204 lines
30 KiB
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204 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="4D7DDCB5787155D1B49BC0702346EF47" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria tumida Miller 2022, sp. nov." docType="treatment" docUuid="984AAC32-D689-4CD8-BD12-382AFEF9DB5E" docUuidSource="ZooBank" docVersion="1" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287389350" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Miller, Kelly B.</mods:namePart>
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<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
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<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1136</mods:number>
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<mods:end>56</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1136-1</mods:identifier>
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<mods:identifier type="ZooBank">11C6BBFB339A4672AE662CE2B1E6321E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5D0E6613B1D05E4293ABEE4BFD1CA18A</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:984AAC32-D689-4CD8-BD12-382AFEF9DB5E" httpUri="http://treatment.plazi.org/id/4D7DDCB5787155D1B49BC0702346EF47" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/984AAC32-D689-4CD8-BD12-382AFEF9DB5E" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tumida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tumida" status="sp. nov.">Desmopachria tumida</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">Figures 23-27</figureCitation>
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<figureCitation captionStart="Figures 76, 77" captionStartId="F7" captionText="Figures 76, 77. Desmopachria species distributions. 76 ungrouped Desmopachria species 77 Desmopachria apicodente species group." figureDoi="10.3897/zookeys.1136.72744.figures76-77" httpUri="https://binary.pensoft.net/fig/782459" pageId="0" pageNumber="1">, 77</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Venezuela, Bolivar State, Gran Sabana, Pauji, Esmeraldes,
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<geoCoordinate degrees="4" direction="north" minutes="28.233" orientation="latitude" precision="1" value="4.47055">4°28.233'N</geoCoordinate>
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,
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<geoCoordinate degrees="17" direction="west" minutes="35.559" orientation="longitude" precision="1" value="-17.59265">17°35.559'W</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is characterized by the distinctive lateral longitudinal tumidity on the elytron (Figs
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">23</figureCitation>
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,
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">24</figureCitation>
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). The humeral angle in dorsal aspect is subsinuate because of lateral expansion of the tumidity. The male genitalia are distinct with the median lobe broad with the lateral margins medially bisinuate and apically narrowed to an apically rounded apex (Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">25</figureCitation>
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) and in lateral aspect with the base large and rounded and the apical portion with the dorsal margin linear and with a broad expansion on the ventral margin with the apex slender and apically narrowly rounded (Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">26</figureCitation>
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). The lateral lobe is broad medially and apically tapered to a pointed apex with a distinct apical socketed
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<normalizedToken originalValue="“tooth”">"tooth"</normalizedToken>
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(Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">27</figureCitation>
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). This tooth is shared with two other species,
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<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria apicodente" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apicodente">
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<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria apicodente</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lateralis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lateralis">
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<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lateralis</emphasis>
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</taxonomicName>
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sp. nov., together making up the
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<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria apicodente" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apicodente">
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<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria apicodente</emphasis>
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</taxonomicName>
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species-group (see Diagnosis under
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<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria apicodente" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apicodente">
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<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria apicodente</emphasis>
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</taxonomicName>
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for discussion of differences).
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<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lateralis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lateralis">
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<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lateralis</emphasis>
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</taxonomicName>
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also has a prominent lateral longitudinal elytral tumidity, but that species has differently shaped male genitalia (Figs
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">20-22</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
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TL = 2.0 mm, GW = 1.5 mm, PW = 1.1 mm, HW = 0.7 mm, EW = 0.4 mm, TL/GW = 1.4, HW/EW = 1.8. Body very broad, laterally broadly rounded, lateral margins nearly continuously curved between pronotum and elytron (Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">23</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration.</emphasis>
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Head and pronotum evenly orange. Elytron brown, laterally and apically somewhat paler brown-orange. Ventral surfaces and appendages orange to orange-brown.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
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Head broad, short; anterior margin of clypeus distinctly margined with continuous flattened bead, broader and flatter medially; surface of head shiny, punctation extremely fine and sparse; eyes large (HW/EW = 1.8); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, gently curved with continuous narrow marginal bead; surface shiny, punctation very fine, of the same size and evenly distributed except more coarsely punctate posteromedially. Elytron broad, laterally broadly curved, lateral margin distinctly sinuate at humeral angle, with distinct lateral tumidity extending posteriorly from humeral angle (Figs
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">23</figureCitation>
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,
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">24</figureCitation>
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); surface shiny, punctation of two sizes, minute and small, evenly punctate. Prosternum extremely short, longitudinally compressed, medially flattened; prosternal process broad, flattened, concave with broad lateral bead, apically pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, impunctate; metaventrite wings extremely slender. Metacoxa with medial portion short, <1/3 length of metaventrite medially, metacoxal lines slightly sinuate, divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, extremely minutely punctate. Metatrochanter very large, longer than length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, surface impunctate.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
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Male median lobe in in lateral aspect with large, broad, rounded base, apical portion with dorsal margin linear, ventral margin with large expansion medially, apically narrowed and slender to narrowly rounded apex (Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">25</figureCitation>
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); in ventral aspect broad, long, lateral margins distinctly sinuate medially, apically narrowed to broadly rounded apex (Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">26</figureCitation>
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). Lateral lobe large, broad medially, narrow basally and apically strongly and evenly narrowed to pointed apex, dorsal margin with long dense series of long setae (Fig.
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<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">27</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism and variation.</emphasis>
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Only a single male specimen was examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is named
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<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tumida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tumida">
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||
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tumida</emphasis>
|
||
</taxonomicName>
|
||
, Latin for swollen, for the laterally tumid, or swollen, elytral margins (Figs
|
||
<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">23</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 13–27" captionStartId="F2" captionText="Figures 13 - 27. Desmopachria apicodente - group species. (13 - 17) Desmopachria apicodente 13 habitus 14 male median lobe, right lateral aspect 15 male median lobe, ventral aspect 16 male median lobe and lateral lobes, ventral aspect 17 right lateral lobe, right lateral aspect (18 - 22) Desmopachria lateralis 18 habitus 19 left elytron, left lateral aspect 20 male median lobe, right lateral aspect 21 male median lobe and right lateral lobe, ventral aspect 22 right lateral lobe, right lateral aspect (23 - 27) Desmopachria tumida 23 habitus 24 left elytron, left lateral aspect 25 male median lobe, right lateral aspect 26 male median lobe, ventral aspect 27 right lateral lobe, right lateral aspect. Scale bars: 1.0 mm for habitus drawings." figureDoi="10.3897/zookeys.1136.72744.figures13-27" httpUri="https://binary.pensoft.net/fig/782454" pageId="0" pageNumber="1">24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species is known only from one site in the Gran Sabana of Bolivar State, Venezuela (Fig.
|
||
<figureCitation captionStart="Figures 76, 77" captionStartId="F7" captionText="Figures 76, 77. Desmopachria species distributions. 76 ungrouped Desmopachria species 77 Desmopachria apicodente species group." figureDoi="10.3897/zookeys.1136.72744.figures76-77" httpUri="https://binary.pensoft.net/fig/782459" pageId="0" pageNumber="1">77</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Holotype in MIZA, male labeled, "VENEZUELA: Bolivar State
|
||
<geoCoordinate degrees="4" direction="north" minutes="28.233" orientation="latitude" precision="1" value="4.47055">4°28.233'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="61" direction="west" minutes="35.559" orientation="longitude" precision="1" value="-61.59265">61°35.559'W</geoCoordinate>
|
||
, 867 m Gran Sabana, Pauji: Esmeraldes 16.vii.2010;leg. Short, Tellez, Arias detrital pools by forested stream VZ10-0716-02A/ SEMC0908227 KUNHM-ENT [barcode label]/ HOLOTYPE
|
||
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tumida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tumida">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tumida</emphasis>
|
||
</taxonomicName>
|
||
Miller, 2021 [red label with black line border]." This species is only known from the holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="checklist of desmopachria apicodente species group">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Checklist of
|
||
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria apicodente" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apicodente">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria apicodente</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria apicodente" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apicodente">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria apicodente</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. - Guyana, Venezuela
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lateralis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lateralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lateralis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. - Venezuela
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria tumida" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tumida">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria tumida</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. - Venezuela
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |