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<document id="49A3E1134C825313D140018796169107" ID-CLB-Dataset="22111" ID-DOI="10.11646/zootaxa.4778.3.4" ID-GBIF-Dataset="6384c0b8-822b-46fa-9de7-1d1752536fd8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3828759" ID-ZooBank="8806ED6F-1556-4C68-8B4A-8012704B3D4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589540929688" checkinUser="plazi" docAuthor="Levey, Brian" docDate="2020" docId="03AD2D3AAF38FFD6FF0636D7FB31CF03" docLanguage="en" docName="zootaxa.4778.3.4.pdf" docOrigin="Zootaxa 4778 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anaspis cooteri Levey 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="514" masterDocId="FF945542AF3CFFD3FF913566FFA5CB34" masterDocTitle="A review of the species of Anaspis (s. str.) similar to A. nigripes Brisout and A. apfelbecki Schilsky, with the description of three new species (Coleoptera: Scraptiidae)" masterLastPageNumber="520" masterPageNumber="509" pageNumber="513" updateTime="1698839823653" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5F67C201B446F92A0096597C764F76B1">A review of the species of Anaspis (s. str.) similar to A. nigripes Brisout and A. apfelbecki Schilsky, with the description of three new species (Coleoptera: Scraptiidae)</mods:title>
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<mods:namePart id="6B2F1B56A3CF22F8D22B1916FE43E40D">Levey, Brian</mods:namePart>
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<mods:title id="517E749B3270C290F9AB438A6456CE24">Zootaxa</mods:title>
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<mods:date id="1BE853832096047B838E377E361B0506">2020</mods:date>
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<mods:number id="C960B108DD2C2FAC3CE4A4048EDE6485">2020-05-15</mods:number>
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<treatment id="03AD2D3AAF38FFD6FF0636D7FB31CF03" ID-DOI="http://doi.org/10.5281/zenodo.3848114" ID-GBIF-Taxon="164159708" ID-Zenodo-Dep="3848114" LSID="urn:lsid:plazi:treatment:03AD2D3AAF38FFD6FF0636D7FB31CF03" httpUri="http://treatment.plazi.org/id/03AD2D3AAF38FFD6FF0636D7FB31CF03" lastPageId="5" lastPageNumber="514" pageId="4" pageNumber="513">
<subSubSection id="C31ECFA7AF38FFD7FF0636D7FE3EC8FF" box="[151,411,945,971]" pageId="4" pageNumber="513" type="nomenclature">
<paragraph id="8BBB9C2CAF38FFD7FF0636D7FE3EC8FF" blockId="4.[151,411,945,1007]" box="[151,411,945,971]" pageId="4" pageNumber="513">
<heading id="D0F32B40AF38FFD7FF0636D7FE3EC8FF" bold="true" box="[151,411,945,971]" fontSize="11" level="1" pageId="4" pageNumber="513" reason="1">
<emphasis id="B970403EAF38FFD7FF0636D7FE3EC8FF" bold="true" box="[151,411,945,971]" pageId="4" pageNumber="513">
<taxonomicName id="4C04E7AFAF38FFD7FF0636D7FEF7C8FF" authority="Levey, 2020" authorityName="Levey" authorityYear="2020" box="[151,338,945,971]" class="Insecta" family="Scraptiidae" genus="Anaspis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="513" phylum="Arthropoda" rank="species" species="cooteri" status="sp. nov.">
<emphasis id="B970403EAF38FFD7FF0636D7FEF7C8FF" bold="true" box="[151,338,945,971]" italics="true" pageId="4" pageNumber="513">Anaspis cooteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A243FD45AF38FFD7FEC836D7FE3EC8FF" box="[345,411,945,971]" pageId="4" pageNumber="513" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31ECFA7AF38FFD7FF0636B3FE80CF4B" pageId="4" pageNumber="513" type="description">
<paragraph id="8BBB9C2CAF38FFD7FF0636B3FEC8C8DB" blockId="4.[151,411,945,1007]" box="[151,365,981,1007]" pageId="4" pageNumber="513">
(
<figureCitation id="133F80A9AF38FFD7FF0E36B3FF42C8DB" box="[159,231,981,1007]" captionStart="FIGURES 19" captionStartId="3.[151,264,1626,1651]" captionTargetBox="[158,1431,242,1595]" captionTargetId="figure@3.[151,1436,237,1601]" captionTargetPageId="3" captionText="FIGURES 19. Fore legs of male: 1. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 2. A. nigripes Brisout de Barneville, France, Côte-dOr, Pouilly en-Auxois (NMGW); 3. A. cooteri sp. n., holotype (NMGW); 4. A. curva sp. n., holotype (NMGW); 5. A. bertrami sp. n., holotype (NMGW); 6. A. graeca Schilsky, syntype (BMNH); 7. A. apfelbecki Schilsky, paralectotype (ZMHB); 8. A. apfelbecki Schilsky, paralectotype (ZMHB); 9. A. marseuli Csiki, Japan, Tochigi, Tsurutanuma, Utsunomiya City (TPMJ)." figureDoi="http://doi.org/10.5281/zenodo.3828761" httpUri="https://zenodo.org/record/3828761/files/figure.png" pageId="4" pageNumber="513">Figs 3</figureCitation>
,
<figureCitation id="133F80A9AF38FFD7FF6536B3FEB7C8DB" box="[244,274,981,1007]" captionStart="FIGURES 1017" captionStartId="8.[151,264,1867,1892]" captionTargetBox="[155,1434,186,1839]" captionTargetId="figure@8.[151,1436,181,1842]" captionTargetPageId="8" captionText="FIGURES 1017. Male genitalia: 10. A. curva sp. n., holotype (NMGW); 11. A. bertrami sp. n., holotype (NMGW); 12. A. apfelbecki Schilsky, paralectotype (ZMHB); 13. A. apfelbecki Schilsky, paralectotype (ZMHB); 14. A. cooteri sp. n. holotype (NMGW); 15. A. nigripes Brisout de Barneville, France, Côte-dOr, Pouilly-en-Auxois (NMGW); 16. A. koenigi Schilsky, paralectotype (ZMHB); 17. A. marseuli Csiki, Japan, Tochigi, Yunishikawa, Kuriyama Vill. (TPMJ)." figureDoi="http://doi.org/10.5281/zenodo.3828763" httpUri="https://zenodo.org/record/3828763/files/figure.png" pageId="4" pageNumber="513">14</figureCitation>
,
<figureCitation id="133F80A9AF38FFD7FE8F36B3FE99C8DB" box="[286,316,981,1007]" captionStart="FIGURES 1827" captionStartId="9.[151,264,1862,1887]" captionTargetBox="[190,1403,190,1824]" captionTargetId="figure@9.[181,1407,181,1837]" captionTargetPageId="9" captionText="FIGURES 1827. Apical abdominal ventrites of male: 18. A. apfelbecki Schilsky, paralectotype (ZMHB); 19. A. curva sp. n., holotype; 20. A. bertrami sp. n., holotype (NMGW); 21. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 22. A. nigripes Brisout de Barneville, Spain: La Rioja, Nieva de Cameros, La Dehesa (NMGW); 23. A. koenigi Schilsky, paralectotype (ZMHB); 24. A. marseuli Csiki, Japan, Tochigi City (TPMJ); 25. A. marseuli Csiki, Japan, Utsunomiya City (TPMJ); 26. A. cooteri sp. n., paratype (NMGW); 27. A. graeca Schilsky, syntype (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3828765" httpUri="https://zenodo.org/record/3828765/files/figure.png" pageId="4" pageNumber="513">26</figureCitation>
,
<figureCitation id="133F80A9AF38FFD7FED936B3FEC0C8DB" box="[328,357,981,1007]" captionStart="FIGURES 2836" captionStartId="10.[151,264,1844,1869]" captionTargetBox="[217,1371,195,1810]" captionTargetId="figure@10.[204,1384,181,1819]" captionTargetPageId="10" captionText="FIGURES 2836. Dorsal habitus: 28. Anaspis bertrami sp. n. female paratype (NMGW); 29. A. curva sp. n., male holotype (NMGW); 30. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 31. A. cooteri sp. n., male paratype (NMGW); 32. A. marseuli Csiki, Japan, Hyogo, Haruki, Onsen Town (TPMJ); 33. A. nigripes Brisout de Barneville, male, France, Côte-dOr, Pouilly-en-Auxois (NMGW); 34. A. apfelbecki Schilsky, paralectotype (ZMHB); 35. A. koenigi Schilsky, paralectotype (ZMHB); 36. A. graeca Schilsky, syntype (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3828767" httpUri="https://zenodo.org/record/3828767/files/figure.png" pageId="4" pageNumber="513">31</figureCitation>
)
</paragraph>
<paragraph id="8BBB9C2CAF38FFD7FF06317BFE80CF4B" blockId="4.[151,1437,1052,2015]" pageId="4" pageNumber="513">
<materialsCitation id="3B6C9671AF38FFD7FF06317BFCBECF6F" ID-GBIF-Occurrence="2620263329" collectingDate="2001-06-16" collectionCode="NMGW" collectorName="Mei Yao Yu, N &amp; J. Cooter. &amp; On &amp; Hydrangea &amp; Dipp" country="China" location="Xialongmen" longitude="115.71667" municipality="Dongling Mts." pageId="4" pageNumber="513" specimenCount="1" specimenCount-male="1" stateProvince="Beijing" typeStatus="holotype">
<typeStatus id="54BF228EAF38FFD7FF06317BFF5ACF03" box="[151,255,1053,1079]" pageId="4" pageNumber="513" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0257A5AF38FFD7FE96317BFEB9CF03" box="[263,284,1053,1079]" pageId="4" pageNumber="513" type="male"></specimenCount>
(
<collectionCode id="ED1504E9AF38FFD7FEBD317BFE2ECF03" box="[300,395,1053,1079]" pageId="4" pageNumber="513">NMGW</collectionCode>
)
<collectingCountry id="F313DCBCAF38FFD7FE0A317BFE47CF03" box="[411,482,1053,1079]" name="China" pageId="4" pageNumber="513">China</collectingCountry>
:
<collectingRegion id="49C052CEAF38FFD7FE7C317BFDE7CF03" box="[493,578,1053,1079]" country="China" name="Beijing" pageId="4" pageNumber="513">Beijing</collectingRegion>
,
<collectingMunicipality id="6BDF0656AF38FFD7FDDE317BFD53CF03" box="[591,758,1053,1079]" pageId="4" pageNumber="513">Dongling Mts.</collectingMunicipality>
,
<location id="8EDBCAF7AF38FFD7FC93317BFC2ACF03" LSID="urn:lsid:plazi:treatment:03AD2D3AAF38FFD6FF0636D7FB31CF03:8EDBCAF7AF38FFD7FC93317BFC2ACF03" box="[770,911,1053,1079]" country="China" longitude="115.71667" municipality="Dongling Mts." name="Xialongmen" pageId="4" pageNumber="513" stateProvince="Beijing">Xialongmen</location>
,
<collectorName id="26F1F9FAAF38FFD7FC0D317BFBE0CF03" box="[924,1093,1053,1079]" pageId="4" pageNumber="513">Mei Yao Yu, N</collectorName>
39ᵒ 96,
<geoCoordinate id="EE30FAEBAF38FFD7FB33317AFAB7CF03" box="[1186,1298,1052,1079]" degrees="115" direction="east" minutes="43" orientation="longitude" pageId="4" pageNumber="513" precision="925" value="115.71667">E115ᵒ 43</geoCoordinate>
.
<date id="FFBABAECAF38FFD7FA8C317BFA3CCF03" box="[1309,1433,1053,1079]" pageId="4" pageNumber="513" value="2001-06-16">
<collectingDate id="EFFE4304AF38FFD7FA8C317BFA3CCF03" box="[1309,1433,1053,1079]" pageId="4" pageNumber="513" value="2001-06-16">16.vi.2001</collectingDate>
</date>
.
<collectorName id="26F1F9FAAF38FFD7FF063127FEA7CF6F" box="[151,258,1089,1115]" pageId="4" pageNumber="513">J. Cooter.</collectorName>
/
<collectorName id="26F1F9FAAF38FFD7FE813127FE97CF6F" box="[272,306,1089,1115]" pageId="4" pageNumber="513">On</collectorName>
flowers of
<emphasis id="B970403EAF38FFD7FE233127FD7DCF6F" box="[434,728,1089,1115]" italics="true" pageId="4" pageNumber="513">
<collectorName id="26F1F9FAAF38FFD7FE233127FD95CF6F" box="[434,560,1089,1115]" pageId="4" pageNumber="513">Hydrangea</collectorName>
bretschniederi
</emphasis>
<collectorName id="26F1F9FAAF38FFD7FD4E3127FCBECF6F" box="[735,795,1089,1115]" pageId="4" pageNumber="513">Dipp</collectorName>
</materialsCitation>
.
<materialsCitation id="3B6C9671AF38FFD7FCB43127FE87CF4B" ID-GBIF-Occurrence="2620263319" collectingDate="2001-06-16" collectionCode="NMGW, BMNH, ZMHB" collectorName="Mei Yao Yu, N &amp; J. Cooter. &amp; On &amp; Hydrangea &amp; Dipp" country="China" location="Xialongmen" longitude="115.71667" municipality="Dongling Mts." pageId="4" pageNumber="513" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Beijing" typeStatus="paratype">
<typeStatus id="54BF228EAF38FFD7FCB43127FC37CF6F" box="[805,914,1089,1115]" pageId="4" pageNumber="513" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D0257A5AF38FFD7FC093127FC18CF6F" box="[920,957,1089,1115]" pageId="4" pageNumber="513" type="male">9♂</specimenCount>
,
<specimenCount id="9D0257A5AF38FFD7FC583127FC49CF6F" box="[969,1004,1089,1115]" pageId="4" pageNumber="513" type="female">6♀</specimenCount>
(
<collectionCode id="ED1504E9AF38FFD7FC6B3127FBFDCF6F" box="[1018,1112,1089,1115]" pageId="4" pageNumber="513">NMGW</collectionCode>
,
<collectionCode id="ED1504E9AF38FFD7FBF23127FB1FCF6F" box="[1123,1210,1089,1115]" pageId="4" pageNumber="513">BMNH</collectionCode>
,
<collectionCode id="ED1504E9AF38FFD7FB543127FABFCF6F" box="[1221,1306,1089,1115]" pageId="4" pageNumber="513">ZMHB</collectionCode>
), same data as Holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31ECFA7AF38FFD6FF5631EEFC75CF27" lastPageId="5" lastPageNumber="514" pageId="4" pageNumber="513" type="diagnosis">
<paragraph id="8BBB9C2CAF38FFD7FF5631EEFB84CE63" blockId="4.[151,1437,1052,2015]" pageId="4" pageNumber="513">
<emphasis id="B970403EAF38FFD7FF5631EEFEE7CF97" bold="true" box="[199,322,1160,1187]" pageId="4" pageNumber="513">Diagnosis.</emphasis>
General diagnosis of
<typeStatus id="54BF228EAF38FFD7FDA031EFFD36CF97" box="[561,659,1161,1187]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
(male): Length (fore margin of pronotum to apex of elytra in mid line)
<quantity id="4CFC31C9AF38FFD7FF0631CBFF54CFF3" box="[151,241,1197,1223]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="513" unit="mm" value="2.7">2.7 mm</quantity>
. Head, except for distal margin of fronto-clypeus, pronotum, scutellum, elytra and underside black; distal margin of fronto-clypeus and labrum yellow brown, labrum darker at centre; maxillary palpi, basal four segments of antennae, fore tibia and tibial spurs of all legs yellow; apical six segments of antennae black; coxae, trochanters and femora of all legs, and tibia of mid and hind legs brown; tarsi brown, first and second segments of mid and hind legs paler in basal half. Entire body densely covered with short grey pubescence.
</paragraph>
<paragraph id="8BBB9C2CAF38FFD7FF563007FA23CD43" blockId="4.[151,1437,1052,2015]" pageId="4" pageNumber="513">
Head. Antennae: first antennomere almost rectangular, about 1.5× as long as wide; second antennomere cylindrical about as long as first; third antennomere cylindrical, slightly widened at apex, as long as second; fourth antennomere like third, but slightly longer; fifth to tenth antennomeres slightly shorter than fourth becoming progressively broader; eleventh antennomere ovate, slightly longer than tenth. Apical segment of maxillary palp securiform. Pronotum: 1.2× as wide at base as long in mid line; hind angles obtuse, rounded; lateral edge straight when viewed from side. Scutellum: not visible in
<typeStatus id="54BF228EAF38FFD7FDB33373FD2DCD1B" box="[546,648,1557,1583]" pageId="4" pageNumber="513" type="holotype">holotype</typeStatus>
. Elytra: 2.9× as long as pronotum. Abdomen: Ventrites 14 convex, but not keeled; 3 with a pair of small, thin, very slightly curved appendages, joined at the base; 4 with a pair of much shorter slightly curved appendages; overlapped by those of segment 3; 5 with a narrow V shaped notch at centre.
</paragraph>
<paragraph id="8BBB9C2CAF38FFD6FF5633E7FB24C92F" blockId="4.[151,1437,1052,2015]" lastBlockId="5.[151,1437,153,1079]" lastPageId="5" lastPageNumber="514" pageId="4" pageNumber="513">
Legs: Fore leg (
<figureCitation id="133F80A9AF38FFD7FEEB33E7FE19CDAF" box="[378,444,1665,1691]" captionStart="FIGURES 19" captionStartId="3.[151,264,1626,1651]" captionTargetBox="[158,1431,242,1595]" captionTargetId="figure@3.[151,1436,237,1601]" captionTargetPageId="3" captionText="FIGURES 19. Fore legs of male: 1. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 2. A. nigripes Brisout de Barneville, France, Côte-dOr, Pouilly en-Auxois (NMGW); 3. A. cooteri sp. n., holotype (NMGW); 4. A. curva sp. n., holotype (NMGW); 5. A. bertrami sp. n., holotype (NMGW); 6. A. graeca Schilsky, syntype (BMNH); 7. A. apfelbecki Schilsky, paralectotype (ZMHB); 8. A. apfelbecki Schilsky, paralectotype (ZMHB); 9. A. marseuli Csiki, Japan, Tochigi, Tsurutanuma, Utsunomiya City (TPMJ)." figureDoi="http://doi.org/10.5281/zenodo.3828761" httpUri="https://zenodo.org/record/3828761/files/figure.png" pageId="4" pageNumber="513">Fig. 3</figureCitation>
) with femora moderately flattened anterior-posterior, and slightly excavated on the posterior surface, the ventral edge concavely excavate; tibia about same length as femora, slightly flattened anteriorposterior, almost parallel sided for most of its length, but abruptly narrowing at basal sixth, with a dense distal fringe of yellow spines and two yellow spurs slightly longer than tarsal segment 1; tarsal segment 1 flattened, cylindrical, about twice as long as wide; 2 about as long and as wide as 1, slightly widening distally; 3 slightly shorter than 2, approximately heart shaped with a deep medial excision distally, about as wide as long, with a long semicircular ventral pad extending distally beneath the entire length of segment 4; 4 very small, almost triangular, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, slightly widened at base. Mid leg with femora strongly anterior-posterior flattened; tibia about as long as femora, regularly widening from base to apex, the ventral margin almost straight; with a dense distal fringe of yellow spines and a pair of long yellow spurs, the longer slightly less than half the length of tarsal segment 1, the shorter about one-third length of segment 1; tarsal segment 1 long, cylindrical, 0.48 length of tibia; 2 shorter, cylindrical, 0.56 length of 1; 3 shorter, 0.55 length of 2, approximately heart shaped with a deep medial excision, with a long semicircular ventral pad; 4 very small oblong, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, slightly widened at base. Hind leg with femora strongly flattened anterior-posterior; tibia about half three quarters length of femora, strongly widened from base to apex, about 4.0× as long as wide at apex, with a dense distal fringe of short yellow spines and two yellow spurs, the longer about half the length and the shorter about onethird the length of tarsal segment 1; tarsal segment 1 moderately widening at apex, about four× as long as wide at apex, about 0.63 length of tibia; 2 slightly widening at apex, about 0.68 length of 1; 3 very slightly widening at apex, about 0.83 length of 2; 4 cylindrical, about 1.18× length of 3, claws small, slightly curved, slightly widened at base. Genitalia (
<figureCitation id="133F80A9AF39FFD6FE9C3767FEC7C92F" box="[269,354,513,539]" captionStart="FIGURES 1017" captionStartId="8.[151,264,1867,1892]" captionTargetBox="[155,1434,186,1839]" captionTargetId="figure@8.[151,1436,181,1842]" captionTargetPageId="8" captionText="FIGURES 1017. Male genitalia: 10. A. curva sp. n., holotype (NMGW); 11. A. bertrami sp. n., holotype (NMGW); 12. A. apfelbecki Schilsky, paralectotype (ZMHB); 13. A. apfelbecki Schilsky, paralectotype (ZMHB); 14. A. cooteri sp. n. holotype (NMGW); 15. A. nigripes Brisout de Barneville, France, Côte-dOr, Pouilly-en-Auxois (NMGW); 16. A. koenigi Schilsky, paralectotype (ZMHB); 17. A. marseuli Csiki, Japan, Tochigi, Yunishikawa, Kuriyama Vill. (TPMJ)." figureDoi="http://doi.org/10.5281/zenodo.3828763" httpUri="https://zenodo.org/record/3828763/files/figure.png" pageId="5" pageNumber="514">Fig. 14</figureCitation>
): Parameres not hooked at apex; basal piece 2.3× as long as parameres.
</paragraph>
<paragraph id="8BBB9C2CAF39FFD6FF563743FB9FC9B3" blockId="5.[151,1437,153,1079]" pageId="5" pageNumber="514">
Variation: Length 2.73.0 mm; the only colour variation noticed is that the apical segment of the maxillary palp is partly black along the dorsal edge in some
<specimenCount id="9D0257A5AF39FFD6FD01372CFD00C950" box="[656,677,586,612]" pageId="5" pageNumber="514" type="male"></specimenCount>
specimens. Pronotum: 1.21.3× as wide at base as long in mid line. Scutellum: Triangular about as wide as long. Elytra: 2.73.2× as long as pronotum.
</paragraph>
<paragraph id="8BBB9C2CAF39FFD6FF5637F6FDE1C893" blockId="5.[151,1437,153,1079]" pageId="5" pageNumber="514">
<emphasis id="B970403EAF39FFD6FF5637F6FDBFC99F" bold="true" box="[199,538,656,683]" pageId="5" pageNumber="514">General diagnosis of female:</emphasis>
Length
<quantity id="4CFC31C9AF39FFD6FDE737F7FCA5C99F" box="[630,768,657,683]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="5" pageNumber="514" unit="mm" value="2.95" valueMax="3.2" valueMin="2.7">2.73.2 mm</quantity>
. Colour as in
<specimenCount id="9D0257A5AF39FFD6FC0937F7FC15C99F" box="[920,944,657,683]" pageId="5" pageNumber="514" type="male"></specimenCount>
. Head: as in
<specimenCount id="9D0257A5AF39FFD6FBAF37F7FBF3C99F" box="[1086,1110,657,683]" pageId="5" pageNumber="514" type="male"></specimenCount>
. Pronotum: 1.31.4× as wide at base as long in mid line. Scutellum: as in male. Elytra: 3.03.3× as long as pronotum. Abdomen: as in male but without appendages and 5
<superScript id="7C713164AF39FFD6FE2E37BEFE69C9D2" attach="left" box="[447,460,728,742]" fontSize="6" pageId="5" pageNumber="514">th</superScript>
ventrite without a notch, broadly rounded at apex. Legs: Fore leg as in male but with the following differences: femora not flattened, not excavate on the posterior surface and with ventral edge not concavely excavate; tibia not obviously flattened anterior-posterior, not parallel sided and more gradually narrowing at basal sixth; tarsal segment 1 elongate cylindrical about three to four times as long as wide, 2 about two-thirds length of 1, slightly widening at apex; 3 about three-quarters length of 2, more parallel sided, less heart shaped than in male. Mid and hind legs as in male.
</paragraph>
<paragraph id="8BBB9C2CAF39FFD6FF5636D6FC75CF27" blockId="5.[151,1437,153,1079]" pageId="5" pageNumber="514">
<emphasis id="B970403EAF39FFD6FF5636D6FE44C8FF" bold="true" box="[199,481,944,971]" pageId="5" pageNumber="514">Comparative diagnosis.</emphasis>
This species appears to be most closely related to
<taxonomicName id="4C04E7AFAF39FFD6FB8836D7FB31C8FF" box="[1049,1172,945,971]" class="Insecta" family="Scraptiidae" genus="Anaspis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="514" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="B970403EAF39FFD6FB8836D7FB31C8FF" box="[1049,1172,945,971]" italics="true" pageId="5" pageNumber="514">A. nigripes</emphasis>
</taxonomicName>
differing in the smaller appendages of the ventrite 3 of the male (compare
<figureCitation id="133F80A9AF39FFD6FD5B36B3FCFAC8DB" box="[714,863,981,1007]" captionStart="FIGURES 1827" captionStartId="9.[151,264,1862,1887]" captionTargetBox="[190,1403,190,1824]" captionTargetId="figure@9.[181,1407,181,1837]" captionTargetPageId="9" captionText="FIGURES 1827. Apical abdominal ventrites of male: 18. A. apfelbecki Schilsky, paralectotype (ZMHB); 19. A. curva sp. n., holotype; 20. A. bertrami sp. n., holotype (NMGW); 21. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 22. A. nigripes Brisout de Barneville, Spain: La Rioja, Nieva de Cameros, La Dehesa (NMGW); 23. A. koenigi Schilsky, paralectotype (ZMHB); 24. A. marseuli Csiki, Japan, Tochigi City (TPMJ); 25. A. marseuli Csiki, Japan, Utsunomiya City (TPMJ); 26. A. cooteri sp. n., paratype (NMGW); 27. A. graeca Schilsky, syntype (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3828765" httpUri="https://zenodo.org/record/3828765/files/figure.png" pageId="5" pageNumber="514">Figs 26 &amp; 22</figureCitation>
); the more elongate pronotum, with the lateral margins almost parallel sided in front of hind angles (compare
<figureCitation id="133F80A9AF39FFD6FCBD369FFC67CF27" box="[812,962,1017,1043]" captionStart="FIGURES 2836" captionStartId="10.[151,264,1844,1869]" captionTargetBox="[217,1371,195,1810]" captionTargetId="figure@10.[204,1384,181,1819]" captionTargetPageId="10" captionText="FIGURES 2836. Dorsal habitus: 28. Anaspis bertrami sp. n. female paratype (NMGW); 29. A. curva sp. n., male holotype (NMGW); 30. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 31. A. cooteri sp. n., male paratype (NMGW); 32. A. marseuli Csiki, Japan, Hyogo, Haruki, Onsen Town (TPMJ); 33. A. nigripes Brisout de Barneville, male, France, Côte-dOr, Pouilly-en-Auxois (NMGW); 34. A. apfelbecki Schilsky, paralectotype (ZMHB); 35. A. koenigi Schilsky, paralectotype (ZMHB); 36. A. graeca Schilsky, syntype (BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3828767" httpUri="https://zenodo.org/record/3828767/files/figure.png" pageId="5" pageNumber="514">Figs 31 &amp; 33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31ECFA7AF39FFD6FF56317BFB31CF03" box="[199,1172,1053,1079]" pageId="5" pageNumber="514" type="etymology">
<paragraph id="8BBB9C2CAF39FFD6FF56317BFB31CF03" blockId="5.[151,1437,153,1079]" box="[199,1172,1053,1079]" pageId="5" pageNumber="514">
<emphasis id="B970403EAF39FFD6FF56317BFEEBCF03" bold="true" box="[199,334,1053,1079]" pageId="5" pageNumber="514">Etymology.</emphasis>
This species is named for Jonathan Cooter, the collector of the
<typeStatus id="54BF228EAF39FFD6FB87317BFBE3CF03" box="[1046,1094,1053,1079]" pageId="5" pageNumber="514">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>