121 lines
20 KiB
XML
121 lines
20 KiB
XML
<document id="6E009789F82382D796A23BC8BECF6F4A" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. & Hughes, Nigel C." docDate="2022" docId="03CB87846E721D2FFF3AC49E4BC4F7E8" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calymene oldhami Brongniart 1822" docType="treatment" docVersion="7" lastPageNumber="340" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="340" updateTime="1699356965331" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="37F351FF8457A97C1FBD2FF08D16D7A7">Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:namePart id="F1945A5EBA8BE1DF285FB1D2A4E626B6">Fortey, Richard A.</mods:namePart>
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<mods:affiliation id="E8CBEA8D1EF99855B9FF7FE9AD9FD7B6">Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="982DF525B724D26317D50459528696A5">Wernette, Shelly J.</mods:namePart>
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<mods:affiliation id="CE9A9CED2CCD158CFE7D87772AA8437C">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. & Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<mods:namePart id="50CE7F09BD32F528FAF5F0F0599D607F">Hughes, Nigel C.</mods:namePart>
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<mods:title id="9C26B726B2B571BB8153C0AB2346269C">Zootaxa</mods:title>
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<mods:date id="1CB1BC13AC2EECADF121B29675B84E5A">2022</mods:date>
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<mods:number id="8A2FB25C612CDCFDA2E98D86AE362EDA">2022-07-08</mods:number>
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<treatment id="03CB87846E721D2FFF3AC49E4BC4F7E8" ID-DOI="http://doi.org/10.5281/zenodo.6816307" ID-GBIF-Taxon="197214830" ID-Zenodo-Dep="6816307" LSID="urn:lsid:plazi:treatment:03CB87846E721D2FFF3AC49E4BC4F7E8" httpUri="http://treatment.plazi.org/id/03CB87846E721D2FFF3AC49E4BC4F7E8" lastPageNumber="340" pageId="39" pageNumber="340">
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<subSubSection id="C37865196E721D2FFF3AC49E4833F5B5" box="[151,552,223,250]" pageId="39" pageNumber="340" type="nomenclature">
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<paragraph id="8BDD36926E721D2FFF3AC49E4833F5B5" blockId="39.[151,552,223,285]" box="[151,552,223,250]" pageId="39" pageNumber="340">
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<heading id="D09581FE6E721D2FFF3AC49E4833F5B5" bold="true" box="[151,552,223,250]" fontSize="11" level="1" pageId="39" pageNumber="340" reason="1">
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<emphasis id="B916EA806E721D2FFF3AC49E4833F5B5" bold="true" box="[151,552,223,250]" pageId="39" pageNumber="340">
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<emphasis id="B916EA806E721D2FFF3AC49E4ABEF5B6" bold="true" box="[151,165,223,250]" italics="true" pageId="39" pageNumber="340">“</emphasis>
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<taxonomicName id="4C624D116E721D2FFF08C49E4833F5B5" ID-CoL="3G5C" authority="Reed, 1915" authorityName="Brongniart" authorityYear="1822" box="[165,552,223,250]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="340" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E721D2FFF08C49E4B8CF5B6" bold="true" box="[165,407,223,250]" italics="true" pageId="39" pageNumber="340">Calymene” oldhami</emphasis>
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<bibRefCitation id="EFF34B636E721D2FFE32C49E4833F5B5" author="Reed, F. R. C." box="[415,552,223,249]" pageId="39" pageNumber="340" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C37865196E721D2FFF3AC5424947F42C" pageId="39" pageNumber="340" type="description">
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<paragraph id="8BDD36926E721D2FFF3AC5424B2BF451" blockId="39.[151,552,223,285]" box="[151,304,259,285]" pageId="39" pageNumber="340">
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<figureCitation id="13592A176E721D2FFF3AC5424B14F451" box="[151,271,259,285]" captionStart="FIGURE 10" captionStartId="31.[151,250,1456,1480]" captionTargetBox="[287,1301,181,1432]" captionTargetId="figure-18@31.[287,1301,181,1432]" captionTargetPageId="31" captionText="FIGURE 10. (1, 2) Lonchodamas? sp. (Reed) from the Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. 1a, b. Pygidium (original of Reed 1915, pl. 5, fig. 6), GSI 11508, original and retrodeformation. 2. Cranidium (refigured from Reed 1915, pl. 5, fig. 5), GSI 11507. (3, 6) Lonchodamas shanensis (Reed) from the Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 3. Pygidium (original of Reed 1915, pl. 5, fig. 3), GSI 11505, Hwe-hok. 6. Cranidium (original of Reed 1915, pl. 5, fig. 2), GSI 11504, Mong Ha. (4) Dionide hybrida (Reed), Cephalon (original of Reed 1915, pl. 5, fig. 7), GSI 11509, Hwe Mawng Beds (uppermost Katian), Hwe-hok, northern Shan State, Myanmar. (5, 8) Ovalocephalus? sp. indet. (Reed) from the Li-Lu Formation (Katian), Mân-ngai, northern part of Shan State, Myanmar. 5. Cranidium (original of Reed 1915, pl. 8, fig. 17), GSI 11563. 8. Thorax (original of Reed 1915, pl. 8, fig. 21), GSI 11567. (7) Pliomerina martellii (Reed), Cranidium (original of Reed 1917, pl. 8, fig. 15), GSI 11915, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (9) Prionocheilus liluensis (Reed), Cranidium (original of Reed 1915, pl. 8 figs 6 and 7), GSI 11554, Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. (10, 13) Ovalocephalus sp. (Reed) from the Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. 10. Pygidium (original of Reed 1915, pl. 8, fig. 9), GSI 11556. 13. Cephalon; not previously figured; no GSI catalogue number. (11, 12) “Calymene” oldhami (Reed) from Lower Naungkangyi Beds (probable Darriwilian), Mâ Shio, northern Shan State, Myanmar. 11. Cephalon and thorax (original of Reed 1915, pl. 8, fig. 11), GSI 11558. 12. Cephalon and partial thorax (original of Reed 1915, pl. 8, fig. 10), GSI 11557. Scale bars = 2 mm for 1–3, 6, 8–13; = 5 mm for 4, 5; = 1 mm for 7." figureDoi="http://doi.org/10.5281/zenodo.6810316" httpUri="https://zenodo.org/record/6810316/files/figure.png" pageId="39" pageNumber="340">Figs 10.11</figureCitation>
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,
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<figureCitation id="13592A176E721D2FFEB9C5424B2BF451" box="[276,304,259,285]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="39" pageNumber="340">12</figureCitation>
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</paragraph>
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<paragraph id="8BDD36926E721D2FFF3AC5064947F42C" blockId="39.[151,860,327,352]" box="[151,860,327,352]" pageId="39" pageNumber="340">
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1915
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<taxonomicName id="4C624D116E721D2FFF5AC5094BADF42C" authority="Fortey & Wernette & Hughes, 2022" authorityName="Fortey & Wernette & Hughes" authorityYear="2022" box="[247,438,328,352]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="oldhami" status="sp. nov.">
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<emphasis id="B916EA806E721D2FFF5AC5094BADF42C" box="[247,438,328,352]" italics="true" pageId="39" pageNumber="340">Calymene oldhami</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22557FB6E721D2FFE11C5094810F413" box="[444,523,328,351]" pageId="39" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
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Reed; p. 47–48, pl. 8, figs 10,11.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E721D2FFF3AC5CC4F2BF481" pageId="39" pageNumber="340" type="materials_examined">
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<paragraph id="8BDD36926E721D2FFF3AC5CC4F2BF481" blockId="39.[151,1437,397,676]" pageId="39" pageNumber="340">
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<emphasis id="B916EA806E721D2FFF3AC5CC4B1DF4EB" bold="true" box="[151,262,397,423]" pageId="39" pageNumber="340">Material.</emphasis>
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<materialsCitation id="3B0A3CCF6E721D2FFEA1C5CF4F2BF481" ID-GBIF-Occurrence="3861306319" accessionNumber="GSI 11557" collectionCode="GSI" location="Lower Naungkangyi Beds" municipality="Man-shio" pageId="39" pageNumber="340" specimenCode="11558" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D988306E721D2FFEA1C5CF4B62F4E4" box="[268,377,398,424]" pageId="39" pageNumber="340" type="holotype">Holotype</typeStatus>
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: cranidium with partial thorax and hypostome in part and counterpart from
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<location id="8EBD60496E721D2FFB1AC5CF4AD4F480" LSID="urn:lsid:plazi:treatment:03CB87846E721D2FFF3AC49E4BC4F7E8:8EBD60496E721D2FFB1AC5CF4AD4F480" municipality="Man-shio" name="Lower Naungkangyi Beds" pageId="39" pageNumber="340">Lower Naungkangyi Beds</location>
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(probable Darriwilian) at
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<collectingMunicipality id="6BB9ACE86E721D2FFE58C5F34878F480" box="[501,611,434,460]" pageId="39" pageNumber="340">Man-shio</collectingMunicipality>
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,
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<figureCitation id="13592A176E721D2FFDC3C5F348FCF480" box="[622,743,434,460]" captionStart="FIGURE 10" captionStartId="31.[151,250,1456,1480]" captionTargetBox="[287,1301,181,1432]" captionTargetId="figure-18@31.[287,1301,181,1432]" captionTargetPageId="31" captionText="FIGURE 10. (1, 2) Lonchodamas? sp. (Reed) from the Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. 1a, b. Pygidium (original of Reed 1915, pl. 5, fig. 6), GSI 11508, original and retrodeformation. 2. Cranidium (refigured from Reed 1915, pl. 5, fig. 5), GSI 11507. (3, 6) Lonchodamas shanensis (Reed) from the Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 3. Pygidium (original of Reed 1915, pl. 5, fig. 3), GSI 11505, Hwe-hok. 6. Cranidium (original of Reed 1915, pl. 5, fig. 2), GSI 11504, Mong Ha. (4) Dionide hybrida (Reed), Cephalon (original of Reed 1915, pl. 5, fig. 7), GSI 11509, Hwe Mawng Beds (uppermost Katian), Hwe-hok, northern Shan State, Myanmar. (5, 8) Ovalocephalus? sp. indet. (Reed) from the Li-Lu Formation (Katian), Mân-ngai, northern part of Shan State, Myanmar. 5. Cranidium (original of Reed 1915, pl. 8, fig. 17), GSI 11563. 8. Thorax (original of Reed 1915, pl. 8, fig. 21), GSI 11567. (7) Pliomerina martellii (Reed), Cranidium (original of Reed 1917, pl. 8, fig. 15), GSI 11915, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (9) Prionocheilus liluensis (Reed), Cranidium (original of Reed 1915, pl. 8 figs 6 and 7), GSI 11554, Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. (10, 13) Ovalocephalus sp. (Reed) from the Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. 10. Pygidium (original of Reed 1915, pl. 8, fig. 9), GSI 11556. 13. Cephalon; not previously figured; no GSI catalogue number. (11, 12) “Calymene” oldhami (Reed) from Lower Naungkangyi Beds (probable Darriwilian), Mâ Shio, northern Shan State, Myanmar. 11. Cephalon and thorax (original of Reed 1915, pl. 8, fig. 11), GSI 11558. 12. Cephalon and partial thorax (original of Reed 1915, pl. 8, fig. 10), GSI 11557. Scale bars = 2 mm for 1–3, 6, 8–13; = 5 mm for 4, 5; = 1 mm for 7." figureDoi="http://doi.org/10.5281/zenodo.6810316" httpUri="https://zenodo.org/record/6810316/files/figure.png" pageId="39" pageNumber="340">Figs 10.11</figureCitation>
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,
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<figureCitation id="13592A176E721D2FFD46C5F3491CF480" box="[747,775,434,460]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="39" pageNumber="340">12</figureCitation>
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(
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<bibRefCitation id="EFF34B636E721D2FFCB8C5F34982F481" author="Reed, F. R. C." box="[789,921,434,461]" pageId="39" pageNumber="340" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 8 figs 11,10),
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<collectionCode id="ED73AE576E721D2FFBC9C5F34E8AF480" box="[1124,1169,434,460]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="39" pageNumber="340">GSI</collectionCode>
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11557,
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<specimenCode id="DBC49EE96E721D2FFB49C5F34F37F481" box="[1252,1324,434,461]" collectionCode="GSI" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="39" pageNumber="340">11558</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37865196E721D2FFF6AC5944BC4F7E8" pageId="39" pageNumber="340" type="discussion">
|
||
<paragraph id="8BDD36926E721D2FFF6AC5944BC4F7E8" blockId="39.[151,1437,397,676]" pageId="39" pageNumber="340">
|
||
<emphasis id="B916EA806E721D2FFF6AC5944B57F4A3" bold="true" box="[199,332,469,495]" pageId="39" pageNumber="340">Discussion.</emphasis>
|
||
A single small cranidium was the basis for this species.
|
||
<bibRefCitation id="EFF34B636E721D2FFC11C5974E5DF4BD" author="Reed, F. R. C." box="[956,1094,470,497]" pageId="39" pageNumber="340" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915)</bibRefCitation>
|
||
illustrated both part and counterpart of this specimen, and this is followed herein. However, the specimens have deteriorated since their original description. For example, the hypostome is quite clearly shown on Brock’s illustration (
|
||
<bibRefCitation id="EFF34B636E721D2FFBDAC65F4EE7F775" author="Reed, F. R. C." box="[1143,1276,542,569]" pageId="39" pageNumber="340" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
|
||
, pl. 8, fig. 11) and displays the median fork consistent with calymenid affinities; this is no longer clearly shown. They are included here for completeness. It is not possible to give a more precise generic assignment to this species, which is best regarded as a
|
||
<emphasis id="B916EA806E721D2FFE9FC6CA4BC3F7E8" box="[306,472,650,676]" italics="true" pageId="39" pageNumber="340">nomen dubium</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |