treatments-xml/data/03/CB/87/03CB8781FFE4FFE32E77F9E52A78FABD.xml
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<document id="54EC7F5029AE2BB52E09A498B2C38DED" ID-DOI="10.5281/zenodo.158781" ID-GBIF-Dataset="22fe8498-8c9e-43b5-a8e8-193af943e51d" ID-ISSN="1175­5326" ID-Zenodo-Dep="158781" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461031685441" checkinUser="plazi" docAuthor="Betancur-R, Ricardo &amp; P, Arturo Acero" docDate="2004" docId="03CB8781FFE4FFE32E77F9E52A78FABD" docLanguage="en" docName="zt00703.pdf" docOrigin="Zootaxa 703" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Notarius" docType="key" docVersion="8" lastPageNumber="18" masterDocId="FFF2FFF9FFF4FFF22F7FFFE52F53FFA0" masterDocTitle="tus (Jordan and Gilbert) (Siluriformes: Ariidae) from the eastern Pacific, with evidence of monophyly and limits of Notarius" masterLastPageNumber="20" masterPageNumber="1" pageNumber="17" updateTime="1698573480147" updateUser="plazi">
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<mods:title id="FAAB229E046D5A1C2F144EC05A372B87">tus (Jordan and Gilbert) (Siluriformes: Ariidae) from the eastern Pacific, with evidence of monophyly and limits of Notarius</mods:title>
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<mods:namePart id="5F8F6DEEA6E8C99644317C3DE62EEFD8">Betancur-R, Ricardo</mods:namePart>
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<mods:namePart id="96FDCE85233716EB4EA33E3C1B953C6D">P, Arturo Acero</mods:namePart>
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<mods:date id="BFABAF9111A67F5B446BC629B2372BD4">2004</mods:date>
<mods:detail id="0F6F14B940462D5E7F80385732E9C075" type="volume">
<mods:number id="E1654A54AB599FFA9C457B831B3EF0AD">703</mods:number>
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<mods:identifier id="A050696BC9847F06A141B51A597CB893" type="Zenodo-Dep">158781</mods:identifier>
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<treatment id="03CB8781FFE4FFE32E77F9E52A78FABD" ID-DOI="http://doi.org/10.5281/zenodo.6273392" ID-GBIF-Taxon="119608183" ID-Zenodo-Dep="6273392" LSID="urn:lsid:plazi:treatment:03CB8781FFE4FFE32E77F9E52A78FABD" httpUri="http://treatment.plazi.org/id/03CB8781FFE4FFE32E77F9E52A78FABD" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C378651CFFE4FFE22E77F9E52D2BF8F4" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BDD3697FFE4FFE22E77F9E52B24F9BA" blockId="16.[264,1143,1536,1562]" box="[264,1143,1536,1562]" pageId="16" pageNumber="17">
<heading id="D09581FBFFE4FFE22E77F9E52B24F9BA" bold="true" box="[264,1143,1536,1562]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B916EA85FFE4FFE22E77F9E52B24F9BA" bold="true" box="[264,1143,1536,1562]" pageId="16" pageNumber="17">
Key to described species of the genus
<taxonomicName id="4C624D14FFE4FFE22DA3F9E52C16F9BA" ID-CoL="64Z5" ID-ENA="286639" box="[732,837,1536,1562]" class="Actinopterygii" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B916EA85FFE4FFE22DA3F9E52C16F9BA" bold="true" box="[732,837,1536,1562]" italics="true" pageId="16" pageNumber="17">Notarius</emphasis>
</taxonomicName>
from the eastern Pacific
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD3697FFE4FFE22E77F9AF2D2BF8F4" blockId="16.[264,1323,1610,1876]" pageId="16" pageNumber="17">
The species of the genus
<taxonomicName id="4C624D14FFE4FFE22D53F9AE2DC0F9C4" box="[556,659,1611,1636]" class="Actinopterygii" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B916EA85FFE4FFE22D53F9AE2DC0F9C4" box="[556,659,1611,1636]" italics="true" pageId="16" pageNumber="17">Notarius</emphasis>
</taxonomicName>
are distinguished from other EP ariid taxa by the following combination of features: humeral process pointed, triangular to elongated, but never fan­shaped; three pairs of barbels present; fleshy furrow between posterior nostrils absent; fleshy groove in median depression of head absent; coarse to sharp granules or spinulations on anterior surface of head shield absent; gill rakers on rear surfaces of first two gill arches absent. Some of the data ranges showed below are based on wider ranges proposed by
<bibRefCitation id="EFF34B66FFE4FFE22E52F8DF2D27F8F4" author="Kailola" box="[301,628,1850,1876]" pageId="16" pageNumber="20" refString="Kailola, P. J. &amp; Bussing, W. A. (1995) Ariidae. In: Fischer, W., Krupp, F., Schneider, W., Sommer, C., Carpenter, K. E. &amp; Niem, V. (Eds.). Guia FAO para identificacion de especies para los fines de la pesca, Pacifico Centro-Oriental. Vol 2, FAO, Rome, pp. 860 - 886." type="journal article" year="1995">Kailola and Bussing (1995)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C378651CFFE5FFE32E77FEF92A78FABD" pageId="17" pageNumber="18" type="key">
<keyStep id="30962E32FFE5FFE32E77FEF92A7FFE26" pageId="17" pageNumber="18">
<paragraph id="8BDD3697FFE5FFE32E77FEF92A78FEFD" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FEF92A78FEFD" pageId="17" pageNumber="18">
1 Predorsal plate large, square or hexagonal and shaped like a forward pointing arrow .. ....................................................................................................................
<taxonomicName id="4C624D14FFE5FFE32BE4FEA12A78FEFD" box="[1179,1323,324,349]" class="Insecta" family="Cerambycidae" genus="Sciades" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="troschelii">
<emphasis id="B916EA85FFE5FFE32BE4FEA12A78FEFD" box="[1179,1323,324,349]" italics="true" pageId="17" pageNumber="18">N. troschelii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD3697FFE5FFE32E77FE892A7FFE26" blockId="17.[264,1324,284,1310]" box="[264,1324,364,390]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FE892A7FFE26" box="[264,1324,364,390]" pageId="17" pageNumber="18">­ Predorsal plate narrow and crescent­shaped ................................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E32FFE5FFE32E77FE712A78FE76" pageId="17" pageNumber="18">
<paragraph id="8BDD3697FFE5FFE32E77FE712A78FE0D" blockId="17.[264,1324,284,1310]" box="[264,1323,404,430]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FE712A78FE0D" box="[264,1323,404,430]" pageId="17" pageNumber="18">
2 Gill rakers on second arch 56; anal fin rays 2328 ..............................
<taxonomicName id="4C624D14FFE5FFE32B04FE712A78FE0D" box="[1147,1323,404,429]" family="Ariidae" genus="Tachisurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="lentiginosus">
<emphasis id="B916EA85FFE5FFE32B04FE712A78FE0D" box="[1147,1323,404,429]" italics="true" pageId="17" pageNumber="18">N. lentiginosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD3697FFE5FFE32E77FE592A78FE76" blockId="17.[264,1324,284,1310]" box="[264,1323,444,470]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FE592A78FE76" box="[264,1323,444,470]" pageId="17" pageNumber="18">­ Gill rakers on second arch 8 or more; anal fin rays 1722 ........................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E32FFE5FFE32E77FE012A78FCB6" pageId="17" pageNumber="18">
<paragraph id="8BDD3697FFE5FFE32E77FE012A79FD3D" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FE012A79FD3D" pageId="17" pageNumber="18">
3 Epioccipital bones extensively invasive over skull surface, and forming with the supraoccipital a basally wide complex process which tapers drastically posteriorly (
<figureCitation id="13592A12FFE5FFE32E3EFDD12EDFFDEE" box="[321,396,564,590]" captionStart="FIGURE 6" captionStartId="7.[264,367,1431,1455]" captionTargetBox="[584,1003,797,1407]" captionTargetId="figure@7.[584,1003,797,1407]" captionTargetPageId="7" captionText="FIGURE 6. Head shield of a female specimen (STRI 5715, 16 mm supraoccipital process length) of Notarius insculptus. CP: complex process; E: epioccipital (= epiotic); S: supraoccipital." httpUri="https://zenodo.org/record/169450/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
); supraoccipital process length shorter than base of complex process width; maxillary barbels relatively long, their length in adult specimens 26.730.3% SL........ ...................................................................................................................
<taxonomicName id="4C624D14FFE5FFE32BEEFD612A79FD3D" ID-CoL="GQVQ" box="[1169,1322,644,669]" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="insculptus">
<emphasis id="B916EA85FFE5FFE32BEEFD612A79FD3D" box="[1169,1322,644,669]" italics="true" pageId="17" pageNumber="18">N. insculptus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD3697FFE5FFE32E77FD492A78FCB6" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FD492A78FCB6" pageId="17" pageNumber="18">
­ Epioccipital bones not invasive or only slightly invasive over skull surface (
<figureCitation id="13592A12FFE5FFE32BB1FD492A4AFD66" box="[1230,1305,684,710]" captionStart="FIGURE 2" captionStartId="3.[264,368,1447,1471]" captionTargetBox="[620,992,920,1428]" captionTargetId="figure@3.[620,992,920,1428]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal view of head of a female paratype (UCR 2451 ­ 2, voucher 95 ­ 7; 72 mm HL) of Notarius biffi, from the Pacific coast of Costa Rica." httpUri="https://zenodo.org/record/169446/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
); supraoccipital process length as long as or longer than its width at base; maxillary barbels relatively short, their length in adult specimens 26.1% SL or less....................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E32FFE5FFE32E77FCC12A7FFC16" pageId="17" pageNumber="18">
<paragraph id="8BDD3697FFE5FFE32E77FCC12A7FFCC6" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FCC12A7FFCC6" pageId="17" pageNumber="18">4 Mouth small, its width 33.942.5% HL; anterior internarial distance 17.924.0% HL; eye relatively large, its diameter 3.5­4.5% SL ............................................................. 5</keyLead>
</paragraph>
<paragraph id="8BDD3697FFE5FFE32E77FC912A7FFC16" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FC912A7FFC16" pageId="17" pageNumber="18">­ Mouth large, its width 44.154.2% HL; anterior internarial distance 25.332.2% HL; eye relatively small, its diameter 2.5­3.7% SL ............................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E32FFE5FFE32E77FC212A79FB05" pageId="17" pageNumber="18">
<paragraph id="8BDD3697FFE5FFE32E77FC212A79FB8D" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FC212A79FB8D" pageId="17" pageNumber="18">
5 Gill rakers on first arch 1113; lips thin; mandibulary barbels comparatively short, their length 10.213.1% (mean 11.6%) SL; caudal peduncle relatively slender, its depth 6.16.7% (mean 6.4%) SL ........................................................................
<taxonomicName id="4C624D14FFE5FFE32BA6FBF12A79FB8D" ID-CoL="6SB5S" box="[1241,1322,1044,1069]" class="Actinopterygii" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="biffi">
<emphasis id="B916EA85FFE5FFE32BA6FBF12A79FB8D" box="[1241,1322,1044,1069]" italics="true" pageId="17" pageNumber="18">N. biffi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDD3697FFE5FFE32E77FBD92A79FB05" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FBD92A79FB05" pageId="17" pageNumber="18">
­ Gill rakers on first arch 810; lips usually thick; mandibulary barbels comparatively long, their length 13.717.7% (mean 16.1%) SL; caudal peduncle relatively deep, its depth 6.87.4% (mean 7.1%) SL
<emphasis id="B916EA85FFE5FFE32DB9FB692BD7FB05" box="[710,1156,1164,1189]" italics="true" pageId="17" pageNumber="18">
........................................
<taxonomicName id="4C624D14FFE5FFE32C8EFB692BD7FB05" ID-CoL="GQY5" box="[1009,1156,1164,1189]" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="planiceps">N. planiceps</taxonomicName>
</emphasis>
/
<taxonomicName id="4C624D14FFE5FFE32BF3FB692A79FB05" ID-CoL="GQY5" box="[1164,1322,1164,1190]" class="Actinopterygii" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="planiceps">
aff.
<emphasis id="B916EA85FFE5FFE32BC5FB692A79FB05" box="[1210,1322,1164,1189]" italics="true" pageId="17" pageNumber="18">planiceps</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30962E32FFE5FFE32E77FB512A78FABD" pageId="17" pageNumber="18">
<paragraph id="8BDD3697FFE5FFE32E77FB512A79FB6D" blockId="17.[264,1324,284,1310]" box="[264,1322,1204,1230]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FB512A79FB6D" box="[264,1322,1204,1230]" pageId="17" pageNumber="18">
6 Supraoccipital process elongated, its base width
<quantity id="4C9A9B72FFE5FFE32C0FFB512CB4FB6E" box="[880,999,1204,1230]" metricMagnitude="-2" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="17" pageNumber="18" unit="in" value="1.65" valueMax="1.7" valueMin="1.6">1.61.7 in</quantity>
its length
<emphasis id="B916EA85FFE5FFE32B1FFB512A79FB6D" box="[1120,1322,1204,1229]" italics="true" pageId="17" pageNumber="18">
............
<taxonomicName id="4C624D14FFE5FFE32BC3FB512A79FB6D" ID-CoL="47S7C" box="[1212,1322,1204,1229]" class="Actinopterygii" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="cookei">N. cookei</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BDD3697FFE5FFE32E77FB392A78FABD" blockId="17.[264,1324,284,1310]" pageId="17" pageNumber="18">
<keyLead id="309395A2FFE5FFE32E77FB392A78FABD" pageId="17" pageNumber="18">
­ Supraoccipital process relatively wide and triangular­shaped, its base width 1.0
<quantity id="4C9A9B72FFE5FFE32B96FB392A78FB56" box="[1257,1323,1244,1270]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="17" pageNumber="18" unit="in" value="1.3">1.3 in</quantity>
its length .......................................................................................................
<taxonomicName id="4C624D14FFE5FFE32BD0FAE12A78FABD" ID-CoL="47S7F" box="[1199,1323,1284,1309]" class="Actinopterygii" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="B916EA85FFE5FFE32BD0FAE12A78FABD" box="[1199,1323,1284,1309]" italics="true" pageId="17" pageNumber="18">N. kessleri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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