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<document id="F77E7F7EA4074A344FB02E374AB335AB" ID-CLB-Dataset="28139" ID-DOI="10.11646/zootaxa.4563.3.1" ID-GBIF-Dataset="cf7fa86d-ae34-4e8d-aa9d-7c15568bc995" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601295" ID-ZooBank="B0A3408F-563A-4DD3-94A4-284A2770B0A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171996401" checkinUser="plazi" docAuthor="Van Hinsbergh, Victor W. M. &amp; Helwerda, Renate A." docDate="2019" docId="C5011D20FFCFFFECFF01FB6DC5BEACEE" docLanguage="en" docName="zootaxa.4563.3.1.pdf" docOrigin="Zootaxa 4563 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hymenocephalus aeger Gilbert &amp; Hubbs 1920" docType="treatment" docVersion="9" lastPageNumber="421" masterDocId="39386558FFDBFFF9FF96FFB2C62CAC3B" masterDocTitle="Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines" masterLastPageNumber="443" masterPageNumber="401" pageNumber="421" updateTime="1698703400501" updateUser="ExternalLinkService">
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<mods:title id="FFFB16CE259AF2BE8854E648AA40BC25">Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines</mods:title>
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<mods:namePart id="90E1F37E126B8346E4B17568CE3EADEB">Van Hinsbergh, Victor W. M.</mods:namePart>
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<mods:namePart id="51004DB021A7FDC22A2CDEC7AE5ED08F">Helwerda, Renate A.</mods:namePart>
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<mods:identifier id="82F7CCAB63BA9F6C2C378ACB9B94C8F4" type="DOI">10.11646/zootaxa.4563.3.1</mods:identifier>
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<treatment id="C5011D20FFCFFFECFF01FB6DC5BEACEE" ID-DOI="http://doi.org/10.5281/zenodo.5937049" ID-GBIF-Taxon="154474753" ID-Zenodo-Dep="5937049" LSID="urn:lsid:plazi:treatment:C5011D20FFCFFFECFF01FB6DC5BEACEE" httpUri="http://treatment.plazi.org/id/C5011D20FFCFFFECFF01FB6DC5BEACEE" lastPageId="21" lastPageNumber="421" pageId="20" pageNumber="421">
<subSubSection id="05B2FFBDFFCFFFEDFF01FB6DC766A920" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="4D17AC36FFCFFFEDFF01FB6DC4F5A8C1" blockId="20.[151,729,1247,1307]" box="[151,729,1247,1274]" pageId="20" pageNumber="421">
<heading id="165F1B5AFFCFFFEDFF01FB6DC4F5A8C1" bold="true" box="[151,729,1247,1274]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<taxonomicName id="8AA8D7B5FFCFFFEDFF01FB6DC4F5A8C1" ID-CoL="3NJ8K" authority="Gilbert &amp; Hubbs, 1920" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[151,729,1247,1274]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="species" species="aeger">
<emphasis id="7FDC7024FFCFFFEDFF01FB6DC4F5A8C1" bold="true" box="[151,729,1247,1274]" pageId="20" pageNumber="421">
<emphasis id="7FDC7024FFCFFFEDFF01FB6DC79FA8C2" bold="true" box="[151,435,1247,1273]" italics="true" pageId="20" pageNumber="421">Hymenocephalus aeger</emphasis>
Gilbert &amp; Hubbs, 1920
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="4D17AC36FFCFFFEDFF01FAB0C766A920" blockId="20.[151,729,1247,1307]" box="[151,330,1282,1307]" pageId="20" pageNumber="421">(Figures 68–69)</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFCFFFECFF01FAF8C5BEACEE" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="description">
<paragraph id="4D17AC36FFCFFFEDFF01FAF8C508A95B" blockId="20.[151,804,1354,1376]" box="[151,804,1354,1376]" pageId="20" pageNumber="421">
2014
<taxonomicName id="8AA8D7B5FFCFFFEDFF46FAF8C752A95B" authorityName="Gigioli" authorityYear="1884" box="[208,382,1354,1376]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFCFFFEDFF46FAF8C752A95B" box="[208,382,1354,1376]" italics="true" pageId="20" pageNumber="421">Hymenocephalus</emphasis>
</taxonomicName>
<taxonomicName id="8AA8D7B5FFCFFFEDFE12FAF9C449A95B" authority="Schwarzhans W" authorityName="Schwarzhans W" box="[388,613,1354,1376]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="species" species="aeger">aeger Schwarzhans W</taxonomicName>
, pp. 58–59, fig. 30
</paragraph>
<paragraph id="4D17AC36FFCFFFEDFF01FA3FC455A9F1" blockId="20.[151,1436,1421,2022]" pageId="20" pageNumber="421">
Material:
<specimenCount id="5BAE67BFFFCFFFEDFE93FA3CC7A3A99D" box="[261,399,1421,1446]" pageId="20" pageNumber="421" type="generic">7 specimens</specimenCount>
in total. Anda1 (1) RGM 962232;
<materialsCitation id="FDC0A66BFFCFFFEDFC83FA3FC240A99D" ID-GBIF-Occurrence="2013952781" box="[789,1132,1421,1446]" collectionCode="RGM" pageId="20" pageNumber="421" specimenCode="RGM 962233" specimenCount="1">
AndaDeVos (3)
<specimenCode id="1D0E044DFFCFFFEDFC5CFA3CC245A99D" box="[970,1129,1421,1446]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="421" type="Museum">RGM 962233</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFCFFFEDFBE5FA3CC33EA99D" ID-GBIF-Occurrence="2013952805" box="[1139,1298,1421,1446]" collectionCode="RGM" isTailEnumeration="true" pageId="20" pageNumber="421" specimenCode="RGM 962234" specimenCount="1">
<specimenCode id="1D0E044DFFCFFFEDFBE5FA3CC33EA99D" box="[1139,1298,1421,1446]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="421" type="Museum">RGM 962234</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFCFFFEDFA8BFA3FC771A9F1" ID-GBIF-Occurrence="2013952783" collectionCode="RGM" pageId="20" pageNumber="421" specimenCode="RGM 962235" specimenCount="1">
AndaCliff3 (1)
<specimenCode id="1D0E044DFFCFFFEDFF28FA00C771A9F1" box="[190,349,1457,1482]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="421" type="Museum">RGM 962235</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFCFFFEDFEFEFA03C45AA9F1" ID-GBIF-Occurrence="2013952789" box="[360,630,1457,1482]" collectionCode="RGM" pageId="20" pageNumber="421" specimenCode="RGM 962236" specimenCount="1">
Tiep1 (2)
<specimenCode id="1D0E044DFFCFFFEDFE41FA00C45AA9F1" box="[471,630,1457,1482]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="421" type="Museum">RGM 962236</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4D17AC36FFCFFFEDFF51FA64C4C6AB35" blockId="20.[151,1436,1421,2022]" pageId="20" pageNumber="421">
Two well-preserved specimens (OL:OH= 0.93–1.00) comply well with the characteristics of members of the genus
<taxonomicName id="8AA8D7B5FFCFFFEDFF77FA4BC78EAA29" authorityName="Gigioli" authorityYear="1884" box="[225,418,1529,1554]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFCFFFEDFF77FA4BC78EAA29" box="[225,418,1529,1554]" italics="true" pageId="20" pageNumber="421">Hymenocephalus</emphasis>
</taxonomicName>
that have a sulcus with a fused colliculum. The dorsal rim has a large moderately broad predorsal lobe that is bordered by two straight and parallel sides and a curved upper rim with four insignificant lobules; accentuated by a small concavity the posterior part runs gradually down towards a posterior point just above a posterior depression of the sulcus. The sulcus is slightly bent, in particular because its posterior part slightly declines to the ventral side. The shape of the colliculum differs between the two specimens. One has a fully fused colliculum with a thin extension towards but not reaching the anterior rim; the fused colliculum of the other specimen still displays that it is a fusion of two collicula. Under the colliculum, both specimens show a large pseudocolliculum. A ventral furrow is hardly visible.
</paragraph>
<paragraph id="4D17AC36FFCFFFECFF51F8ABC5BEACEE" blockId="20.[151,1436,1421,2022]" lastBlockId="21.[151,1436,151,213]" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
The posterior bluntly pointed part is slightly stronger in the largest specimen, which also has the fully fused colliculum. Notwithstanding, we consider the otoliths conspecific, because of the identity in the shape of the predorsal extension and characteristic shape of the sulcus.
<taxonomicName id="8AA8D7B5FFCFFFEDFCD0F8D3C22BAB41" authorityName="Gigioli" authorityYear="1884" box="[838,1031,1889,1914]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFCFFFEDFCD0F8D3C22BAB41" box="[838,1031,1889,1914]" italics="true" pageId="20" pageNumber="421">Hymenocephalus</emphasis>
</taxonomicName>
is encountered in the Indo-Pacific marine milieu (
<bibRefCitation id="2939D1C7FFCFFFEDFED3F837C40BABA5" author="Schwarzhans, W." box="[325,551,1925,1950]" pageId="20" pageNumber="421" pagination="1 - 73" refId="ref19846" refString="Schwarzhans, W. (2014) Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species. Zootaxa, 3888 (1), 1 - 73. http: // dx. doi. org / 10.11646 / zootaxa. 3888.1.1" type="journal article" year="2014">Schwarzhans, 2014</bibRefCitation>
). The specimens have a high similarity to the extant species
<taxonomicName id="8AA8D7B5FFCFFFEDFB4DF837C6FAABFA" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="species" species="aeger">
<emphasis id="7FDC7024FFCFFFEDFB4DF837C6FAABFA" italics="true" pageId="20" pageNumber="421">Hymenocephalus aeger</emphasis>
</taxonomicName>
, to which species we consider them to belong. They are reminiscent of
<taxonomicName id="8AA8D7B5FFCFFFEDFB9CF818C31CABF9" authority="Giglioni, 1884" authorityName="Giglioni" authorityYear="1884" box="[1034,1328,1961,1986]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="species" species="italicus">
<emphasis id="7FDC7024FFCFFFEDFB9CF818C2ACABF9" box="[1034,1152,1961,1986]" italics="true" pageId="20" pageNumber="421">H. italicus</emphasis>
Giglioni, 1884
</taxonomicName>
, which is known from the Atlantic Ocean and Mediterranean Sea, and
<taxonomicName id="8AA8D7B5FFCFFFEDFC1EF87CC31AABDD" authority="Smith &amp; Radcliffe, 1912" authorityName="Smith &amp; Radcliffe" authorityYear="1912" box="[904,1334,1997,2022]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="species" species="torvus">
<emphasis id="7FDC7024FFCFFFEDFC1EF87CC5D4ABDE" box="[904,1016,1998,2021]" italics="true" pageId="20" pageNumber="421">H. torvus</emphasis>
Smith &amp; Radcliffe, 1912
</taxonomicName>
, and
<taxonomicName id="8AA8D7B5FFCFFFECFA17F87CC709AC8B" baseAuthorityName="non Jordan &amp; Gilbert" baseAuthorityYear="1904" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" lastPageId="21" lastPageNumber="422" order="Gadiformes" pageId="20" pageNumber="421" phylum="Chordata" rank="species" species="striatissimus">
<emphasis id="7FDC7024FFCFFFECFA17F87CC709AC8B" italics="true" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">H. striatissimus</emphasis>
</taxonomicName>
<collectingCountry id="35BFECA6FFCEFFECFEBAFF25C75BAC8B" box="[300,375,151,176]" name="Jordan" pageId="21" pageNumber="422">Jordan</collectingCountry>
&amp; Gilbert, 1904, which dwell in
<collectingCountry id="35BFECA6FFCEFFECFD79FF25C541AC8B" box="[751,877,151,176]" name="Philippines" pageId="21" pageNumber="422">Philippines</collectingCountry>
and Japanese seas, but differ in OL:OH ratio, the shape and ornamentation of the dorsal extension, and curved sulcus.
</paragraph>
</subSubSection>
</treatment>
</document>
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