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<document id="D198FE698B9985F90D2C6882914648C2" ID-CLB-Dataset="28139" ID-DOI="10.11646/zootaxa.4563.3.1" ID-GBIF-Dataset="cf7fa86d-ae34-4e8d-aa9d-7c15568bc995" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601295" ID-ZooBank="B0A3408F-563A-4DD3-94A4-284A2770B0A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171996401" checkinUser="plazi" docAuthor="Van Hinsbergh, Victor W. M. &amp; Helwerda, Renate A." docDate="2019" docId="C5011D20FFD1FFF3FF01FF25C53EAFE0" docLanguage="en" docName="zootaxa.4563.3.1.pdf" docOrigin="Zootaxa 4563 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benthosema fibulatum" docType="treatment" docVersion="10" lastPageNumber="411" masterDocId="39386558FFDBFFF9FF96FFB2C62CAC3B" masterDocTitle="Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines" masterLastPageNumber="443" masterPageNumber="401" pageNumber="411" updateTime="1698703400501" updateUser="ExternalLinkService">
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<mods:title id="1FF37B400C0BA11E6B97606E059EC859">Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines</mods:title>
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<mods:namePart id="AC983C98316AC87CF786833A1B62EF26">Van Hinsbergh, Victor W. M.</mods:namePart>
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<mods:namePart id="82D732FBC9FB90FE810F539A7A77606A">Helwerda, Renate A.</mods:namePart>
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<mods:number id="BFE03C5C86A66D66B38BF5EAAC2DE436">4563</mods:number>
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<mods:identifier id="19ED469AA140529F139D9D6EBF523116" type="DOI">10.11646/zootaxa.4563.3.1</mods:identifier>
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<treatment id="C5011D20FFD1FFF3FF01FF25C53EAFE0" ID-DOI="http://doi.org/10.5281/zenodo.5937011" ID-GBIF-Taxon="154474775" ID-Zenodo-Dep="5937011" LSID="urn:lsid:plazi:treatment:C5011D20FFD1FFF3FF01FF25C53EAFE0" httpUri="http://treatment.plazi.org/id/C5011D20FFD1FFF3FF01FF25C53EAFE0" lastPageNumber="411" pageId="10" pageNumber="411">
<subSubSection id="05B2FFBDFFD1FFF3FF01FF25C7BEACE8" pageId="10" pageNumber="411" type="nomenclature">
<paragraph id="4D17AC36FFD1FFF3FF01FF25C4DDAC89" blockId="10.[151,753,151,211]" box="[151,753,151,178]" pageId="10" pageNumber="411">
<heading id="165F1B5AFFD1FFF3FF01FF25C4DDAC89" bold="true" box="[151,753,151,178]" fontSize="11" level="1" pageId="10" pageNumber="411" reason="1">
<taxonomicName id="8AA8D7B5FFD1FFF3FF01FF25C4DDAC89" ID-CoL="LHZJ" authority="(Gilbert &amp; Cramer, 1897)" baseAuthorityName="Gilbert &amp; Cramer" baseAuthorityYear="1897" box="[151,753,151,178]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="fibulatum">
<emphasis id="7FDC7024FFD1FFF3FF01FF25C4DDAC89" bold="true" box="[151,753,151,178]" pageId="10" pageNumber="411">
<emphasis id="7FDC7024FFD1FFF3FF01FF25C787AC8A" bold="true" box="[151,427,151,177]" italics="true" pageId="10" pageNumber="411">Benthosema fibulatum</emphasis>
(Gilbert &amp; Cramer, 1897)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4D17AC36FFD1FFF3FF01FF08C7BEACE8" blockId="10.[151,753,151,211]" box="[151,402,186,211]" pageId="10" pageNumber="411">(Figure 27 and 36–38)</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFD1FFF3FF01FEB0C53EAFE0" pageId="10" pageNumber="411" type="description">
<paragraph id="4D17AC36FFD1FFF3FF01FEB0C529AD23" blockId="10.[151,1123,258,342]" box="[151,773,258,280]" pageId="10" pageNumber="411">
1980
<taxonomicName id="8AA8D7B5FFD1FFF3FF65FEB0C4ACAD23" authority="Schwarzhans W" authorityName="Schwarzhans W" box="[243,640,258,280]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="fibulatum">
<emphasis id="7FDC7024FFD1FFF3FF65FEB0C7FAAD23" box="[243,470,258,280]" italics="true" pageId="10" pageNumber="411">Benthosema fibulatum</emphasis>
Schwarzhans W
</taxonomicName>
, p.42, fig. 89
</paragraph>
<paragraph id="4D17AC36FFD1FFF3FF01FE93C24FAD0C" blockId="10.[151,1123,258,342]" box="[151,1123,289,311]" pageId="10" pageNumber="411">
1995
<taxonomicName id="8AA8D7B5FFD1FFF3FF65FE93C437AD0C" authority="Smale" authorityName="Smale" box="[243,539,289,311]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="fibulatum">
<emphasis id="7FDC7024FFD1FFF3FF65FE93C7FAAD0C" box="[243,470,289,311]" italics="true" pageId="10" pageNumber="411">Benthosema fibulatum</emphasis>
Smale
</taxonomicName>
MJ, Watson G, Hecht T, pp. 41–42, plate 16, fig. G1–G3.
</paragraph>
<paragraph id="4D17AC36FFD1FFF3FF01FEF2C56EAD6D" blockId="10.[151,1123,258,342]" box="[151,834,320,342]" pageId="10" pageNumber="411">
1999
<taxonomicName id="8AA8D7B5FFD1FFF3FF65FEF2C4B6AD6D" authority="Rivaton &amp; Bourret" authorityName="Rivaton &amp; Bourret" box="[243,666,320,342]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="fibulatum">
<emphasis id="7FDC7024FFD1FFF3FF65FEF2C7FAAD6D" box="[243,470,320,342]" italics="true" pageId="10" pageNumber="411">Benthosema fibulatum</emphasis>
Rivaton &amp; Bourret
</taxonomicName>
pl. 105, fig. 1–2
</paragraph>
<paragraph id="4D17AC36FFD1FFF3FF01FE30C7B7AD84" blockId="10.[151,1436,386,987]" pageId="10" pageNumber="411">
Material:
<specimenCount id="5BAE67BFFFD1FFF3FE9DFE31C785ADA0" box="[267,425,386,411]" pageId="10" pageNumber="411" type="generic">30 specimens</specimenCount>
in total. Roxas (3) RGM 962171, 962172, 962349;
<materialsCitation id="FDC0A66BFFD1FFF3FBB3FE30C6DEAD84" ID-GBIF-Occurrence="2013952787" collectionCode="RGM" pageId="10" pageNumber="411" specimenCode="RGM 962345" specimenCount="1">
AndaDeVos (27) 962348,
<specimenCode id="1D0E044DFFD1FFF3FACAFE31C6C2AD84" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="411" type="Museum">RGM 962345</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFD1FFF3FF6FFE15C7B4AD84" ID-GBIF-Occurrence="2013952778" box="[249,408,422,447]" collectionCode="RGM" isTailEnumeration="true" pageId="10" pageNumber="411" specimenCode="RGM 962346" specimenCount="1">
<specimenCode id="1D0E044DFFD1FFF3FF6FFE15C7B4AD84" box="[249,408,422,447]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="411" type="Museum">RGM 962346</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4D17AC36FFD1FFF3FF51FE79C420AF1D" blockId="10.[151,1436,386,987]" pageId="10" pageNumber="411">
One well-conserved adult oval-shaped Myctophid specimen and
<specimenCount id="5BAE67BFFFD1FFF3FC23FE79C219ADD8" box="[949,1077,458,483]" pageId="10" pageNumber="411" type="juvenile">29 juvenile</specimenCount>
specimens were recognized as
<taxonomicName id="8AA8D7B5FFD1FFF3FF01FE5CC7A9AE3C" box="[151,389,494,519]" class="Actinopterygii" family="Myctophidae" genus="Bethosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="fibulatum">
<emphasis id="7FDC7024FFD1FFF3FF01FE5CC7A9AE3C" box="[151,389,494,519]" italics="true" pageId="10" pageNumber="411">Bethosema fibulatum</emphasis>
</taxonomicName>
. The adult specimen is characterized by a
<taxonomicName id="8AA8D7B5FFD1FFF3FCFBFE5CC5DAAE3C" authorityName="Goode &amp; Bean" authorityYear="1896" box="[877,1014,494,519]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFD1FFF3FCFBFE5CC5DAAE3C" box="[877,1014,494,519]" italics="true" pageId="10" pageNumber="411">Benthosema</emphasis>
</taxonomicName>
-characteristic ventral rim consisting of a blunt massive denticle at the end of the posterior part and 4–6 poorly developed rounded denticles that are restricted to the anterior part of the ventral rim. The dorsal rim is ornamented with regular broad crenulations; it is rather regularly curved with a feeble postdorsal angle and a vertical posterior part. The specimen (OL:OH ratio=1.26) has a OCL/CCL ratio amounting 1.92. A bluntly pointed rostrum exceeds a short antirostrum. There is a distinct but short excisural notch. The inner surface is slightly convex. The outer surface shows ridges and furrows along the ventral and dorsal rims that run perpendicular to the rims, but do not reach the central part. The posterior part of the outer surface is convex and slightly thicker than the anterior part; the transition from anterior to posterior part is slightly concave.
</paragraph>
<paragraph id="4D17AC36FFD1FFF3FF51FC80C53EAFE0" blockId="10.[151,1436,386,987]" pageId="10" pageNumber="411">
The ventral rim with its robust posterior-ventral denticle is characteristic for otoliths of the genus
<taxonomicName id="8AA8D7B5FFD1FFF3FA9AFC80C3B9AF70" authorityName="Goode &amp; Bean" authorityYear="1896" box="[1292,1429,818,843]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFD1FFF3FA9AFC80C3B9AF70" box="[1292,1429,818,843]" italics="true" pageId="10" pageNumber="411">Benthosema</emphasis>
</taxonomicName>
. The relatively high L:H ratio (1.26) and the high OCL/CCL ratio (1.92) fit with the extant
<taxonomicName id="8AA8D7B5FFD1FFF3FB12FCE4C352AF54" box="[1156,1406,854,879]" class="Actinopterygii" family="Myctophidae" genus="Benthosema" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="fibulatum">
<emphasis id="7FDC7024FFD1FFF3FB12FCE4C352AF54" box="[1156,1406,854,879]" italics="true" pageId="10" pageNumber="411">Benthosema fibulatum</emphasis>
</taxonomicName>
as reported by
<bibRefCitation id="2939D1C7FFD1FFF3FEB6FCC8C468AFA8" author="Rivaton, J. &amp; Bourret, P." box="[288,580,890,915]" pageId="10" pageNumber="411" refId="ref19612" refString="Rivaton, J. &amp; Bourret, P. (1999) Otoliths of the Indo-Pacific fishes. Documents Scientifiques et Techniques, Centre IRD de Noumea, Noumea, 378 pp." type="book" year="1999">Rivaton &amp; Bourett (1999)</bibRefCitation>
from the
<collectingCountry id="35BFECA6FFD1FFF3FD23FCC8C51FAFA8" box="[693,819,890,915]" name="Philippines" pageId="10" pageNumber="411">Philippines</collectingCountry>
environment. Together with the shape of the mid- and posterior dorsal rim and the typical dentition along the ventral rim this species discriminates from
<taxonomicName id="8AA8D7B5FFD1FFF3FA87FC2CC79DAFE0" authority="Taning, 1932" authorityName="Taning" authorityYear="1932" family="Myctophidae" genus="Benthosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="panamense">
<emphasis id="7FDC7024FFD1FFF3FA87FC2CC739AFE0" italics="true" pageId="10" pageNumber="411">Benthosoma panamense</emphasis>
Tåning, 1932
</taxonomicName>
and other
<taxonomicName id="8AA8D7B5FFD1FFF3FDBEFC70C49FAFE0" box="[552,691,962,987]" family="Myctophidae" genus="Benthosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Myctophiformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFD1FFF3FDBEFC70C49FAFE0" box="[552,691,962,987]" italics="true" pageId="10" pageNumber="411">Benthosoma</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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