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<document ID-DOI="http://doi.org/10.5281/zenodo.3788773" ID-GBIF-Dataset="6dfdec5c-2e9d-4e1f-be4c-a120edc779e0" ID-GBIF-Taxon="163747513" ID-ISSN="1313–2970" ID-Zenodo-Dep="3788773" ID-ZooBank="6D59834F-82C0-4DCD-8F65-202AE8F03965" LSID="urn:lsid:zoobank.org:act:85959B2B-9561-4B4F-9070-4E7906F0AA72" approvalRequired="61" approvalRequired_for_textStreams="61" checkinTime="1587954401351" checkinUser="plazi" docAuthor="Hawks, David" docDate="2010" docId="777587FF090CFFA7FF43538CB3E9FA34" docLanguage="en" docName="ZK_article_2166.pdf" docOrigin="ZooKeys 39 (39)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@7e86ff3b" docStyleName="zookeys.2008.journal_article" docTitle="Catocala chelidonia subsp. uniforma Hawks 2010, ssp. n." docType="treatment" docUuid="85959B2B-9561-4B4F-9070-4E7906F0AA72" docUuidSource="ZooBank" docVersion="5" lastPageId="19" lastPageNumber="32" masterDocId="8B4CFF87091EFFB4FFCF5615B045FFA5" masterDocTitle="Review of the Catocala delilah species complex (Lepidoptera, Erebidae)" masterLastPageNumber="35" masterPageNumber="13" pageId="18" pageNumber="31" updateTime="1643557595621" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Catocala delilah species complex (Lepidoptera, Erebidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hawks, David</mods:namePart>
<mods:affiliation>University of California, Riverside ,, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-03-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>39</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>39</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>35</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.39.439</mods:identifier>
<mods:identifier type="GBIF-Dataset">6dfdec5c-2e9d-4e1f-be4c-a120edc779e0</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576631</mods:identifier>
<mods:identifier type="ZooBank">6D59834F-82C0-4DCD-8F65-202AE8F03965</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788773" ID-GBIF-Taxon="163747513" ID-Zenodo-Dep="3788773" LSID="urn:lsid:zoobank.org:act:85959B2B-9561-4B4F-9070-4E7906F0AA72" httpUri="http://treatment.plazi.org/id/777587FF090CFFA7FF43538CB3E9FA34" lastPageId="19" lastPageNumber="32" pageId="18" pageNumber="31">
<subSubSection pageId="18" pageNumber="31" type="nomenclature">
<paragraph blockId="18.[140,949,1433,1531]" box="[140,676,1433,1460]" pageId="18" pageNumber="31">
<heading box="[140,676,1433,1460]" fontSize="11" level="2" pageId="18" pageNumber="31" reason="3">
<emphasis bold="true" box="[140,676,1433,1460]" pageId="18" pageNumber="31">
<taxonomicName authority="Hawks" authorityName="Hawks" authorityYear="2010" box="[140,589,1433,1460]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="chelidonia" status="ssp. n." subSpecies="uniforma">
<emphasis bold="true" box="[140,497,1433,1460]" italics="true" pageId="18" pageNumber="31">
<emphasis box="[140,246,1433,1460]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Catocala</emphasis>
chelidonia uniforma
</emphasis>
Hawks
</taxonomicName>
,
<taxonomicNameLabel box="[601,676,1434,1460]" pageId="18" pageNumber="31" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[140,949,1433,1531]" box="[140,949,1469,1496]" pageId="18" pageNumber="31">
<uri box="[140,949,1469,1496]" pageId="18" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid box="[416,949,1469,1496]" pageId="18" pageNumber="31">85959B2B-9561-4B4F-9070-4E7906F0AA72</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,265,1504,1531]" pageId="18" pageNumber="31" type="description">
<paragraph blockId="18.[140,949,1433,1531]" box="[140,265,1504,1531]" pageId="18" pageNumber="31">Figs 25, 26</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="32" pageId="18" pageNumber="31" type="materials_examined">
<paragraph blockId="18.[140,1107,1574,1672]" pageId="18" pageNumber="31">
<emphasis bold="true" box="[140,313,1574,1601]" pageId="18" pageNumber="31">
<emphasis box="[140,198,1575,1601]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608811627" collectionCode="PMNH" county="Cochise County" elevation="1700" location="Huachuca Mountains" municipality="Ash Canyon" pageId="18" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Arizona" typeStatus="holotype">
<emphasis bold="true" box="[322,435,1575,1601]" pageId="18" pageNumber="31">
<typeStatus box="[322,435,1575,1601]" pageId="18" pageNumber="31">Holotype</typeStatus>
</emphasis>
♁ (Fig. 25,
<collectionCode box="[579,668,1575,1601]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="18" pageNumber="31" type="Museum">PMNH</collectionCode>
specimen #ENT 721053; FW length 20.0 mm),
<emphasis bold="true" box="[273,369,1610,1636]" pageId="18" pageNumber="31">
<typeStatus box="[273,369,1610,1636]" pageId="18" pageNumber="31">allotype</typeStatus>
</emphasis>
<specimenCount box="[379,394,1610,1636]" pageId="18" pageNumber="31" type="female">♀</specimenCount>
(F ig. 26,
<collectionCode box="[519,608,1610,1636]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="18" pageNumber="31" type="Museum">PMNH</collectionCode>
specimen #ENT 721054; FW length 19.5 mm):
<collectingMunicipality box="[210,354,1645,1671]" pageId="18" pageNumber="31">Ash Canyon</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090CFFA6FEA25078B222F922" box="[365,615,1645,1671]" county="Cochise County" municipality="Ash Canyon" name="Huachuca Mountains" pageId="18" pageNumber="31" stateProvince="Arizona">Huachuca Mountains</location>
,
<collectingCounty box="[627,812,1645,1671]" pageId="18" pageNumber="31">Cochise County</collectingCounty>
,
<collectingRegion box="[824,915,1645,1671]" country="United States of America" name="Arizona" pageId="18" pageNumber="31">Arizona</collectingRegion>
,
<quantity box="[926,1018,1645,1672]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="31" unit="m" value="1700.0">
<elevation box="[926,1018,1645,1672]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="31" unit="m" value="1700.0">1700 m</elevation>
</quantity>
, leg. N.
</materialsCitation>
</paragraph>
<paragraph blockId="19.[140,1108,166,932]" pageId="19" pageNumber="32">
<materialsCitation ID-GBIF-Occurrence="2608811628" box="[140,266,167,193]" location="McFarland" pageId="19" pageNumber="32" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FF4356B2B14FFF64" box="[140,266,167,193]" name="McFarland" pageId="19" pageNumber="32">McFarland</location>
</materialsCitation>
;
<typeStatus box="[276,375,167,193]" pageId="19" pageNumber="32">holotype</typeStatus>
<date box="[382,520,167,193]" pageId="19" pageNumber="32" value="1981-06-02">2 June 1981</date>
,
<typeStatus box="[531,620,167,193]" pageId="19" pageNumber="32">allotype</typeStatus>
<date box="[626,781,166,193]" pageId="19" pageNumber="32" value="1983-06-15">15 June 1983</date>
.
<emphasis bold="true" box="[791,906,167,193]" pageId="19" pageNumber="32">
<typeStatus box="[791,906,167,193]" pageId="19" pageNumber="32">Paratypes</typeStatus>
</emphasis>
(13 ♁
<specimenCount box="[981,1031,167,193]" pageId="19" pageNumber="32" type="female">26 ♀</specimenCount>
): 8 ♁
<specimenCount box="[140,191,202,228]" pageId="19" pageNumber="32" type="female">12 ♀</specimenCount>
from the type locality, varying dates and collectors;
<materialsCitation ID-GBIF-Occurrence="2608811620" county="Cochise County" location="Sierra Vista" pageId="19" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Arizona">
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FCCD56DFB3C3FF41" box="[770,902,202,228]" county="Cochise County" name="Sierra Vista" pageId="19" pageNumber="32" stateProvince="Arizona">Sierra Vista</location>
,
<collectingCounty box="[912,1096,202,228]" pageId="19" pageNumber="32">Cochise County</collectingCounty>
,
<collectingRegion box="[140,228,237,263]" country="United States of America" name="Arizona" pageId="19" pageNumber="32">Arizona</collectingRegion>
(
<specimenCount box="[250,289,237,264]" pageId="19" pageNumber="32" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811623" county="Cochise County" location="Huachuca Mountains" municipality="Miller Canyon" pageId="19" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Arizona">
<collectingMunicipality box="[316,490,237,263]" pageId="19" pageNumber="32">Miller Canyon</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FE3656F8B2B3FEA2" box="[505,758,237,263]" county="Cochise County" municipality="Miller Canyon" name="Huachuca Mountains" pageId="19" pageNumber="32" stateProvince="Arizona">Huachuca Mountains</location>
,
<collectingCounty box="[774,963,237,263]" pageId="19" pageNumber="32">Cochise County</collectingCounty>
,
<collectingRegion box="[977,1065,237,263]" country="United States of America" name="Arizona" pageId="19" pageNumber="32">Arizona</collectingRegion>
(
<specimenCount pageId="19" pageNumber="32" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811630" collectingDate="2007-06-11" collectingDateMax="2007-06-12" collectingDateMin="2007-06-11" collectorName="M. L. Raschko" county="Cochise County" location="Sunny Flat Campground" municipality="Chiricahua Mountains" pageId="19" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Arizona">
<collectingCounty box="[181,372,272,298]" pageId="19" pageNumber="32">Cochise County</collectingCounty>
:
<collectingMunicipality box="[385,648,272,298]" pageId="19" pageNumber="32">Chiricahua Mountains</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FD595705B3FCFE8F" box="[662,953,272,298]" county="Cochise County" municipality="Chiricahua Mountains" name="Sunny Flat Campground" pageId="19" pageNumber="32" stateProvince="Arizona">Sunny Flat Campground</location>
,
<date pageId="19" pageNumber="32" value="2007-06-11" valueMax="2007-06-12" valueMin="2007-06-11">
<collectingDate pageId="19" pageNumber="32" value="2007-06-11" valueMax="2007-06-12" valueMin="2007-06-11">11–12 June 2007</collectingDate>
</date>
,
<collectorName box="[215,376,307,334]" pageId="19" pageNumber="32">M.L. Raschko</collectorName>
(1 ♁,
<specimenCount box="[454,492,308,334]" pageId="19" pageNumber="32" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811622" location="South Fork Cave Creek" municipality="Chiricahua Mountains" pageId="19" pageNumber="32" specimenCount="2" specimenCount-female="2" stateProvince="Arizona">
<collectingMunicipality box="[518,780,307,333]" pageId="19" pageNumber="32">Chiricahua Mountains</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FCD65726B46DFEE8" box="[793,1064,307,334]" municipality="Chiricahua Mountains" name="South Fork Cave Creek" pageId="19" pageNumber="32" stateProvince="Arizona">South Fork Cave Creek</location>
(
<specimenCount pageId="19" pageNumber="32" type="female">2 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811640" box="[183,1084,342,370]" location="Southwestern Research Station" municipality="Chiricahua Mountains" pageId="19" pageNumber="32" specimenCount="5" specimenCount-female="5" stateProvince="Arizona">
<collectingMunicipality box="[183,447,343,369]" pageId="19" pageNumber="32">Chiricahua Mountains</collectingMunicipality>
, vicinity of
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FD9F5742B3F0FED4" box="[592,949,343,369]" municipality="Chiricahua Mountains" name="Southwestern Research Station" pageId="19" pageNumber="32" stateProvince="Arizona">Southwestern Research Station</location>
(3 ♁,
<specimenCount box="[1032,1073,342,369]" pageId="19" pageNumber="32" type="female">5 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811657" box="[140,468,378,404]" location="Chiricahua Mountains" pageId="19" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Arizona">
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FF43576FB1CAFE31" box="[140,399,378,404]" name="Chiricahua Mountains" pageId="19" pageNumber="32" stateProvince="Arizona">Chiricahua Mountains</location>
(
<specimenCount box="[420,457,378,404]" pageId="19" pageNumber="32" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811629" collectorName="Clanton Draw" county="Hidalgo County" location="Peloncillo Mountains" pageId="19" pageNumber="32" specimenCount="3" specimenCount-female="3" stateProvince="New Mexico">
<collectingRegion box="[483,633,378,404]" country="United States of America" name="New Mexico" pageId="19" pageNumber="32">New Mexico</collectingRegion>
:
<collectingCounty box="[646,835,378,404]" pageId="19" pageNumber="32">Hidalgo County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FC9E576FB40DFE31" box="[849,1096,378,404]" county="Hidalgo County" name="Peloncillo Mountains" pageId="19" pageNumber="32" stateProvince="New Mexico">Peloncillo Mountains</location>
,
<collectorName box="[140,304,413,440]" pageId="19" pageNumber="32">Clanton Draw</collectorName>
(
<specimenCount box="[321,358,413,440]" pageId="19" pageNumber="32" type="female">3 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811624" collectionCode="PMNH, LACM" country="United States of America" location="Bear Canyon" municipality="Animas Mountains" pageId="19" pageNumber="32" specimenCount="3" typeStatus="holotype">
<collectingMunicipality box="[382,602,413,439]" pageId="19" pageNumber="32">Animas Mountains</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA7FF43538CB3E9FA34:FA036032090DFFA7FDA9578BB2BFFE12" box="[614,762,413,440]" country="United States of America" municipality="Animas Mountains" name="Bear Canyon" pageId="19" pageNumber="32">Bear Canyon</location>
(1 ♁).
<typeStatus box="[844,951,413,439]" pageId="19" pageNumber="32">Holotype</typeStatus>
and
<typeStatus box="[1008,1097,413,439]" pageId="19" pageNumber="32">allotype</typeStatus>
deposited at
<collectionCode box="[285,377,449,475]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="19" pageNumber="32" type="Museum">PMNH</collectionCode>
,
<typeStatus box="[389,495,449,475]" pageId="19" pageNumber="32">paratypes</typeStatus>
deposited at
<collectionCode box="[648,730,448,474]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="19" pageNumber="32" type="Museum">LACM</collectionCode>
and other
<collectingCountry box="[856,911,448,474]" name="United States of America" pageId="19" pageNumber="32">USA</collectingCountry>
museums as well as in private collections
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="diagnosis">
<paragraph blockId="19.[140,1108,166,932]" pageId="19" pageNumber="32">
<emphasis bold="true" box="[185,312,518,544]" pageId="19" pageNumber="32">
<emphasis box="[185,307,518,544]" inLineHeading="true" pageId="19" pageNumber="32" reason="1">Diagnosis</emphasis>
.
</emphasis>
This subspecies has uniform gray forewings with substantially reduced hoary mottling. Th e antemedial and postmedial forewing lines vary from inconspicuous to very distinct. Hindwing as in the nominate subspecies.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="description">
<paragraph blockId="19.[140,1108,166,932]" pageId="19" pageNumber="32">
<emphasis bold="true" box="[185,335,624,650]" pageId="19" pageNumber="32">
<emphasis box="[185,330,624,650]" inLineHeading="true" pageId="19" pageNumber="32" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[346,446,625,650]" italics="true" pageId="19" pageNumber="32">Forewing</emphasis>
: dorsal surface ground color charcoal gray, concolorous, with few hoary scales; antemedial and postmedial lines black; antemedial line fades away toward the inner margin; subreniform whitish to yellow, usually visible.
<emphasis italics="true" pageId="19" pageNumber="32">Hindwing</emphasis>
: pale areas dull yellow orange; bands black, apical spot dull yellow orange; anal spot separated from outer band; few dark scales at base.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="32" type="distribution">
<paragraph blockId="19.[140,1108,166,932]" pageId="19" pageNumber="32">
<emphasis bold="true" box="[185,497,800,826]" pageId="19" pageNumber="32">
<emphasis box="[185,336,800,826]" inLineHeading="true" pageId="19" pageNumber="32" reason="1">Distribution</emphasis>
and biology.
</emphasis>
<taxonomicName box="[508,837,800,826]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="chelidonia" subSpecies="uniforma">
<emphasis box="[508,837,800,826]" italics="true" pageId="19" pageNumber="32">Catocala chelidonia uniforma</emphasis>
</taxonomicName>
is known from mountains of southeastern
<collectingRegion box="[377,465,835,861]" country="United States of America" name="Arizona" pageId="19" pageNumber="32">Arizona</collectingRegion>
and southwestern
<collectingRegion box="[675,822,835,862]" country="United States of America" name="New Mexico" pageId="19" pageNumber="32">New Mexico</collectingRegion>
. Nothing is known of the early stages of this subspecies. Adults have been collected from late May to mid-July, with peak abundance in mid-June (N. McFarland, pers. comm.).
</paragraph>
<paragraph blockId="19.[140,393,1026,1053]" box="[140,393,1026,1053]" pageId="19" pageNumber="32">
<emphasis bold="true" box="[140,393,1026,1053]" pageId="19" pageNumber="32">Acknowledgments</emphasis>
</paragraph>
<paragraph blockId="19.[140,1108,1082,1425]" pageId="19" pageNumber="32">
I thank J.W. Johnson, W.W. Mayhew, the late K.W. Cooper, J.D. Pinto, and the late M.M. Barnes (University of California, Riverside) for their support of this research over the years. M.L. Raschko (Temecula, CA), D.L. Wikle (
<collectingCountry box="[801,937,1152,1178]" name="San Marino" pageId="19" pageNumber="32">San Marino</collectingCountry>
, CA) and K.H. Osborne (Riverside, CA) offered useful discussions of western
<taxonomicName authorityName=", Barnes and McDunnough" authorityYear="1918" box="[853,948,1187,1213]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis box="[853,948,1187,1213]" italics="true" pageId="19" pageNumber="32">Catocala</emphasis>
</taxonomicName>
and provided access to specimens in their personal collections. L.F. Gall (PMNH), C. Schmidt, and D. Lafontaine provided editorial assistance and helpful suggestions on the manuscript. J.D. Gill (CNC) and A. M. Burg (PMNH) helped prepare the Figures. Most of the text was condensed from a 1986 Masters Th esis submitted by DCH to the Graduate Program in Evolutionary Biology at the University of California, Riverside. Partial financial assistance provided by the Goelet-Cary Fund, Yale University.
</paragraph>
</subSubSection>
</treatment>
</document>
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