FF645DB0FE1EFE13" bold="true" box="[151,472,438,465]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF645DB0FEDDFE12" bold="true" box="[151,283,438,465]" italics="true" pageId="1" pageNumber="426">Eloeophila</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FED05DB0FE1EFE13" author="Rondani, C." box="[291,472,438,464]" pageId="1" pageNumber="426" refId="ref19091" refString="Rondani, C. (1856) Dipterologiae Italicae Prodromus. Vol. 1. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. Ex Tipographia Alexandri Stocche, Parmae [Parma], 228 pp. https: // doi. org / 10.5962 / bhl. title. 8160" type="book" year="1856">Rondani, 1856</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BDD368F121CFA12FF645DFDFE1AFDD0" blockId="1.[151,1437,507,716]" box="[151,476,507,531]" pageId="1" pageNumber="426">
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<treatmentCitation id="0AC3109E121CFA12FF645DFDFE1EFDD0" author="Rondani, C." box="[151,472,507,531]" page="182" pageId="1" pageNumber="426" year="1856">
<taxonomicName id="4C624D0C121CFA12FF645DFDFE1EFDD0" ID-CoL="87H3T" authority="Rondani, 1856: 182" authorityName="Rondani" authorityPageNumber="182" authorityYear="1856" box="[151,472,507,531]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D121CFA12FF645DFDFEC0FDD0" box="[151,262,507,531]" italics="true" pageId="1" pageNumber="426">Eloeophila</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FEFE5DFDFE1EFDD0" author="Rondani, C." box="[269,472,507,531]" pageId="1" pageNumber="426" refId="ref19091" refString="Rondani, C. (1856) Dipterologiae Italicae Prodromus. Vol. 1. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. Ex Tipographia Alexandri Stocche, Parmae [Parma], 228 pp. https: // doi. org / 10.5962 / bhl. title. 8160" type="book" year="1856">Rondani, 1856: 182</bibRefCitation>
</taxonomicName>
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.
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<subSubSection id="C3786504121CFA12FF645E1CFDFEFBB8" pageId="1" pageNumber="426" type="materials_examined">
<paragraph id="8BDD368F121CFA12FF645E1CFB6FFDF1" blockId="1.[151,1437,507,716]" box="[151,1193,538,562]" pageId="1" pageNumber="426">
<typeStatus id="54D9882D121CFA12FF645E1CFF0FFDF1" box="[151,201,538,562]" pageId="1" pageNumber="426">Type</typeStatus>
species:
<taxonomicName id="4C624D0C121CFA12FED55E1CFD55FDF1" authority="Meigen, 1818" authorityName="Meigen" authorityYear="1818" box="[294,659,538,562]" class="Insecta" family="Limoniidae" genus="Limnobia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis id="B916EA9D121CFA12FED55E1CFDC6FDF1" box="[294,512,538,562]" italics="true" pageId="1" pageNumber="426">Limnobia marmorata</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FDF55E1CFD55FDF1" author="Meigen, J. W." box="[518,659,538,562]" pageId="1" pageNumber="426" refId="ref18745" refString="Meigen, J. W. (1818) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Vol. 1. Bei Friedrich Wilhelm Forstmann, Aachen, xxxvi + 324 pp. https: // doi. org / 10.5962 / bhl. title. 13731" type="book" year="1818">Meigen, 1818</bibRefCitation>
</taxonomicName>
[=
<emphasis id="B916EA9D121CFA12FD455E1CFCD2FDF1" box="[694,788,538,562]" italics="true" pageId="1" pageNumber="426">maculata</emphasis>
(
<bibRefCitation id="EFF34B7E121CFA12FCD15E1CFC75FDF1" author="Meigen, J. W." box="[802,947,538,562]" pageId="1" pageNumber="426" refId="ref18704" refString="Meigen, J. W. (1804) Klassifikazion und Beschreibung der europaischen zweiflugeligen Insekten (Diptera Linn.). Erster Band. Abt. I. Reichard, Braunschweig [Brunswick], xxviii + 152 pp." type="book" year="1804">Meigen, 1804</bibRefCitation>
)] (original designation).
</paragraph>
<paragraph id="8BDD368F121CFA12FF645E3EFE75FD92" blockId="1.[151,1437,507,716]" box="[151,435,568,593]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF645E3EFF20FD93" box="[151,230,568,592]" italics="true" pageId="1" pageNumber="426">Ephelia</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FF1F5E3EFE68FD92" author="Schiner, J. R." box="[236,430,568,593]" pageId="1" pageNumber="426" pagination="217 - 226" refId="ref19237" refString="Schiner, J. R. (1863) Vorlaufiger Commentar zum dipterologischen Theile der Fauna austriaca. V [concl.]. Wiener Entomologische Monatschrift, 7, 217 - 226." type="journal article" year="1863">Schiner, 1863: 222</bibRefCitation>
.
</paragraph>
<paragraph id="8BDD368F121CFA12FF645E5EFE73FD4D" blockId="1.[151,1437,507,716]" pageId="1" pageNumber="426">
<typeStatus id="54D9882D121CFA12FF645E5EFF0FFDB3" box="[151,201,600,624]" pageId="1" pageNumber="426">Type</typeStatus>
species:
<taxonomicName id="4C624D0C121CFA12FEDE5E51FD62FDAC" authority="Meigen, 1818" authorityName="Meigen" authorityYear="1818" box="[301,676,599,623]" class="Insecta" family="Limoniidae" genus="Limnobia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis id="B916EA9D121CFA12FEDE5E51FDCCFDAC" box="[301,522,599,623]" italics="true" pageId="1" pageNumber="426">Limnobia marmorata</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FDE05E51FD62FDAC" author="Meigen, J. W." box="[531,676,599,623]" pageId="1" pageNumber="426" refId="ref18745" refString="Meigen, J. W. (1818) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Vol. 1. Bei Friedrich Wilhelm Forstmann, Aachen, xxxvi + 324 pp. https: // doi. org / 10.5962 / bhl. title. 13731" type="book" year="1818">Meigen, 1818</bibRefCitation>
</taxonomicName>
[=
<emphasis id="B916EA9D121CFA12FD3E5E51FCEDFDAC" box="[717,811,599,623]" italics="true" pageId="1" pageNumber="426">maculata</emphasis>
(
<bibRefCitation id="EFF34B7E121CFA12FCCF5E51FC17FDAC" author="Meigen, J. W." box="[828,977,599,623]" pageId="1" pageNumber="426" refId="ref18704" refString="Meigen, J. W. (1804) Klassifikazion und Beschreibung der europaischen zweiflugeligen Insekten (Diptera Linn.). Erster Band. Abt. I. Reichard, Braunschweig [Brunswick], xxviii + 152 pp." type="book" year="1804">Meigen, 1804</bibRefCitation>
)] (original designation). Synonymic note in
<bibRefCitation id="EFF34B7E121CFA12FF345E70FE77FD4D" author="Alexander, C. P." box="[199,433,630,654]" pageId="1" pageNumber="426" pagination="105 - 136" refId="ref18048" refString="Alexander, C. P. (1932) New or little-known Tipulidae from eastern Asia (Diptera). X. Philippine Journal of Science, 49, 105 - 136." type="journal article" year="1932">Alexander (1932: 122)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDD368F121CFA12FF645E92FC69FD6E" blockId="1.[151,1437,507,716]" box="[151,943,660,685]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF645E92FECCFD6F" box="[151,266,660,684]" italics="true" pageId="1" pageNumber="426">Trichehelia</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FEE35E92FE27FD6E" author="Alexander, C. P." box="[272,481,660,685]" pageId="1" pageNumber="426" pagination="71 - 78" refId="ref18182" refString="Alexander, C. P. (1938) New or insufficiently-known crane-flies from the Nearctic region (Tipulidae, Diptera). Part IV. Bulletin of the Brooklyn Entomological Society, 33, 71 - 78." type="journal article" year="1938">Alexander, 1938: 73</bibRefCitation>
. As subgenus of
<taxonomicName id="4C624D0C121CFA12FD7F5E92FC6DFD6E" authority="Macquart, 1834" authorityName="Macquart" authorityYear="1834" box="[652,939,660,685]" class="Insecta" family="Limoniidae" genus="Limnophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D121CFA12FD7F5E92FCC7FD6F" box="[652,769,660,684]" italics="true" pageId="1" pageNumber="426">Limnophila</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FCF45E93FC6DFD6E" author="Macquart, J." box="[775,939,661,685]" pageId="1" pageNumber="426" refId="ref18658" refString="Macquart, J. (1834) Histoire naturelle des Insectes. Dipteres. Ouvrage accompagne de planches. Tome premier. Librairie Encyclopedique de Roret, Paris, 578 pp. https: // doi. org / 10.5962 / bhl. title. 14274" type="book" year="1834">Macquart, 1834</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDD368F121CFA12FF645EB2FA90FD08" blockId="1.[151,1437,507,716]" box="[151,1366,691,716]" pageId="1" pageNumber="426">
<typeStatus id="54D9882D121CFA12FF645EB2FF0FFD0F" box="[151,201,692,716]" pageId="1" pageNumber="426">Type</typeStatus>
species:
<taxonomicName id="4C624D0C121CFA12FED55EB5FD71FD08" authority="Alexander, 1938" authorityName="Alexander" authorityYear="1938" box="[294,695,691,715]" class="Insecta" family="Limoniidae" genus="Limnophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="species" species="seticellula">
<emphasis id="B916EA9D121CFA12FED55EB5FDCFFD08" box="[294,521,691,715]" italics="true" pageId="1" pageNumber="426">Limnophila seticellula</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FDFC5EB5FD71FD08" author="Alexander, C. P." box="[527,695,691,715]" pageId="1" pageNumber="426" pagination="71 - 78" refId="ref18182" refString="Alexander, C. P. (1938) New or insufficiently-known crane-flies from the Nearctic region (Tipulidae, Diptera). Part IV. Bulletin of the Brooklyn Entomological Society, 33, 71 - 78." type="journal article" year="1938">Alexander, 1938</bibRefCitation>
</taxonomicName>
(original designation). Synonymic note in
<bibRefCitation id="EFF34B7E121CFA12FB9B5EB5FA95FD08" author="Alexander, C. P." box="[1128,1363,691,715]" pageId="1" pageNumber="426" pagination="349 - 388" refId="ref18400" refString="Alexander, C. P. (1957) Records and descriptions of Japanese Tipulidae (Diptera). Part V. The crane-flies of Honshu. I. Philippine Journal of Science, 85, 349 - 388." type="journal article" year="1957">Alexander (1957: 363)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDD368F121CFA12FF645EFFFB3DFC9F" blockId="1.[151,1437,761,1364]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF645EFFFD59FCD0" bold="true" box="[151,671,761,788]" pageId="1" pageNumber="426">
General description of Japanese
<taxonomicName id="4C624D0C121CFA12FDE95EFFFD5EFCD7" authorityName="Rondani" authorityYear="1856" box="[538,664,761,788]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D121CFA12FDE95EFFFD5EFCD7" bold="true" box="[538,664,761,788]" italics="true" pageId="1" pageNumber="426">Eloeophila</emphasis>
</taxonomicName>
.
</emphasis>
<emphasis id="B916EA9D121CFA12FD565EFFFD24FCD0" box="[677,738,761,787]" italics="true" pageId="1" pageNumber="426">Head</emphasis>
. Eye dichoptic; anterior part of vertex about 1/3 width of head. Rostrum about half length of eye in lateral view. Antenna 16-segmented; scape cylindrical, about twice as long as pedicel; pedicel globular; flagellomeres covered with pubescence and verticils. Palpus 5-segmented.
</paragraph>
<paragraph id="8BDD368F121CFA12FF345F63FAE6FC04" blockId="1.[151,1437,761,1364]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF345F63FEDEFCBC" box="[199,280,869,895]" italics="true" pageId="1" pageNumber="426">Thorax</emphasis>
. Prescutum bearing distinct prescutal pit. Tuberculate pit absent. Pleuron with meron small; mid and hind coxae close to each other. Wing often sparsely or heavily spotted. M
<subScript id="17E634CA121CFA12FC2E5F9AFC30FC69" attach="left" box="[989,1014,924,938]" fontSize="6" pageId="1" pageNumber="426">1+2</subScript>
forked. Cell bm with supernumerary m-cu near middle. Legs with tibial spurs 1 + 2 + 2. Halter usually 0.7 times as long as thorax or shorter.
</paragraph>
<paragraph id="8BDD368F121CFA12FF345FD7FDFEFBB8" blockId="1.[151,1437,761,1364]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF345FD7FEFFFC28" box="[199,313,977,1003]" italics="true" pageId="1" pageNumber="426">Abdomen.</emphasis>
Male terminalia with tergite 9 not separated from sternite 9. Gonocoxite with outer and inner gonostylus. Outer gonostylus sclerotized, usually flattened dorsoventrally. Inner gonostylus freshly. Parameres extending medially, fused with each other at center, forming bridge. Mesal-apical lobe of paramere bare at tip. Lateral apodeme of lower shelf about as long as lower shelf. Ovipositor (
<figureCitation id="13592A0A121CFA12FC8E583BFC13FB94" box="[893,981,1085,1111]" captionStart="FIGURE 1" captionStartId="2.[151,250,1599,1623]" captionTargetBox="[172,1415,201,1547]" captionTargetId="figure-16@2.[151,1435,179,1574]" captionTargetPageId="2" captionText="FIGURE 1. Eloeophila amamiana (Alexander, 1956). A. habitus, lateral view; B. wing; C. male terminalia, dorsal view; D. outer gonostylus, ventral view; E. aedeagal complex, dorsal view; F. same, lateral view (left = dorsal); G. paramere, dorsal surface; H. female terminalia, lateral view. Scale bars: AB, 2 mm; CH, 0.1 mm. Abbreviations: ad, aedeagus; av, apodeme of vesica; cr, cercus; hv, hypogynial valve; ig, inner gonostylus; lp, lateral apodeme; ls, lower shelf; outer gonostylus; pm, paramere; s m-cu, supernumerary m-cu; sX, sternite X; tX, tergite X; vs, vesica." figureDoi="http://doi.org/10.5281/zenodo.5057237" httpUri="https://zenodo.org/record/5057237/files/figure.png" pageId="1" pageNumber="426">Fig. 1H</figureCitation>
) with cercus longer than tergite 10. Hypogynial valve longer than sternite 8.
</paragraph>
</subSubSection>
<subSubSection id="C3786504121CFA12FF345883FBB5FA97" pageId="1" pageNumber="426" type="discussion">
<paragraph id="8BDD368F121CFA12FF345883FBB5FA97" blockId="1.[151,1437,761,1364]" pageId="1" pageNumber="426">
<emphasis id="B916EA9D121CFA12FF345883FEFDFB5C" bold="true" box="[199,315,1157,1183]" pageId="1" pageNumber="426">Remarks.</emphasis>
<taxonomicName id="4C624D0C121CFA12FEB75883FE79FB5C" authorityName="Rondani" authorityYear="1856" box="[324,447,1157,1183]" class="Insecta" family="Limoniidae" genus="Eloeophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D121CFA12FEB75883FE79FB5C" box="[324,447,1157,1183]" italics="true" pageId="1" pageNumber="426">Eloeophila</emphasis>
</taxonomicName>
is similar to
<emphasis id="B916EA9D121CFA12FDA85883FD05FB5C" box="[603,707,1157,1183]" italics="true" pageId="1" pageNumber="426">Idioptera</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FD3E5880FC47FB5C" author="Macquart, J." box="[717,897,1157,1184]" pageId="1" pageNumber="426" refId="ref18658" refString="Macquart, J. (1834) Histoire naturelle des Insectes. Dipteres. Ouvrage accompagne de planches. Tome premier. Librairie Encyclopedique de Roret, Paris, 578 pp. https: // doi. org / 10.5962 / bhl. title. 14274" type="book" year="1834">
Macquart,
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in terms of having a supernumerary m-cu, but differenciated from the genus by the following characters: outer gonostylus of male terminalia heavily sclerotized, dark, usually flattened dorsoventrally, tip usually pointed (not heavily sclerotized, almost as pale as inner gonostylus, not flattened dorsoventrally, obtuse at tip in
<emphasis id="B916EA9D121CFA12FD4758F7FCDAFAC8" box="[692,796,1265,1291]" italics="true" pageId="1" pageNumber="426">Idioptera</emphasis>
). In general appearance and structure of male terminalia, this genus resembles
<taxonomicName id="4C624D0C121CFA12FE765913FD38FAF3" authority="Alexander, 1956 b" authorityName="Alexander" authorityYear="1956" box="[389,766,1301,1328]" class="Insecta" family="Limoniidae" genus="Afrolimnophila" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA9D121CFA12FE765913FDE9FAEC" box="[389,559,1301,1327]" italics="true" pageId="1" pageNumber="426">Afrolimnophila</emphasis>
<bibRefCitation id="EFF34B7E121CFA12FDC75913FD38FAF3" author="Alexander, C. P." box="[564,766,1301,1328]" pageId="1" pageNumber="426" pagination="129 - 380" refId="ref18376" refString="Alexander, C. P. (1956 b) Tipulidae. Ruwenzori Expedition 1934 - 35, 1, 129 - 380." type="journal article" year="1956">Alexander, 1956b</bibRefCitation>
</taxonomicName>
, but species of the latter genus do not have a supernumerary m-cu. (rarely the crossvein present, but differing in the structure of the male terminalia).
</paragraph>
</subSubSection>
</treatment>
</document>