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<document id="B93A81AB9DC9DDD038B1AC5C9E8B21BC" ID-CLB-Dataset="28139" ID-DOI="10.11646/zootaxa.4563.3.1" ID-GBIF-Dataset="cf7fa86d-ae34-4e8d-aa9d-7c15568bc995" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601295" ID-ZooBank="B0A3408F-563A-4DD3-94A4-284A2770B0A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171996401" checkinUser="plazi" docAuthor="Van Hinsbergh, Victor W. M. &amp; Helwerda, Renate A." docDate="2019" docId="C5011D20FFDEFFFFFF01FC66C78AA834" docLanguage="en" docName="zootaxa.4563.3.1.pdf" docOrigin="Zootaxa 4563 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maurolicus muelleri" docType="treatment" docVersion="10" lastPageNumber="407" masterDocId="39386558FFDBFFF9FF96FFB2C62CAC3B" masterDocTitle="Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines" masterLastPageNumber="443" masterPageNumber="401" pageNumber="406" updateTime="1698703400501" updateUser="ExternalLinkService">
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<mods:title id="C41B15B83B4512CB1D126E604041F03D">Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines</mods:title>
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<mods:namePart id="96FCCCDD31BC13C562238D7ED328999D">Van Hinsbergh, Victor W. M.</mods:namePart>
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<mods:identifier id="9D8DBC26487816645441BDE6A8514F90" type="DOI">10.11646/zootaxa.4563.3.1</mods:identifier>
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<treatment id="C5011D20FFDEFFFFFF01FC66C78AA834" ID-DOI="http://doi.org/10.5281/zenodo.5936993" ID-GBIF-Taxon="154474727" ID-Zenodo-Dep="5936993" LSID="urn:lsid:plazi:treatment:C5011D20FFDEFFFFFF01FC66C78AA834" httpUri="http://treatment.plazi.org/id/C5011D20FFDEFFFFFF01FC66C78AA834" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406">
<subSubSection id="05B2FFBDFFDEFFFCFF01FC66C710A834" pageId="5" pageNumber="406" type="nomenclature">
<paragraph id="4D17AC36FFDEFFFCFF01FC66C459AFD5" blockId="5.[151,629,980,1039]" box="[151,629,980,1006]" pageId="5" pageNumber="406">
<heading id="165F1B5AFFDEFFFCFF01FC66C459AFD5" bold="true" box="[151,629,980,1006]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
<taxonomicName id="8AA8D7B5FFDEFFFCFF01FC66C459AFD5" ID-CoL="3YG5C" authority="(Gmelin, 1789)" baseAuthorityName="Gmelin" baseAuthorityYear="1789" box="[151,629,980,1006]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF01FC66C459AFD5" bold="true" box="[151,629,980,1006]" pageId="5" pageNumber="406">
<emphasis id="7FDC7024FFDEFFFCFF01FC66C70FAFD5" bold="true" box="[151,291,980,1006]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDEFFFCFED9FC66C798AFD5" bold="true" box="[335,436,980,1006]" italics="true" pageId="5" pageNumber="406">muelleri</emphasis>
(Gmelin, 1789)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FC44C710A834" blockId="5.[151,629,980,1039]" box="[151,316,1014,1039]" pageId="5" pageNumber="406">(Figures 9–13)</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFDEFFFFFF01FB8CC78AA834" lastPageId="6" lastPageNumber="407" pageId="5" pageNumber="406" type="description">
<paragraph id="4D17AC36FFDEFFFCFF01FB8CC5F1A86F" blockId="5.[151,1066,1086,1292]" box="[151,989,1086,1108]" pageId="5" pageNumber="406">
1934
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FB8CC4B4A86F" authority="Chaine &amp; Duvergier" authorityName="Chaine &amp; Duvergier" box="[245,664,1086,1108]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FB8CC7EFA86F" box="[245,451,1086,1108]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Chaine &amp; Duvergier
</taxonomicName>
(re-pictured in Nolf etal., 2009)
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FBEFC4D6A848" blockId="5.[151,1066,1086,1292]" box="[151,762,1117,1139]" pageId="5" pageNumber="406">
1980
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FBEFC441A848" authority="Schwarzhans W" authorityName="Schwarzhans W" box="[245,621,1117,1139]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FBEFC7EFA848" box="[245,451,1117,1139]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Schwarzhans W
</taxonomicName>
, p. 31, fig. 54
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FBCEC5F4A8A9" blockId="5.[151,1066,1086,1292]" box="[151,984,1148,1170]" pageId="5" pageNumber="406">
1995
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FBCEC425A8A9" authority="Smale" authorityName="Smale" box="[245,521,1148,1170]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FBCEC7EFA8A9" box="[245,451,1148,1170]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Smale
</taxonomicName>
MJ, Watson G, Hecht T, p. 29, plate 11, E1–3
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FB28C5FDA88B" blockId="5.[151,1066,1086,1292]" box="[151,977,1178,1200]" pageId="5" pageNumber="406">
1999
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FB28C4B3A88B" authority="Rivaton J, Bourret P" authorityName="Rivaton J, Bourret P" box="[245,671,1178,1200]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FB28C7EFA88B" box="[245,451,1178,1200]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Rivaton J, Bourret P
</taxonomicName>
, pp. 204-205, pl.93 fig. 13–16
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FB0BC206A8F4" blockId="5.[151,1066,1086,1292]" box="[151,1066,1209,1231]" pageId="5" pageNumber="406">
2008
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FB0BC427A8F4" authority="Tusset" authorityName="Tusset" box="[245,523,1209,1231]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FB0BC7EFA8F4" box="[245,451,1209,1231]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Tusset
</taxonomicName>
VM, Lombarte A, Assis CA, pp. 38-39, fig. 19 A1–3
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FB6AC56BA8D5" blockId="5.[151,1066,1086,1292]" box="[151,839,1240,1262]" pageId="5" pageNumber="406">
2012
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FB6AC7C1A8D5" authority="Lin" authorityName="Lin" box="[245,493,1240,1262]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FB6AC7EFA8D5" box="[245,451,1240,1262]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Lin
</taxonomicName>
<collectingCountry id="35BFECA6FFDEFFFCFE65FB6AC43AA8D5" box="[499,534,1240,1262]" name="Switzerland" pageId="5" pageNumber="406">CH</collectingCountry>
and Chang CW, p. 75, plate 7
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FB44C443A937" blockId="5.[151,1066,1086,1292]" box="[151,623,1270,1292]" pageId="5" pageNumber="406">
2013
<taxonomicName id="8AA8D7B5FFDEFFFCFF63FB44C43BA937" authority="Nolf D." authorityName="Nolf D." box="[245,535,1270,1292]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF63FB44C7EFA937" box="[245,451,1270,1292]" italics="true" pageId="5" pageNumber="406">Maurolicus muelleri</emphasis>
Nolf D.
</taxonomicName>
plate 43
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FAEFC72FA948" blockId="5.[151,1436,1373,2008]" box="[151,259,1373,1395]" pageId="5" pageNumber="406">
<emphasis id="7FDC7024FFDEFFFCFF01FAEFC72FA948" bold="true" box="[151,259,1373,1395]" pageId="5" pageNumber="406">PLATE 1.</emphasis>
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FACEC28CA9F4" blockId="5.[151,1436,1373,2008]" pageId="5" pageNumber="406">
1.
<taxonomicName id="8AA8D7B5FFDEFFFCFF22FACEC73BA9AA" authorityName="Swainson" authorityYear="1838" box="[180,279,1404,1425]" class="Actinopterygii" family="Congridae" genus="Ariosoma" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF22FACEC73BA9AA" box="[180,279,1404,1425]" italics="true" pageId="5" pageNumber="406">Ariosoma</emphasis>
</taxonomicName>
sp., RGM 962092, locality AndaDeVos. For this figure and remaining figures, all material derives from the Piacenzian or Gelasian Santa Cruz Formation of Cabarruyan Island or nearby Luzon Island,
<collectingCountry id="35BFECA6FFDEFFFCFBACFA28C283A98B" box="[1082,1199,1434,1456]" name="Philippines" pageId="5" pageNumber="406">Philippines</collectingCountry>
. Figures display otolith medial faces, which were imaged with light microscope Z-stack photography, unless stated otherwise.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FA6AC58DA9D5" blockId="5.[151,1436,1373,2008]" box="[151,929,1496,1518]" pageId="5" pageNumber="406">
2.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26FA6AC711A9D5" authorityName="Ogilby" authorityYear="1898" box="[176,317,1496,1518]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF26FA6AC711A9D5" box="[176,317,1496,1518]" italics="true" pageId="5" pageNumber="406">Bathycongrus</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="2939D1C7FFDEFFFCFEF0FA6BC7F0A9D5" author="Frost, G. A." box="[358,476,1496,1518]" pageId="5" pageNumber="406" pagination="133 - 142" refId="ref18329" refString="Frost, G. A. (1933) Otoliths of fishes from the Tertiary formations of New Zealand. Transactions and Proceedings of the Royal Society of New Zealand, 63, 133 - 142. [figures redrawn in Schwarzhans, W. (1980) pages 167 - 169]" type="journal article" year="1933">Frost, 1933</bibRefCitation>
, RGM 962094, locality Anda2, SEM image.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01FA44C5D3AA37" blockId="5.[151,1436,1373,2008]" box="[151,1023,1526,1549]" pageId="5" pageNumber="406">
3.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26FA44C711AA37" authorityName="Ogilby" authorityYear="1898" box="[176,317,1526,1548]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF26FA44C711AA37" box="[176,317,1526,1548]" italics="true" pageId="5" pageNumber="406">Bathycongrus</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="2939D1C7FFDEFFFCFEF0FA45C7F6AA37" author="Frost, G. A." box="[358,474,1526,1548]" pageId="5" pageNumber="406" pagination="133 - 142" refId="ref18329" refString="Frost, G. A. (1933) Otoliths of fishes from the Tertiary formations of New Zealand. Transactions and Proceedings of the Royal Society of New Zealand, 63, 133 - 142. [figures redrawn in Schwarzhans, W. (1980) pages 167 - 169]" type="journal article" year="1933">Frost, 1933</bibRefCitation>
, juvenile, RGM 962095, locality Anda2, SEM image.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F9A7C537AA71" blockId="5.[151,1436,1373,2008]" pageId="5" pageNumber="406">
4.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26F9A7C711AA10" authorityName="Ogilby" authorityYear="1898" box="[176,317,1557,1579]" class="Actinopterygii" family="Congridae" genus="Bathycongrus" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF26F9A7C711AA10" box="[176,317,1557,1579]" italics="true" pageId="5" pageNumber="406">Bathycongrus</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="2939D1C7FFDEFFFCFEF0F9A4C7F6AA10" author="Frost, G. A." box="[358,474,1557,1579]" pageId="5" pageNumber="406" pagination="133 - 142" refId="ref18329" refString="Frost, G. A. (1933) Otoliths of fishes from the Tertiary formations of New Zealand. Transactions and Proceedings of the Royal Society of New Zealand, 63, 133 - 142. [figures redrawn in Schwarzhans, W. (1980) pages 167 - 169]" type="journal article" year="1933">Frost, 1933</bibRefCitation>
juvenile, RGM 962096, locality Anda3. (a) SEM image, (b) light microscopy image. 5.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26F986C716AA71" authorityName="Kaup" authorityYear="1856" box="[176,314,1588,1610]" class="Actinopterygii" family="Congridae" genus="Congridarum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF26F986C716AA71" box="[176,314,1588,1610]" italics="true" pageId="5" pageNumber="406">Congridarum</emphasis>
</taxonomicName>
sp., RGM 962093, locality Tiep2, SEM image.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F9E0C791AABC" blockId="5.[151,1436,1373,2008]" pageId="5" pageNumber="406">
6.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26F9E0C799AA53" authorityName="Temminck &amp; Schlegel" authorityYear="1846" box="[176,437,1618,1640]" class="Actinopterygii" family="Gonorynchidae" genus="Gonorynchus" kingdom="Animalia" order="Gonorynchiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="abbreviatus">
<emphasis id="7FDC7024FFDEFFFCFF26F9E0C799AA53" box="[176,437,1618,1640]" italics="true" pageId="5" pageNumber="406">Gonorynchus abbreviatus</emphasis>
</taxonomicName>
, RGM 962097, locality Roxas. (a) SEM image, (b) light microscopy image, (c) ventral profile, (d) lateral face, (e) ostial profile.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F923C4D2AA9D" blockId="5.[151,1436,1373,2008]" box="[151,766,1680,1702]" pageId="5" pageNumber="406">
7.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26F923C739AA9D" authorityName="Linnaeus" authorityYear="1758" box="[176,277,1681,1702]" class="Actinopterygii" family="Argentinidae" genus="Argentina" kingdom="Animalia" order="Osmeriformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF26F923C739AA9D" box="[176,277,1681,1702]" italics="true" pageId="5" pageNumber="406">Argentina</emphasis>
</taxonomicName>
sp., RGM 962098, locality Anda4, SEM image.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F91CC4C2AAFF" blockId="5.[151,1436,1373,2008]" box="[151,750,1710,1732]" pageId="5" pageNumber="406">
8.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26F91CC79CAAFF" box="[176,432,1710,1732]" class="Actinopterygii" family="Opisthoproctidae" genus="Opisthoproctus" kingdom="Animalia" order="Osmeriformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="grimaldii">
<emphasis id="7FDC7024FFDEFFFCFF26F91CC79CAAFF" box="[176,432,1710,1732]" italics="true" pageId="5" pageNumber="406">Opisthoproctus grimaldii</emphasis>
</taxonomicName>
, RGM 962099, locality Roxas.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F97FC4CDAAD8" blockId="5.[151,1436,1373,2008]" box="[151,737,1741,1763]" pageId="5" pageNumber="406">
9.
<taxonomicName id="8AA8D7B5FFDEFFFCFF26F97FC7B3AAD8" box="[176,415,1741,1763]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF26F97FC708AAD8" box="[176,292,1741,1763]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDEFFFCFEDDF97FC7B3AAD8" box="[331,415,1741,1763]" italics="true" pageId="5" pageNumber="406">muelleri</emphasis>
</taxonomicName>
, RGM 962104, locality Anda3.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F95EC4C2AB39" blockId="5.[151,1436,1373,2008]" box="[151,750,1772,1794]" pageId="5" pageNumber="406">
10.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF95EC787AB39" box="[189,427,1772,1794]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF2BF95EC71DAB39" box="[189,305,1772,1794]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDEFFFCFEC1F95EC787AB39" box="[343,427,1772,1794]" italics="true" pageId="5" pageNumber="406">muelleri</emphasis>
</taxonomicName>
, RGM 962105, locality Anda3.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F8B8C4C1AB1B" blockId="5.[151,1436,1373,2008]" box="[151,749,1802,1824]" pageId="5" pageNumber="406">
11.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2AF8B8C786AB1B" box="[188,426,1802,1824]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF2AF8B8C71CAB1B" box="[188,304,1802,1824]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDEFFFCFEC0F8B8C786AB1B" box="[342,426,1802,1824]" italics="true" pageId="5" pageNumber="406">muelleri</emphasis>
</taxonomicName>
, RGM 962101, locality Anda1.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F89BC55FAB04" blockId="5.[151,1436,1373,2008]" box="[151,883,1833,1855]" pageId="5" pageNumber="406">
12.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF89BC787AB04" box="[189,427,1833,1855]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF2BF89BC71DAB04" box="[189,305,1833,1855]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDEFFFCFEC1F89BC787AB04" box="[343,427,1833,1855]" italics="true" pageId="5" pageNumber="406">muelleri</emphasis>
</taxonomicName>
, RGM 962100, locality Anda1, SEM image.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F8FAC4C2AB65" blockId="5.[151,1436,1373,2008]" box="[151,750,1864,1886]" pageId="5" pageNumber="406">
13.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF8FAC787AB65" box="[189,427,1864,1886]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDEFFFCFF2BF8FAC71DAB65" box="[189,305,1864,1886]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDEFFFCFEC1F8FAC787AB65" box="[343,427,1864,1886]" italics="true" pageId="5" pageNumber="406">muelleri</emphasis>
</taxonomicName>
, RGM 962103, locality Anda3.
</paragraph>
<paragraph id="4D17AC36FFDEFFFCFF01F8D4C4B9AB47" blockId="5.[151,1436,1373,2008]" box="[151,661,1894,1916]" pageId="5" pageNumber="406">
14.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF8D4C71DAB47" authorityName="Cocco" authorityYear="1838" box="[189,305,1894,1916]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF2BF8D4C71DAB47" box="[189,305,1894,1916]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
</taxonomicName>
sp., RGM 962109, locality Anda1.
</paragraph>
<footnote id="2EB3B038FFDEFFFCFF01F837C536ABA0" box="[151,794,1925,1947]" pageId="5" pageNumber="406">
<paragraph id="4D17AC36FFDEFFFCFF01F837C536ABA0" blockId="5.[151,1436,1373,2008]" box="[151,794,1925,1947]" pageId="5" pageNumber="406">
15.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF837C71DABA0" authorityName="Cocco" authorityYear="1838" box="[189,305,1925,1947]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF2BF837C71DABA0" box="[189,305,1925,1947]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
</taxonomicName>
sp., RGM 962107, locality Anda1, SEM image.
</paragraph>
</footnote>
<footnote id="2EB3B038FFDEFFFCFF01F816C4BDAB81" box="[151,657,1956,1978]" pageId="5" pageNumber="406">
<paragraph id="4D17AC36FFDEFFFCFF01F816C4BDAB81" blockId="5.[151,1436,1373,2008]" box="[151,657,1956,1978]" pageId="5" pageNumber="406">
16.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF816C71DAB81" authorityName="Cocco" authorityYear="1838" box="[189,305,1956,1978]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF2BF816C71DAB81" box="[189,305,1956,1978]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
</taxonomicName>
sp., RGM 962112, locality Roxas.
</paragraph>
</footnote>
<footnote id="2EB3B038FFDEFFFCFF01F870C4A7ABE3" box="[151,651,1986,2008]" pageId="5" pageNumber="406">
<paragraph id="4D17AC36FFDEFFFCFF01F870C4A7ABE3" blockId="5.[151,1436,1373,2008]" box="[151,651,1986,2008]" pageId="5" pageNumber="406">
17.
<taxonomicName id="8AA8D7B5FFDEFFFCFF2BF870C71DABE3" authorityName="Cocco" authorityYear="1838" box="[189,305,1986,2008]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="5" pageNumber="406" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDEFFFCFF2BF870C71DABE3" box="[189,305,1986,2008]" italics="true" pageId="5" pageNumber="406">Maurolicus</emphasis>
</taxonomicName>
sp., RGM 962115, locality Tiep2.
</paragraph>
</footnote>
<paragraph id="4D17AC36FFDDFFFFFF01FF25C441ACC3" blockId="6.[151,1437,151,1040]" pageId="6" pageNumber="407">
Material:
<specimenCount id="5BAE67BFFFDDFFFFFE93FF2AC7B1AC8B" box="[261,413,151,176]" pageId="6" pageNumber="407" type="generic">24 specimens</specimenCount>
in total. Anda1 (2) RGM 962100,
<materialsCitation id="FDC0A66BFFDDFFFFFCB0FF2AC5EBAC8B" ID-GBIF-Occurrence="2013952855" box="[806,967,151,176]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962101" specimenCount="1">
<specimenCode id="1D0E044DFFDDFFFFFCB0FF2AC5EBAC8B" box="[806,967,151,176]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962101</specimenCode>
</materialsCitation>
,;
<materialsCitation id="FDC0A66BFFDDFFFFFC41FF25C2D8AC8B" ID-GBIF-Occurrence="2013952865" box="[983,1268,151,176]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962102" specimenCount="1">
Anda2 (7)
<specimenCode id="1D0E044DFFDDFFFFFBC4FF2AC2DCAC8B" box="[1106,1264,151,176]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962102</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFDDFFFFFB6DFF2AC3B4AC8B" ID-GBIF-Occurrence="2013952853" box="[1275,1432,151,176]" collectionCode="RGM" isTailEnumeration="true" pageId="6" pageNumber="407" specimenCode="RGM 962111" specimenCount="1">
<specimenCode id="1D0E044DFFDDFFFFFB6DFF2AC3B4AC8B" box="[1275,1432,151,176]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962111</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFDDFFFFFF01FF0EC79BACEE" ID-GBIF-Occurrence="2013952856" box="[151,439,188,213]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962103" specimenCount="1">
Anda3 (4)
<specimenCode id="1D0E044DFFDDFFFFFE82FF0EC798ACEE" box="[276,436,188,213]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962103</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFDDFFFFFE56FF0EC448ACEE" ID-GBIF-Occurrence="2013952860" box="[448,612,188,213]" collectionCode="RGM" isTailEnumeration="true" pageId="6" pageNumber="407" specimenCode="RGM 962104" specimenCount="1">
<specimenCode id="1D0E044DFFDDFFFFFE56FF0EC44CACEE" box="[448,608,188,213]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962104</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFDDFFFFFDFBFF0EC53DACEE" ID-GBIF-Occurrence="2013952874" box="[621,785,188,213]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962105" specimenCount="1">
<specimenCode id="1D0E044DFFDDFFFFFDFBFF0EC521ACEE" box="[621,781,188,213]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962105</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFDDFFFFFC8CFF0EC596ACEE" ID-GBIF-Occurrence="2013952848" box="[794,954,188,213]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962106" specimenCount="1">
<specimenCode id="1D0E044DFFDDFFFFFC8CFF0EC596ACEE" box="[794,954,188,213]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962106</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFDDFFFFFC51FF0EC30CACEE" ID-GBIF-Occurrence="2013952862" box="[967,1312,188,213]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962114" specimenCount="1">
AndaDeVos (2)
<specimenCode id="1D0E044DFFDDFFFFFB16FF0EC30CACEE" box="[1152,1312,188,213]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962114</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFDDFFFFFABBFF0FC718ACC3" ID-GBIF-Occurrence="2013952842" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962113" specimenCount="1">
Roxas (8)
<specimenCode id="1D0E044DFFDDFFFFFF01FF52C718ACC3" box="[151,308,223,248]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962113</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFDDFFFFFEA9FF6DC445ACC3" ID-GBIF-Occurrence="2013952867" box="[319,617,223,248]" collectionCode="RGM" pageId="6" pageNumber="407" specimenCode="RGM 962116" specimenCount="1">
Tiep2 (1) in
<specimenCode id="1D0E044DFFDDFFFFFE5DFF52C445ACC3" box="[459,617,223,248]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="407" type="Museum">RGM 962116</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4D17AC36FFDDFFFFFF51FEB6C520AE93" blockId="6.[151,1437,151,1040]" pageId="6" pageNumber="407">
Small otoliths (OL:OH=1.19–1.35) with a long pointed rostrum a sharp to rectangular postdorsal angle that separates two rather straight parts of the dorsal rim, of which the posterior one is considerably steeper than the anterior one. The anterior part of the dorsal rim starts at 1/3 of the sulcus, leaving the upper part of the rostrum covered by the ostium. The ventral rim is separated in two parts by a pointed postventral angle, which is accentuated by a prominent flag-like extension. The posterior part of the dorsal rim and that of the ventral rim form a variably rounded angle that can be prominent or shallow. The ventral rim contains irregular lobes; the dorsal rim varies in ornamentation from smooth to lobed. The slightly supramedian sulcus is divided in a deep ostium and a longer cauda that ends at considerable distance from the posterior rim. A crista superior lines the cauda only. The crista inferior consists of two separated parts that line either from ostium to the anterior part of the cauda or the posterior part of the cauda. The outer surface is smooth with a marked uprising at the central part. These otoliths are reminiscent of the extant
<taxonomicName id="8AA8D7B5FFDDFFFFFE7CFDDEC4FDAEBE" box="[490,721,620,645]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFE7CFDDEC4FDAEBE" box="[490,721,620,645]" italics="true" pageId="6" pageNumber="407">Maurolicus muelleri</emphasis>
</taxonomicName>
, which is also known from Pliocene and Miocene deposits in Europe (
<bibRefCitation id="2939D1C7FFDDFFFFFF60FD3DC42DAE93" author="Nolf, D. &amp; Capetta, H." box="[246,513,655,680]" pageId="6" pageNumber="407" pagination="209 - 271" refId="ref19426" refString="Nolf, D. &amp; Capetta, H. (1989) Otolithes de poisons pliocenes du Sud-Est de la France. Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 58, 209 - 271." type="journal article" year="1989">Nolf and Capetta, 1989</bibRefCitation>
; Steurbaut
<emphasis id="7FDC7024FFDDFFFFFDE8FD23C483AE93" box="[638,687,655,680]" italics="true" pageId="6" pageNumber="407">et al</emphasis>
. 1979).
</paragraph>
<paragraph id="4D17AC36FFDDFFFFFF51FD06C78AA834" blockId="6.[151,1437,151,1040]" pageId="6" pageNumber="407">
At present 15 species of
<taxonomicName id="8AA8D7B5FFDDFFFFFE7EFD06C444AEF6" authorityName="Cocco" authorityYear="1838" box="[488,616,692,717]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDDFFFFFE7EFD06C444AEF6" box="[488,616,692,717]" italics="true" pageId="6" pageNumber="407">Maurolicus</emphasis>
</taxonomicName>
are known, of which
<taxonomicName id="8AA8D7B5FFDDFFFFFCFCFD07C5DCAEF6" box="[874,1008,692,717]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFCFCFD07C5DCAEF6" box="[874,1008,692,717]" italics="true" pageId="6" pageNumber="407">M. muelleri</emphasis>
</taxonomicName>
and
<taxonomicName id="8AA8D7B5FFDDFFFFFBBDFD07C292AEF7" authorityName="Parin &amp; Kobyliansky" authorityYear="1993" box="[1067,1214,693,716]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="javanicus">
<emphasis id="7FDC7024FFDDFFFFFBBDFD07C292AEF7" box="[1067,1214,693,716]" italics="true" pageId="6" pageNumber="407">M. javanicus</emphasis>
</taxonomicName>
are encountered in deep West Central Pacific waters, and
<taxonomicName id="8AA8D7B5FFDDFFFFFDD3FD6BC4F4AECB" authorityName="Ishikawa" authorityYear="1915" box="[581,728,729,752]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="japonicus">
<emphasis id="7FDC7024FFDDFFFFFDD3FD6BC4F4AECB" box="[581,728,729,752]" italics="true" pageId="6" pageNumber="407">M. japonicus</emphasis>
</taxonomicName>
is known from the North West Pacific,
<collectingCountry id="35BFECA6FFDDFFFFFB03FD6AC2F4AED4" box="[1173,1240,728,751]" name="Japan" pageId="6" pageNumber="407">Japan</collectingCountry>
, and Hawaii. The other
<taxonomicName id="8AA8D7B5FFDDFFFFFF4FFD4EC775AF2E" authorityName="Cocco" authorityYear="1838" box="[217,345,764,789]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDDFFFFFF4FFD4EC775AF2E" box="[217,345,764,789]" italics="true" pageId="6" pageNumber="407">Maurolicus</emphasis>
</taxonomicName>
species are only known from areas remote of the
<collectingCountry id="35BFECA6FFDDFFFFFC0BFD4EC233AF2E" box="[925,1055,764,789]" name="Philippines" pageId="6" pageNumber="407">Philippines</collectingCountry>
. However, many of these species have only been recognized from 1993, when Parin and Kobyliansky (1993) reported that the known pan-global
<taxonomicName id="8AA8D7B5FFDDFFFFFAEBFC93C6D8AF66" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFAEBFC93C6D8AF66" italics="true" pageId="6" pageNumber="407">M. muelleri</emphasis>
</taxonomicName>
represented a mixture of different species. While this asks for caution in using old references to
<taxonomicName id="8AA8D7B5FFDDFFFFFAEBFCF7C6D8AFBB" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFAEBFCF7C6D8AFBB" italics="true" pageId="6" pageNumber="407">M. muelleri</emphasis>
</taxonomicName>
, the presence of
<taxonomicName id="8AA8D7B5FFDDFFFFFE51FCDBC463AFBB" box="[455,591,871,896]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFE51FCDBC463AFBB" box="[455,591,871,896]" italics="true" pageId="6" pageNumber="407">M. muelleri</emphasis>
</taxonomicName>
is well documented from the Mediterranean Sea, from which no other
<taxonomicName id="8AA8D7B5FFDDFFFFFF01FC3EC73BAF9E" authorityName="Cocco" authorityYear="1838" box="[151,279,908,933]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFDDFFFFFF01FC3EC73BAF9E" box="[151,279,908,933]" italics="true" pageId="6" pageNumber="407">Maurolicus</emphasis>
</taxonomicName>
species have been reported (
<bibRefCitation id="2939D1C7FFDDFFFFFDFAFC3EC51BAF9F" author="Tuset, V. M. &amp; Lombarte, A. &amp; Assis, C. A." box="[620,823,908,933]" pageId="6" pageNumber="407" pagination="7 - 171" refId="ref20081" refString="Tuset, V. M., Lombarte, A. &amp; Assis, C. A. (2008) Otolith atlas for the western Mediterranean, north and central eastern Atlantic. Scientia Marina, Barcelona, 72 (Supplement 1), 7 - 171." type="journal article" year="2008">
Tuset
<emphasis id="7FDC7024FFDDFFFFFD20FC3FC4C7AF9E" box="[694,747,908,933]" italics="true" pageId="6" pageNumber="407">et al</emphasis>
, 2008
</bibRefCitation>
). Comparison of the published data on
<taxonomicName id="8AA8D7B5FFDDFFFFFA83FC3FC3B0AF9E" box="[1301,1436,908,933]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFA83FC3FC3B0AF9E" box="[1301,1436,908,933]" italics="true" pageId="6" pageNumber="407">M. muelleri</emphasis>
</taxonomicName>
otoliths provide a consistent picture. Several of our specimens (Figures 9–13) are identical to
<taxonomicName id="8AA8D7B5FFDDFFFFFB30FC03C30BAFF3" box="[1190,1319,943,968]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFB30FC03C30BAFF3" box="[1190,1319,943,968]" italics="true" pageId="6" pageNumber="407">M. muelleri</emphasis>
</taxonomicName>
, while the ornamentation of a few specimens is rather pronounced. As they appear as one species, we have indicated it as
<taxonomicName id="8AA8D7B5FFDDFFFFFF01FC45C7B3A82B" box="[151,415,1015,1040]" class="Actinopterygii" family="Sternoptychidae" genus="Maurolicus" kingdom="Animalia" order="Stomiiformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7FDC7024FFDDFFFFFF01FC45C73BA82B" box="[151,279,1015,1040]" italics="true" pageId="6" pageNumber="407">Maurolicus</emphasis>
cf.
<emphasis id="7FDC7024FFDDFFFFFED4FC45C7B3A82B" box="[322,415,1015,1040]" italics="true" pageId="6" pageNumber="407">muelleri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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