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<document id="F83F92F5C3E69C6F88C6E088E57CEC40" ID-CLB-Dataset="28139" ID-DOI="10.11646/zootaxa.4563.3.1" ID-GBIF-Dataset="cf7fa86d-ae34-4e8d-aa9d-7c15568bc995" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601295" ID-ZooBank="B0A3408F-563A-4DD3-94A4-284A2770B0A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171996401" checkinUser="plazi" docAuthor="Van Hinsbergh, Victor W. M. &amp; Helwerda, Renate A." docDate="2019" docId="C5011D20FFC5FFD9FF01FD7DC564AD96" docLanguage="en" docName="zootaxa.4563.3.1.pdf" docOrigin="Zootaxa 4563 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parascombrops schwarzhansi Van Hinsbergh &amp; Helwerda 2019, n.sp." docType="treatment" docVersion="10" lastPageNumber="433" masterDocId="39386558FFDBFFF9FF96FFB2C62CAC3B" masterDocTitle="Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines" masterLastPageNumber="443" masterPageNumber="401" pageNumber="431" updateTime="1698703400501" updateUser="ExternalLinkService">
<mods:mods id="156A37DD8900615C1D7EF2B253A067EF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E5573426DA42BC6C2D175E22A89840C9">
<mods:title id="BCC0CF76BE3200DC0E683D8A6148EC79">Fish Otoliths from the Cabarruyan Piacenzian-Gelasian fauna found in the Philippines</mods:title>
</mods:titleInfo>
<mods:name id="8F0BF06B1153F942AD0C31F278566C19" type="personal">
<mods:role id="B4475ABD85371ED039D47036FE65DA74">
<mods:roleTerm id="38833FD54DE1BD84921933F4917B0FE6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B58CF63F77490BB29763DAF9329EC68">Van Hinsbergh, Victor W. M.</mods:namePart>
</mods:name>
<mods:name id="19EDC7EE63699FCC68C03828964A989C" type="personal">
<mods:role id="78BF6167D9C4A6AD44369F30FE7D96F0">
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</mods:role>
<mods:namePart id="7811F970C4C230684B697D12253BCECA">Helwerda, Renate A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="20379AB33451C1AD62BCE8C2F5BB7584">text</mods:typeOfResource>
<mods:relatedItem id="6270338A37AB5B8AD6BB8243013BB42C" type="host">
<mods:titleInfo id="09960AFB9492726B8FFF760F3391BC24">
<mods:title id="B7C7C74762CFB3C0E3D10B72CD32FC5B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="279CD7ADAFB9061DB736F1DF727965F6">
<mods:date id="8D7B7A45B69CCDF9CE9C62CD2A6B328A">2019</mods:date>
<mods:detail id="6BBF4C61399F112ADD4F4AA49A69F752" type="pubDate">
<mods:number id="178EFE138436679A4764B96F5D4B28F6">2019-03-04</mods:number>
</mods:detail>
<mods:detail id="3F9D73B1054D43E18FC67261CDDA5395" type="volume">
<mods:number id="01345A863CAC02BD1B828F6CFDB13350">4563</mods:number>
</mods:detail>
<mods:detail id="7B30F810EC515C944A98D466DAE98987" type="issue">
<mods:number id="59E1C5611BB44872E5D07B86B8240364">3</mods:number>
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<mods:start id="1BC608714B0DB9FDFBCF8380FCDD4102">401</mods:start>
<mods:end id="63F5E09C32E5E696720AB7901BD2A98D">443</mods:end>
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<mods:classification id="90C7D702CB607A222C7E761AB99A3634">journal article</mods:classification>
<mods:identifier id="3FF25954E378F6DC61B7599835216EAB" type="CLB-Dataset">28139</mods:identifier>
<mods:identifier id="1AE0B3BDAFC9377025E6F7AD7B4D28AF" type="DOI">10.11646/zootaxa.4563.3.1</mods:identifier>
<mods:identifier id="234B1C2FFBA751E3B033CD295D8FACB8" type="GBIF-Dataset">cf7fa86d-ae34-4e8d-aa9d-7c15568bc995</mods:identifier>
<mods:identifier id="250147EC78E6808E965E326F949E398B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B500A1318BA6DED67F96AA4926419D71" type="Zenodo-Dep">2601295</mods:identifier>
<mods:identifier id="415D4D1A9C800B945E785B5A263FE442" type="ZooBank">B0A3408F-563A-4DD3-94A4-284A2770B0A6</mods:identifier>
</mods:mods>
<treatment id="C5011D20FFC5FFD9FF01FD7DC564AD96" ID-DOI="http://doi.org/10.5281/zenodo.5937093" ID-GBIF-Taxon="154474794" ID-Zenodo-Dep="5937093" LSID="urn:lsid:plazi:treatment:C5011D20FFC5FFD9FF01FD7DC564AD96" httpUri="http://treatment.plazi.org/id/C5011D20FFC5FFD9FF01FD7DC564AD96" lastPageId="32" lastPageNumber="433" pageId="30" pageNumber="431">
<subSubSection id="05B2FFBDFFC5FFE7FF01FD7DC74AAF30" pageId="30" pageNumber="431" type="nomenclature">
<paragraph id="4D17AC36FFC5FFE7FF01FD7DC46CAED1" blockId="30.[151,576,719,779]" box="[151,576,719,746]" pageId="30" pageNumber="431">
<heading id="165F1B5AFFC5FFE7FF01FD7DC46CAED1" bold="true" box="[151,576,719,746]" fontSize="11" level="1" pageId="30" pageNumber="431" reason="1">
<emphasis id="7FDC7024FFC5FFE7FF01FD7DC46CAED1" bold="true" box="[151,576,719,746]" pageId="30" pageNumber="431">
<taxonomicName id="8AA8D7B5FFC5FFE7FF01FD7DC7D2AED2" authority="Van Hinsbergh &amp; Helwerda, 2019" authorityName="Van Hinsbergh &amp; Helwerda" authorityYear="2019" box="[151,510,719,745]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="schwarzhansi" status="sp. nov.">
<emphasis id="7FDC7024FFC5FFE7FF01FD7DC7D2AED2" bold="true" box="[151,510,719,745]" italics="true" pageId="30" pageNumber="431">Parascombrops schwarzhansi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64EFCD5FFFC5FFE7FD93FD62C46CAED1" box="[517,576,720,746]" pageId="30" pageNumber="431" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01FD40C74AAF30" blockId="30.[151,576,719,779]" box="[151,358,754,779]" pageId="30" pageNumber="431">(Figures 102–107)</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFC5FFE7FF01FC88C507AFD8" pageId="30" pageNumber="431" type="materials_examined">
<paragraph id="4D17AC36FFC5FFE7FF01FC88C424AF68" blockId="30.[151,1436,826,1571]" box="[151,520,826,851]" pageId="30" pageNumber="431">
<materialsCitation id="FDC0A66BFFC5FFE7FF01FC88C424AF68" ID-GBIF-Occurrence="2013952731" box="[151,520,826,851]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962352" specimenCount="1" typeStatus="holotype">
<emphasis id="7FDC7024FFC5FFE7FF01FC88C721AF68" bold="true" box="[151,269,826,851]" pageId="30" pageNumber="431">
<typeStatus id="92131294FFC5FFE7FF01FC88C724AF68" box="[151,264,826,851]" pageId="30" pageNumber="431" type="holotype">Holotype</typeStatus>
:
</emphasis>
Anda2,
<specimenCode id="1D0E044DFFC5FFE7FEFBFC89C424AF68" box="[365,520,826,851]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962352</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF51FCECC784AFA0" blockId="30.[151,1436,826,1571]" pageId="30" pageNumber="431">
<materialsCitation id="FDC0A66BFFC5FFE7FF51FCECC53EAF4C" ID-GBIF-Occurrence="2013952726" box="[199,786,862,887]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962243" specimenCount="5" typeStatus="paratype">
<emphasis id="7FDC7024FFC5FFE7FF51FCECC764AF4C" bold="true" box="[199,328,862,887]" pageId="30" pageNumber="431">
<typeStatus id="92131294FFC5FFE7FF51FCECC76FAF4C" box="[199,323,862,887]" pageId="30" pageNumber="431" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="5BAE67BFFFC5FFE7FEC7FCECC7CEAF4C" box="[337,482,862,887]" pageId="30" pageNumber="431" type="generic">5 specimens</specimenCount>
: Anda1 (3)
<specimenCode id="1D0E044DFFC5FFE7FDFBFCEDC522AF4C" box="[621,782,862,887]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962243</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFC5FFE7FC8DFCEDC5ECAF4C" ID-GBIF-Occurrence="2013952729" box="[795,960,862,887]" collectionCode="RGM" isTailEnumeration="true" pageId="30" pageNumber="431" specimenCode="RGM 962244" specimenCount="1" typeStatus="paratype">
<specimenCode id="1D0E044DFFC5FFE7FC8DFCEDC590AF4C" box="[795,956,862,887]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962244</specimenCode>
,
</materialsCitation>
<materialsCitation id="FDC0A66BFFC5FFE7FC5FFCEDC247AF4C" ID-GBIF-Occurrence="2013952722" box="[969,1131,862,887]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962353" specimenCount="1" typeStatus="paratype">
<specimenCode id="1D0E044DFFC5FFE7FC5FFCEDC247AF4C" box="[969,1131,862,887]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962353</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFC5FFE7FBEEFCECC3B4AF4C" ID-GBIF-Occurrence="2013952719" box="[1144,1432,862,887]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962354" specimenCount="1" typeStatus="paratype">
Anda2 (1)
<specimenCode id="1D0E044DFFC5FFE7FB61FCEDC3B4AF4C" box="[1271,1432,862,887]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962354</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFC5FFE7FF01FC30C789AFA0" ID-GBIF-Occurrence="2013952727" box="[151,421,898,923]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962249" specimenCount="1" typeStatus="paratype">
Tiep2 (1)
<specimenCode id="1D0E044DFFC5FFE7FE90FC31C789AFA0" box="[262,421,898,923]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962249</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF51FC14C507AFD8" blockId="30.[151,1436,826,1571]" pageId="30" pageNumber="431">
<emphasis id="7FDC7024FFC5FFE7FF51FC14C7A9AF84" bold="true" box="[199,389,934,959]" pageId="30" pageNumber="431">Other material:</emphasis>
<materialsCitation id="FDC0A66BFFC5FFE7FE1BFC15C586AF84" ID-GBIF-Occurrence="2013952733" box="[397,938,934,959]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962245" specimenCount="13">
<specimenCount id="5BAE67BFFFC5FFE7FE1BFC15C40BAF84" box="[397,551,934,959]" pageId="30" pageNumber="431" type="generic">13 specimens</specimenCount>
in total. Anda1 (4)
<specimenCode id="1D0E044DFFC5FFE7FC9DFC15C586AF84" box="[779,938,934,959]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962245</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFC5FFE7FC21FC14C2FFAF84" ID-GBIF-Occurrence="2013952720" box="[951,1235,934,959]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962246" specimenCount="1">
Anda2 (4)
<specimenCode id="1D0E044DFFC5FFE7FBA5FC15C2FFAF84" box="[1075,1235,934,959]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962246</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFC5FFE7FB76FC14C6C3AFD8" ID-GBIF-Occurrence="2013952723" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962247" specimenCount="1">
Anda3 (2)
<specimenCode id="1D0E044DFFC5FFE7FACAFC15C6C3AFD8" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962247</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFC5FFE7FF6CFC78C42BAFD8" ID-GBIF-Occurrence="2013952728" box="[250,519,970,995]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962248" specimenCount="1">
Tiep1 (1)
<specimenCode id="1D0E044DFFC5FFE7FEFFFC79C42BAFD8" box="[361,519,970,995]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962248</specimenCode>
</materialsCitation>
;
<materialsCitation id="FDC0A66BFFC5FFE7FD84FC78C50BAFD8" ID-GBIF-Occurrence="2013952748" box="[530,807,970,995]" collectionCode="RGM" pageId="30" pageNumber="431" specimenCode="RGM 962250" specimenCount="1">
Tiep2 (2),
<specimenCode id="1D0E044DFFC5FFE7FD1EFC79C50BAFD8" box="[648,807,970,995]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RGM 962250</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFC5FFE7FF51FC5CC473A810" pageId="30" pageNumber="431" type="diagnosis">
<paragraph id="4D17AC36FFC5FFE7FF51FC5CC473A810" blockId="30.[151,1436,826,1571]" pageId="30" pageNumber="431">
<emphasis id="7FDC7024FFC5FFE7FF51FC5CC76FA83C" bold="true" box="[199,323,1006,1031]" pageId="30" pageNumber="431">Diagnosis:</emphasis>
A moderately elongated oval-shaped
<taxonomicName id="8AA8D7B5FFC5FFE7FD69FC5CC583A83C" authorityName="Alcock" authorityYear="1889" box="[767,943,1006,1031]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFC5FFE7FD69FC5CC583A83C" box="[767,943,1006,1031]" italics="true" pageId="30" pageNumber="431">Parascombrops</emphasis>
</taxonomicName>
otolith with distinct crenulation along the regularly curved ventral and dorsal rims.
</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFC5FFE7FF51FB84C2EEA874" box="[199,1218,1078,1103]" pageId="30" pageNumber="431" type="etymology">
<paragraph id="4D17AC36FFC5FFE7FF51FB84C2EEA874" blockId="30.[151,1436,826,1571]" box="[199,1218,1078,1103]" pageId="30" pageNumber="431">
<emphasis id="7FDC7024FFC5FFE7FF51FB84C77DA874" bold="true" box="[199,337,1078,1103]" pageId="30" pageNumber="431">Etymology:</emphasis>
In honor of Dr. Werner Schwarzhans for his contributions on paleoichthology.
</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFC5FFE6FF51FBE8C2ACABEF" lastPageId="31" lastPageNumber="432" pageId="30" pageNumber="431" type="description">
<paragraph id="4D17AC36FFC5FFE7FF51FBE8C431AA18" blockId="30.[151,1436,826,1571]" pageId="30" pageNumber="431">
<emphasis id="7FDC7024FFC5FFE7FF51FBE8C777A848" bold="true" box="[199,347,1114,1139]" pageId="30" pageNumber="431">Description:</emphasis>
The
<typeStatus id="92131294FFC5FFE7FE34FBE8C428A848" box="[418,516,1114,1139]" pageId="30" pageNumber="431" type="holotype">holotype</typeStatus>
is an moderately elongated oval-shaped specimen with a OL=
<quantity id="8A5001D3FFC5FFE7FA9CFBE9C34BA849" box="[1290,1383,1115,1139]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="30" pageNumber="431" unit="mm" value="3.2">3.2 mm</quantity>
and OL:OH= 1.91. The ventral rim is regularly bent with its strongest bending at its anterior part. The dorsal rim also bends regularly, but weaker, until a weak postdorsal angle at its very posterior part. Dorsal and ventral rims are regularly ornamented with small furrows and lobes. The very posterior part of the
<typeStatus id="92131294FFC5FFE7FBDEFB74C286A8E4" box="[1096,1194,1222,1247]" pageId="30" pageNumber="431" type="holotype">holotype</typeStatus>
bends downwards to the ventral rim, a feature that is more rounded in most
<typeStatus id="92131294FFC5FFE7FCB9FB59C5B2A938" box="[815,926,1259,1283]" pageId="30" pageNumber="431" type="paratype">paratypes</typeStatus>
. There is a blunt rostrum and an indistinct excisural notch. The straight sulcus displays a large oval-shaped ostium and a narrower nearly straight cauda. It spans nearly the whole length of the otolith and ends in a brief postcaudal depression, which slightly bends in posterior-ventral direction. The outer surface is decorated with many lobes and furrows that run perpendicular to the dorsal and ventral rims to a central flat area that runs in the mid-central part along the otolith. The
<typeStatus id="92131294FFC5FFE7FAA7FAC9C3B0A9A8" box="[1329,1436,1403,1427]" pageId="30" pageNumber="431" type="paratype">paratypes</typeStatus>
resemble the
<typeStatus id="92131294FFC5FFE7FEA6FA2CC7BEA98C" box="[304,402,1438,1463]" pageId="30" pageNumber="431" type="holotype">holotype</typeStatus>
very well; the OL:OH ratio ranges between 1.73 and ~1.90, and the posterior rim is more rounded, with a shallow postdorsal angle above the end of the cauda. In small specimens the postdorsal angle is not developed, and the postdorsal field form a bluntly pointed extension behind the cauda, which gives the otolith a rounded spindle-shape impression.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F928C72EAA8B" blockId="30.[151,1435,1690,2019]" box="[151,258,1690,1712]" pageId="30" pageNumber="431">
<emphasis id="7FDC7024FFC5FFE7FF01F928C72EAA8B" bold="true" box="[151,258,1690,1712]" pageId="30" pageNumber="431">PLATE 9.</emphasis>
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F90BC4C5AAF4" blockId="30.[151,1435,1690,2019]" box="[151,745,1721,1743]" pageId="30" pageNumber="431">
93.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF90BC73AAAF4" authorityName="Rafinesque" authorityYear="1818" box="[189,278,1721,1743]" class="Actinopterygii" family="Sparidae" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="family">Sparidae</taxonomicName>
indeterminate, RGM 962339, locality Anda2.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F96AC53DAAD5" blockId="30.[151,1435,1690,2019]" box="[151,785,1752,1774]" pageId="30" pageNumber="431">
94.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF96BC7E0AAD5" authorityName="Fourmanoir" authorityYear="1985" box="[189,460,1753,1774]" class="Actinopterygii" family="Cepolidae" genus="Owstonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="7FDC7024FFC5FFE7FF2BF96BC7E0AAD5" box="[189,460,1753,1774]" italics="true" pageId="30" pageNumber="431">Owstonia nigromarginatus</emphasis>
</taxonomicName>
, RGM 962277, locality Roxas.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F944C539AB37" blockId="30.[151,1435,1690,2019]" box="[151,789,1782,1805]" pageId="30" pageNumber="431">
95.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF945C7E0AB36" authorityName="Fourmanoir" authorityYear="1985" box="[189,460,1783,1805]" class="Actinopterygii" family="Cepolidae" genus="Owstonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="7FDC7024FFC5FFE7FF2BF945C7E0AB36" box="[189,460,1783,1805]" italics="true" pageId="30" pageNumber="431">Owstonia nigromarginatus</emphasis>
</taxonomicName>
, RGM 962275, locality Anda2.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F8A7C526AB10" blockId="30.[151,1435,1690,2019]" box="[151,778,1813,1835]" pageId="30" pageNumber="431">
96.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF8A4C7E0AB10" authorityName="Fourmanoir" authorityYear="1985" box="[189,460,1814,1835]" class="Actinopterygii" family="Cepolidae" genus="Owstonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="7FDC7024FFC5FFE7FF2BF8A4C7E0AB10" box="[189,460,1814,1835]" italics="true" pageId="30" pageNumber="431">Owstonia nigromarginatus</emphasis>
</taxonomicName>
, RGM 962278, locality Roxas.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F886C5B5AB71" blockId="30.[151,1435,1690,2019]" box="[151,921,1844,1866]" pageId="30" pageNumber="431">
97.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF886C7E0AB71" authorityName="Fourmanoir" authorityYear="1985" box="[189,460,1844,1866]" class="Actinopterygii" family="Cepolidae" genus="Owstonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="7FDC7024FFC5FFE7FF2BF886C7E0AB71" box="[189,460,1844,1866]" italics="true" pageId="30" pageNumber="431">Owstonia nigromarginatus</emphasis>
</taxonomicName>
, RGM 962274, locality Anda2, SEM image.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F8E0C4AFAB53" blockId="30.[151,1435,1690,2019]" box="[151,643,1874,1896]" pageId="30" pageNumber="431">
98.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF8E1C732AB53" authorityName="Tanaka" authorityYear="1908" box="[189,286,1875,1896]" class="Actinopterygii" family="Cepolidae" genus="Owstonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFC5FFE7FF2BF8E1C732AB53" box="[189,286,1875,1896]" italics="true" pageId="30" pageNumber="431">Owstonia</emphasis>
</taxonomicName>
sp., RGM 962272, locality Anda6.
</paragraph>
<paragraph id="4D17AC36FFC5FFE7FF01F8C3C4AFABBC" blockId="30.[151,1435,1690,2019]" box="[151,643,1905,1927]" pageId="30" pageNumber="431">
99.
<taxonomicName id="8AA8D7B5FFC5FFE7FF2BF8C0C732ABBC" authorityName="Tanaka" authorityYear="1908" box="[189,286,1906,1927]" class="Actinopterygii" family="Cepolidae" genus="Owstonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFC5FFE7FF2BF8C0C732ABBC" box="[189,286,1906,1927]" italics="true" pageId="30" pageNumber="431">Owstonia</emphasis>
</taxonomicName>
sp., RGM 962271, locality Anda2.
</paragraph>
<footnote id="2EB3B038FFC5FFE7FF01F822C4CDAB9D" box="[151,737,1936,1958]" pageId="30" pageNumber="431">
<paragraph id="4D17AC36FFC5FFE7FF01F822C4CDAB9D" blockId="30.[151,1435,1690,2019]" box="[151,737,1936,1958]" pageId="30" pageNumber="431">
100.
<taxonomicName id="8AA8D7B5FFC5FFE7FF5FF823C754AB9D" box="[201,376,1937,1958]" class="Actinopterygii" family="Caproidae" genus="Antigonia" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="capros">
<emphasis id="7FDC7024FFC5FFE7FF5FF823C754AB9D" box="[201,376,1937,1958]" italics="true" pageId="30" pageNumber="431">Antigonia capros</emphasis>
</taxonomicName>
, RGM 962309, locality AndaClif3.
</paragraph>
</footnote>
<footnote id="2EB3B038FFC5FFE7FF01F81CC6CEABD8" pageId="30" pageNumber="431">
<paragraph id="4D17AC36FFC5FFE7FF01F81CC6CEABD8" blockId="30.[151,1435,1690,2019]" pageId="30" pageNumber="431">
101.
<taxonomicName id="8AA8D7B5FFC5FFE7FF58F81CC46FABFE" baseAuthorityName="Smith &amp; Radcliffe" baseAuthorityYear="1912" box="[206,579,1966,1989]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="30" pageNumber="431" phylum="Chordata" rank="species" species="serratospinosus">
<emphasis id="7FDC7024FFC5FFE7FF58F81CC46FABFE" box="[206,579,1966,1989]" italics="true" pageId="30" pageNumber="431">Parascombrops aff. serratospinosus</emphasis>
</taxonomicName>
, RGM 962253, locality AndaClif3. (a) Lateral face, (b) medial face, (c) ventral profile.
</paragraph>
</footnote>
<caption id="19D7FCBEFFC4FFE6FF01F897C6CEAB62" httpUri="https://zenodo.org/record/2601303/files/figure.png" pageId="31" pageNumber="432" startId="31.[151,234,1829,1851]" targetBox="[164,1415,193,1811]" targetPageId="31">
<paragraph id="4D17AC36FFC4FFE6FF01F897C6CEAB62" blockId="31.[151,1392,1829,2035]" pageId="31" pageNumber="432">
<emphasis id="7FDC7024FFC4FFE6FF01F897C73CAB00" bold="true" box="[151,272,1829,1851]" pageId="31" pageNumber="432">PLATE 10.</emphasis>
102.
<taxonomicName id="8AA8D7B5FFC4FFE6FEDEF897C41AAB00" box="[328,566,1829,1851]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="432" phylum="Chordata" rank="species" species="schwar">
<emphasis id="7FDC7024FFC4FFE6FEDEF897C41AAB00" box="[328,566,1829,1851]" italics="true" pageId="31" pageNumber="432">Parascombrops schwar</emphasis>
</taxonomicName>
z
<emphasis id="7FDC7024FFC4FFE6FDD4F897C455AB00" box="[578,633,1829,1851]" italics="true" pageId="31" pageNumber="432">hansi</emphasis>
, RGM 962352, locality Anda2. (a) Medial face, (b) lateral face, (c) ventral profile.
</paragraph>
</caption>
<paragraph id="4D17AC36FFC4FFE6FF01F8D0C2ACABEF" blockId="31.[151,1392,1829,2035]" pageId="31" pageNumber="432">
103.
<taxonomicName id="8AA8D7B5FFC4FFE6FF5FF8D0C794AB43" box="[201,440,1890,1912]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="432" phylum="Chordata" rank="species" species="schwar">
<emphasis id="7FDC7024FFC4FFE6FF5FF8D0C794AB43" box="[201,440,1890,1912]" italics="true" pageId="31" pageNumber="432">Parascombrops schwar</emphasis>
</taxonomicName>
z
<emphasis id="7FDC7024FFC4FFE6FE52F8D0C7D6AB43" box="[452,506,1890,1912]" italics="true" pageId="31" pageNumber="432">hansi</emphasis>
, RGM 962353, locality Anda1. (a) Medial face, (b) lateral face, (c) ventral profile. 104.
<taxonomicName id="8AA8D7B5FFC4FFE6FF5FF833C794ABAC" box="[201,440,1921,1943]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="432" phylum="Chordata" rank="species" species="schwar">
<emphasis id="7FDC7024FFC4FFE6FF5FF833C794ABAC" box="[201,440,1921,1943]" italics="true" pageId="31" pageNumber="432">Parascombrops schwar</emphasis>
</taxonomicName>
z
<emphasis id="7FDC7024FFC4FFE6FE52F833C7D6ABAC" box="[452,506,1921,1943]" italics="true" pageId="31" pageNumber="432">hansi</emphasis>
, RGM 962244, locality Anda1. (a) Medial face, (b) lateral face, (c) ventral profile. 105.
<taxonomicName id="8AA8D7B5FFC4FFE6FF5FF82DC794AB8E" box="[201,440,1951,1973]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="432" phylum="Chordata" rank="species" species="schwar">
<emphasis id="7FDC7024FFC4FFE6FF5FF82DC794AB8E" box="[201,440,1951,1973]" italics="true" pageId="31" pageNumber="432">Parascombrops schwar</emphasis>
</taxonomicName>
z
<emphasis id="7FDC7024FFC4FFE6FE52F82DC7D6AB8E" box="[452,506,1951,1973]" italics="true" pageId="31" pageNumber="432">hansi</emphasis>
, RGM 962243, locality Anda1. (a) Medial face, (b) lateral face, (c) ventral profile. 106.
<taxonomicName id="8AA8D7B5FFC4FFE6FF5FF80CC794ABEF" box="[201,440,1982,2004]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="432" phylum="Chordata" rank="species" species="schwar">
<emphasis id="7FDC7024FFC4FFE6FF5FF80CC794ABEF" box="[201,440,1982,2004]" italics="true" pageId="31" pageNumber="432">Parascombrops schwar</emphasis>
</taxonomicName>
z
<emphasis id="7FDC7024FFC4FFE6FE52F80CC7D6ABEF" box="[452,506,1982,2004]" italics="true" pageId="31" pageNumber="432">hansi</emphasis>
, RGM 962249, locality Tiep2. (a) Lateral face, (b) medial face.
</paragraph>
</subSubSection>
<footnote id="2EB3B038FFC4FFE6FF01F86FC2A5ABC8" box="[151,1161,2013,2035]" pageId="31" pageNumber="432">
<paragraph id="4D17AC36FFC4FFE6FF01F86FC2A5ABC8" blockId="31.[151,1392,1829,2035]" box="[151,1161,2013,2035]" pageId="31" pageNumber="432">
107.
<taxonomicName id="8AA8D7B5FFC4FFE6FF5FF86FC794ABC8" box="[201,440,2013,2035]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="31" pageNumber="432" phylum="Chordata" rank="species" species="schwar">
<emphasis id="7FDC7024FFC4FFE6FF5FF86FC794ABC8" box="[201,440,2013,2035]" italics="true" pageId="31" pageNumber="432">Parascombrops schwar</emphasis>
</taxonomicName>
z
<emphasis id="7FDC7024FFC4FFE6FE52F86FC7D6ABC8" box="[452,506,2013,2035]" italics="true" pageId="31" pageNumber="432">hansi</emphasis>
, RGM 962354, locality Anda2. (a) Lateral face, (b) medial face.
</paragraph>
</footnote>
<subSubSection id="05B2FFBDFFFBFFD9FF51FF25C46FADB3" pageId="32" pageNumber="433" type="discussion">
<paragraph id="4D17AC36FFFBFFD9FF51FF25C46FADB3" blockId="32.[151,1436,151,429]" pageId="32" pageNumber="433">
<emphasis id="7FDC7024FFFBFFD9FF51FF25C762AC8B" bold="true" box="[199,334,151,176]" pageId="32" pageNumber="433">Discussion:</emphasis>
The specimens are reminiscent of
<taxonomicName id="8AA8D7B5FFFBFFD9FD42FF2BC2BCAC8B" authority="Schwarzhans &amp; Prokofiev, 2017" authorityName="Schwarzhans &amp; Prokofiev" authorityYear="2017" box="[724,1168,151,176]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="433" phylum="Chordata" rank="species" species="ohei">
<emphasis id="7FDC7024FFFBFFD9FD42FF2BC533AC8B" box="[724,799,151,176]" italics="true" pageId="32" pageNumber="433">P. ohei</emphasis>
<bibRefCitation id="2939D1C7FFFBFFD9FCB0FF25C2BCAC8B" author="Schwarzhans, W. &amp; Prokofiev, A. M." box="[806,1168,151,176]" pageId="32" pageNumber="433" pagination="1 - 74" refId="ref19934" refString="Schwarzhans, W. &amp; Prokofiev, A. M. (2017) Reappraisal of Synagrops Gunther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae). Zootaxa, 4260 (1), 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 4260.1.1" type="journal article" year="2017">Schwarzhans &amp; Prokofiev, 2017</bibRefCitation>
</taxonomicName>
and juvenile specimens of
<taxonomicName id="8AA8D7B5FFFBFFD9FF23FF0FC489ACEF" authority="Schwarzhans &amp; Prokofiev, 2017" authorityName="Schwarzhans &amp; Prokofiev" authorityYear="2017" box="[181,677,188,213]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="433" phylum="Chordata" rank="species" species="argyreus">
<emphasis id="7FDC7024FFFBFFD9FF23FF0FC71EACEF" box="[181,306,189,212]" italics="true" pageId="32" pageNumber="433">P. argyreus</emphasis>
<bibRefCitation id="2939D1C7FFFBFFD9FEACFF0EC489ACEF" author="Schwarzhans, W. &amp; Prokofiev, A. M." box="[314,677,188,213]" pageId="32" pageNumber="433" pagination="1 - 74" refId="ref19934" refString="Schwarzhans, W. &amp; Prokofiev, A. M. (2017) Reappraisal of Synagrops Gunther, 1887 with rehabilitation and revision of Parascombrops Alcock, 1889 including description of seven new species and two new genera (Perciformes: Acropomatidae). Zootaxa, 4260 (1), 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 4260.1.1" type="journal article" year="2017">Schwarzhans &amp; Prokofiev, 2017</bibRefCitation>
</taxonomicName>
(also depicted by
<bibRefCitation id="2939D1C7FFFBFFD9FCE2FF0EC2B4ACEF" author="Rivaton, J. &amp; Bourret, P." box="[884,1176,188,213]" pageId="32" pageNumber="433" refId="ref19612" refString="Rivaton, J. &amp; Bourret, P. (1999) Otoliths of the Indo-Pacific fishes. Documents Scientifiques et Techniques, Centre IRD de Noumea, Noumea, 378 pp." type="book" year="1999">Rivaton &amp; Bourret (1999)</bibRefCitation>
as
<taxonomicName id="8AA8D7B5FFFBFFD9FB28FF0FC339ACEE" box="[1214,1301,188,213]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="433" phylum="Chordata" rank="species" species="adeni">
<emphasis id="7FDC7024FFFBFFD9FB28FF0FC339ACEE" box="[1214,1301,188,213]" italics="true" pageId="32" pageNumber="433">P. adeni</emphasis>
</taxonomicName>
). However, they are more rounded and less elongated than
<taxonomicName id="8AA8D7B5FFFBFFD9FD2FFF53C528ACC3" authorityName="Schwarzhans &amp; Prokofiev" authorityYear="2017" box="[697,772,223,248]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="433" phylum="Chordata" rank="species" species="ohei">
<emphasis id="7FDC7024FFFBFFD9FD2FFF53C528ACC3" box="[697,772,223,248]" italics="true" pageId="32" pageNumber="433">P. ohei</emphasis>
</taxonomicName>
, and, display a very characteristic crenulation along their dorsal and ventral rims. One larger otolith (
<quantity id="8A5001D3FFFBFFD9FD37FEB6C52EAD27" box="[673,770,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="32" pageNumber="433" unit="mm" value="7.2">7.2 mm</quantity>
), RGM 962251, was encountered in Anda4, which may represent a strongly polished and therewith poorly preserved adult specimen of
<taxonomicName id="8AA8D7B5FFFBFFD9FBD9FE95C2D3AD7B" authorityName="Alcock" authorityYear="1889" box="[1103,1279,295,320]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="433" phylum="Chordata" rank="genus">
<emphasis id="7FDC7024FFFBFFD9FBD9FE95C2D3AD7B" box="[1103,1279,295,320]" italics="true" pageId="32" pageNumber="433">Parascombrops</emphasis>
</taxonomicName>
. Its size and estimated shape may suggest
<taxonomicName id="8AA8D7B5FFFBFFD9FE74FEFEC237AD5F" authority="(Gilbert &amp; Cramer, 1897)" baseAuthorityName="Gilbert &amp; Cramer" baseAuthorityYear="1897" box="[482,1051,332,357]" class="Actinopterygii" family="Acropomatidae" genus="Parascombrops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Perciformes" pageId="32" pageNumber="433" phylum="Chordata" rank="species" species="argyreus">
<emphasis id="7FDC7024FFFBFFD9FE74FEFEC4D7AD5F" box="[482,763,332,357]" italics="true" pageId="32" pageNumber="433">Parascombrops argyreus</emphasis>
(Gilbert &amp;Cramer, 1897)
</taxonomicName>
, but its preservation state does not allow definitive species identification.
</paragraph>
</subSubSection>
<subSubSection id="05B2FFBDFFFBFFD9FF51FE26C564AD96" box="[199,840,404,429]" pageId="32" pageNumber="433" type="distribution">
<paragraph id="4D17AC36FFFBFFD9FF51FE26C564AD96" blockId="32.[151,1436,151,429]" box="[199,840,404,429]" pageId="32" pageNumber="433">
<emphasis id="7FDC7024FFFBFFD9FF51FE26C74EAD96" bold="true" box="[199,354,404,429]" pageId="32" pageNumber="433">Distribution:</emphasis>
Plio-/Pleistocene of northwest
<collectingCountry id="35BFECA6FFFBFFD9FD54FE26C568AD96" box="[706,836,404,429]" name="Philippines" pageId="32" pageNumber="433">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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