treatments-xml/data/0E/36/BF/0E36BF5DFFFFFF9F947DD58C0E4FFB50.xml
2024-06-21 12:29:25 +02:00

177 lines
22 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="AA1DB5E8855307812D5568838459441D" ID-CLB-Dataset="8112" ID-DOI="10.37520/aemnp.2020.25" ID-GBIF-Dataset="bd0522ba-e842-4416-82ac-98640acd8e45" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489312" ID-ZooBank="94935C98-664E-4D3C-87FA-F3FC6513512B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612221404353" checkinUser="felipe" docAuthor="Nabozhenko, Maxim, Keskin, Bekir &amp; Papadopoulou, Anna" docDate="2020" docId="0E36BF5DFFFFFF9F947DD58C0E4FFB50" docLanguage="en" docName="ActaEntMusNatPrag.60.2.411-417.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Nalassus (Nalassus) becvari Nabozhenko &amp; Keskin &amp; Papadopoulou 2020, sp. nov." docType="treatment" docVersion="5" lastPageNumber="415" masterDocId="F20FC725FFFBFF9B9550D5510A0AFFBF" masterDocTitle="Two new species and new records of darkling beetles of the tribe Helopini from Turkey and Cyprus (Coleoptera: Tenebrionidae)" masterLastPageNumber="417" masterPageNumber="411" pageNumber="415" updateTime="1698926180628" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="299774976E788F7678E39D2952A29881" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="69A0B84AE5F267B43E7CB22F6490A394">
<mods:title id="2163E607F8309DADA5957E3762E75B19">Two new species and new records of darkling beetles of the tribe Helopini from Turkey and Cyprus (Coleoptera: Tenebrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="96E6A778C3F61C5C5664AA28605D8E06" type="personal">
<mods:role id="B62FB1926BE6CE2B2AB30BBBB5F81E8A">
<mods:roleTerm id="35A9C426C14040A552D4931C6B41AC37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06497F1BB284A9934BCA3268049EB4E7">Nabozhenko, Maxim</mods:namePart>
<mods:affiliation id="D53FC05B81BCCD51BF2D6034DF459764">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan 367000 Russia; e-mail: nalassus @ mail. ru &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia</mods:affiliation>
</mods:name>
<mods:name id="944DDBACA7A4AE724856187BF77A8EF8" type="personal">
<mods:role id="D4ED240611E61EF925D4899F1322B35F">
<mods:roleTerm id="41F8EC0C1C2643CC1148ED53CFD544C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA7359CC3CA3DB1BD312678F00F40FB7">Keskin, Bekir</mods:namePart>
<mods:affiliation id="DFA927F1DE909EE7AA8028021F7D8180">Ege University, Bornova-Izmir 35100 Turkey; e-mail: bekir. keskin. phd @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="1FCAB4ED4D766CE54E65748B4A8918DC" type="personal">
<mods:role id="E59D8EE1E64AF55C1D3CB5B17CC23C3D">
<mods:roleTerm id="BD18014BFBFC92CB0F4D182F283E224B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2BD807A9FEA9946B2076D8F55F7A65D5">Papadopoulou, Anna</mods:namePart>
<mods:affiliation id="D3D46EA47DC5B5DC3281DB4F0F7612C1">University of Cyprus, 1678 Nicosia P. O. Box 20537 Cyprus; e-mail: papadopoulou. g. anna @ ucy. ac. cy</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B5C7DB828ED2B694DC3ABAFD9422758C">text</mods:typeOfResource>
<mods:relatedItem id="E022E7DBFB8303DD34A126781DA0BB69" type="host">
<mods:titleInfo id="E6927DD85B6B30756CF1EFACF20D7D1F">
<mods:title id="355F277AE6828526F75AEBDD94D3A319">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="B32A11B25F284DE50B9F6D9141AEAD66">
<mods:date id="2AE61A46EB7EA74795FDDDA0F6FD4E8B">2020</mods:date>
<mods:detail id="A6FE5DC317267861000FBACC7FEE7468" type="pubDate">
<mods:number id="3E0661057FCDF56DCE8911878CA94E99">2020-06-25</mods:number>
</mods:detail>
<mods:detail id="D0712017E01A7FAFC2004FC9EB999E04" type="volume">
<mods:number id="DE29025F45D3F78D042B330F64559D8A">60</mods:number>
</mods:detail>
<mods:detail id="E9FCF26F415A9DCAEE3870F089E62012" type="issue">
<mods:number id="D8389DE6C5D771532033A4E5175F1450">2</mods:number>
</mods:detail>
<mods:extent id="5504303742B18F17F49400270F057B0D" unit="page">
<mods:start id="9628B145382379D3D67732C3E18200D7">411</mods:start>
<mods:end id="BB6C8F262A1157D4A40A074957D0D845">417</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A7C2C08A5D792502123534B3D7EE2657">journal article</mods:classification>
<mods:identifier id="6CD05B7E3E0E8FD5F99CFC73E1824E8A" type="CLB-Dataset">8112</mods:identifier>
<mods:identifier id="499F72A1A2E3B7EF905C44B14CEEB0A5" type="DOI">10.37520/aemnp.2020.25</mods:identifier>
<mods:identifier id="CC7D88F66A384D80D742013EEC8FF666" type="GBIF-Dataset">bd0522ba-e842-4416-82ac-98640acd8e45</mods:identifier>
<mods:identifier id="DF56F2E90DCDA0C0EC51E7B7AD22E38A" type="ISSN">1804-6487</mods:identifier>
<mods:identifier id="6516093FE242090B87CB9DDC3784B6D5" type="Zenodo-Dep">4489312</mods:identifier>
<mods:identifier id="95DA28EFAD79AB79A42ACB2863518F20" type="ZooBank">94935C98-664E-4D3C-87FA-F3FC6513512B</mods:identifier>
</mods:mods>
<treatment id="0E36BF5DFFFFFF9F947DD58C0E4FFB50" ID-DOI="http://doi.org/10.5281/zenodo.4550757" ID-GBIF-Taxon="177426383" ID-Zenodo-Dep="4550757" LSID="urn:lsid:plazi:treatment:0E36BF5DFFFFFF9F947DD58C0E4FFB50" httpUri="http://treatment.plazi.org/id/0E36BF5DFFFFFF9F947DD58C0E4FFB50" lastPageNumber="415" pageId="4" pageNumber="415">
<subSubSection id="CE855DC0FFFFFF9F947DD58C0870FEAA" pageId="4" pageNumber="415" type="nomenclature">
<paragraph id="86200E4BFFFFFF9F947DD58C0870FEAA" blockId="4.[151,760,221,398]" pageId="4" pageNumber="415">
<heading id="DD68B927FFFFFF9F947DD58C0870FEAA" centered="true" fontSize="10" level="2" pageId="4" pageNumber="415" reason="3">
<emphasis id="B4EBD259FFFFFF9F947DD58C0870FEAA" bold="true" pageId="4" pageNumber="415">
<taxonomicName id="419F75C8FFFFFF9F947DD58C0813FEAA" authority="Nabozhenko &amp; Keskin" authorityName="Nabozhenko &amp; Keskin &amp; Papadopoulou" authorityYear="2020" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="becvari" status="sp. nov." subGenus="Nalassus">
<emphasis id="B4EBD259FFFFFF9F947DD58C0B9AFF4A" bold="true" box="[301,400,221,245]" italics="true" pageId="4" pageNumber="415">Nalassus</emphasis>
(
<emphasis id="B4EBD259FFFFFF9F94CFD58C0808FF4A" bold="true" box="[415,514,221,245]" italics="true" pageId="4" pageNumber="415">Nalassus</emphasis>
)
<emphasis id="B4EBD259FFFFFF9F9742D58C0868FF4A" bold="true" box="[530,610,221,245]" italics="true" pageId="4" pageNumber="415">becvari</emphasis>
Nabozhenko &amp; Keskin
</taxonomicName>
,
<taxonomicNameLabel id="AFD86F22FFFFFF9F9774D5AF0870FEAA" box="[548,634,254,277]" pageId="4" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CE855DC0FFFFFF9F94C6D44D0BF2FE90" box="[406,504,284,303]" pageId="4" pageNumber="415" type="description">
<paragraph id="86200E4BFFFFFF9F94C6D44D0BF2FE90" blockId="4.[151,760,221,398]" box="[406,504,284,303]" pageId="4" pageNumber="415">
(
<figureCitation id="1EA412CEFFFFFF9F94CCD44D0BF8FE90" box="[412,498,284,303]" captionStart="Figs 812" captionStartId="3.[151,188,2036,2055]" captionTargetBox="[185,1391,1085,2009]" captionTargetId="figure-400@3.[185,1395,1063,2022]" captionTargetPageId="3" captionText="Figs 812. Nalassus becvari sp. nov., holotype. 8 male habitus; 9 aedeagus, ventral view; 10 aedeagus, lateral view; 11 male inner sternite VIII; 12 spiculum gastrale, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4489320" httpUri="https://zenodo.org/record/4489320/files/figure.png" pageId="4" pageNumber="415">Figs 812</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CE855DC0FFFFFF9F95C7D4170867FE31" pageId="4" pageNumber="415" type="materials_examined">
<paragraph id="86200E4BFFFFFF9F95C7D4170867FE31" blockId="4.[151,760,221,398]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F95C7D4170B11FEE6" bold="true" box="[151,283,326,345]" pageId="4" pageNumber="415">Type material.</emphasis>
HοΓοτΥΡΕ: ♁ (
<collectionCode id="E08E968EFFFFFF9F94CED4170BD5FEE6" box="[414,479,326,345]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="415">NMPC</collectionCode>
), labelled: Anatol. or. /
<collectingRegion id="445BC0A9FFFFFF9F97FBD41708F2FEE6" box="[683,760,326,345]" country="Turkey" name="Elazig" pageId="4" pageNumber="415">ELAZIG</collectingRegion>
// Coll. Kadlec. The
<typeStatus id="5924B0E9FFFFFF9F9403D4310B94FECC" box="[339,414,352,371]" pageId="4" pageNumber="415" type="holotype">holotype</typeStatus>
was previously deposited in SBCP, but Stanislav Bečvář kindly agreed to transfer it to NMPC.
</paragraph>
</subSubSection>
<subSubSection id="CE855DC0FFFFFF9F95C7D4F70F0AFDD0" pageId="4" pageNumber="415" type="description">
<paragraph id="86200E4BFFFFFF9F95C7D4F70850FE62" blockId="4.[151,761,422,2077]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F95C7D4F70B28FE02" bold="true" box="[151,290,422,445]" pageId="4" pageNumber="415">Description.</emphasis>
Body small (length
<quantity id="4167A3AEFFFFFF9F94A8D4F70841FE02" box="[504,587,422,445]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="4" pageNumber="415" unit="mm" value="6.4">6.4 mm</quantity>
, width
<quantity id="4167A3AEFFFFFF9F97C7D4F708E6FE02" box="[663,748,422,445]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="415" unit="mm" value="2.5">2.5 mm</quantity>
), slender, brown, shiny, moderately convex.
</paragraph>
<paragraph id="86200E4BFFFFFF9F95E5D4B708CAFC42" blockId="4.[151,761,422,2077]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F95E5D4B70AE5FE42" box="[181,239,486,509]" italics="true" pageId="4" pageNumber="415">Head</emphasis>
widest at eye level, 1.52 times as wide as interocular space of frons, covered with recumbent setae on vertex, genae and near eyes. Eyes convex, transversely elongate, bean-shaped. Epistomal anterior margin straight. Outer margin of genae angulate, strongly rounded in basal third and straight in apical two thirds. Lateral margin of head without emargination between genae and epistoma. Punctation of head coarse and dense, puncture diameter 1.52.0 times as long as interpunctural distance; punctures deep, round. Ventral aspect of eye with a weak posterior ventral impression (not groove). Head ventrally with very coarse punctation and subrecumbent pubescence. Apical maxillary palpomeres strongly widened and flattened, securiform, transverse, with rounded anterior margin.Antennae with strongly thickened oval antennomeres 38. Ratio of length/width of antennomeres 211: 0.7/0.6, 1.6/0.8, 1.2/0.9. 1.2/0.9, 1.3/1, 1.2/1, 1.3/1, 1/1, 1/1, 1.2/0.9.
</paragraph>
<paragraph id="86200E4BFFFFFF9F95E5D1570B37F9E2" blockId="4.[151,761,422,2077]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F95E5D1570B20FBA2" box="[181,298,1030,1053]" italics="true" pageId="4" pageNumber="415">Prothorax.</emphasis>
Pronotum transverse (1.32 times as wide as long), widest slightly before middle, 1.54 times as wide as head. Lateral margins weakly rounded, slightly sinuate near posterior angles; anterior margin weakly widely emarginate; base weakly rounded, with short emargination along scutellum.Angles narrowly rounded at apex, anterior ones rectangular, posterior ones weakly obtuse. Lateral margins and base narrowly beaded, anterior margin beaded only near angles. Disc weakly evenly convex, sides narrowly flattened, punctation slightly finer and sparser than on head, sparser in middle and near margins (puncture diameter 24 times as short as interpunctural distance) and denser on sides (puncture diameter subequal or little wider than interpunctural distance). Prosternum with coarse irregular wrinkles (but punctured near procoxae) and erected setae. Prosternal process very weakly convex. Prothoracic hypomera strongly flattened along outer margin, longitudinally wrinkled and vested with long recumbent setae in anterior and basal parts.
</paragraph>
<paragraph id="86200E4BFFFFFF9F95E5D3370E5BFED0" blockId="4.[151,761,422,2077]" lastBlockId="4.[828,1437,184,1263]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F95E5D3370B34F9C2" box="[181,318,1638,1661]" italics="true" pageId="4" pageNumber="415">Pterothorax.</emphasis>
Elytra elongate, oval, widest in middle (1.58 times as long as wide), 1.81 times as wide as head, 1.17 times as wide and 2.46 times as long as pronotum. Strial punctures elongate, merged in interrupted grooves. Interstriae flat, with coarse and sparse punctation (punctures slightly smaller than in striae). Eighth interval not more convex than others and apically connected with second interval. Lateral deflected margin of elytra visible dorsally. Epipleura strongly depressed along whole length, not reaching sutural angle (ended slightly before apex). Mesoventrite with coarse and dense punctation of round punctures and recumbent pubescence. Wings reduced, small. Metaventrite and metepisterna with coarse, not dense punctation (puncture diameter subequal to interpunctural distance) and short recumbent setae. Legs slender; protrochanters with 3 long setae, other trochanters with 1 long seta; femora with erected setae at base of inner side; tibiae straight, margins around apex with long setae (not spines); tarsomeres elongate, not widened, densely pubescent on plantar side.
</paragraph>
<paragraph id="86200E4BFFFFFF9F960AD4290F0AFDD0" blockId="4.[828,1437,184,1263]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F960AD42909CFFE30" box="[858,965,376,399]" italics="true" pageId="4" pageNumber="415">Abdomen.</emphasis>
Abdominal ventrites with coarse, moderately dense punctation (puncture diameter subequal to interpunctural distance) of round punctures and very short recumbent setae, ventrite 1 without brush of long setae; ventrite 5 more finely punctured, with beaded margin. Aedeagus nalassoid, weakly sclerotized, with laterally flattened keel at apex (
<figureCitation id="1EA412CEFFFFFF9F96C3D7690E0BFDF0" box="[915,1025,568,591]" captionStart="Figs 812" captionStartId="3.[151,188,2036,2055]" captionTargetBox="[185,1391,1085,2009]" captionTargetId="figure-400@3.[185,1395,1063,2022]" captionTargetPageId="3" captionText="Figs 812. Nalassus becvari sp. nov., holotype. 8 male habitus; 9 aedeagus, ventral view; 10 aedeagus, lateral view; 11 male inner sternite VIII; 12 spiculum gastrale, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4489320" httpUri="https://zenodo.org/record/4489320/files/figure.png" pageId="4" pageNumber="415">Figs 910</figureCitation>
). Male inner sternite VIII and spiculum gastrale typical for
<taxonomicName id="419F75C8FFFFFF9F915AD7090E60FDD0" authorityName="Mulsant" authorityYear="1854" box="[1034,1130,600,623]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B4EBD259FFFFFF9F915AD7090E60FDD0" box="[1034,1130,600,623]" italics="true" pageId="4" pageNumber="415">Nalassus</emphasis>
</taxonomicName>
(
<figureCitation id="1EA412CEFFFFFF9F9129D7090EFEFDD0" box="[1145,1268,600,623]" captionStart="Figs 812" captionStartId="3.[151,188,2036,2055]" captionTargetBox="[185,1391,1085,2009]" captionTargetId="figure-400@3.[185,1395,1063,2022]" captionTargetPageId="3" captionText="Figs 812. Nalassus becvari sp. nov., holotype. 8 male habitus; 9 aedeagus, ventral view; 10 aedeagus, lateral view; 11 male inner sternite VIII; 12 spiculum gastrale, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4489320" httpUri="https://zenodo.org/record/4489320/files/figure.png" pageId="4" pageNumber="415">Figs 1112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CE855DC0FFFFFF9F966CD7290E5CFBD0" pageId="4" pageNumber="415" type="diagnosis">
<paragraph id="86200E4BFFFFFF9F966CD7290E5CFBD0" blockId="4.[828,1437,184,1263]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F966CD7290E25FD30" bold="true" box="[828,1071,632,655]" pageId="4" pageNumber="415">Differential diagnosis.</emphasis>
With strongly depressed, epipleura almost reaching apex, the species is externally similar to
<taxonomicName id="419F75C8FFFFFF9F966CD7E90ECEFD70" authority="Nabozhenko, 2001" authorityName="Nabozhenko" authorityYear="2001" box="[828,1220,696,719]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B4EBD259FFFFFF9F966CD7E909F9FD70" box="[828,1011,696,719]" italics="true" pageId="4" pageNumber="415">Nalassus kaszabi</emphasis>
<bibRefCitation id="E20E73BAFFFFFF9F96AAD7E90ECEFD70" author="NABOZHENKO M. V." box="[1018,1220,696,719]" pageId="4" pageNumber="415" pagination="113 - 117" refId="ref5164" refString="NABOZHENKO M. V. 2001: Taxonomic notes on the genus Zophohelops Reitter, 1901 with description of new species from Tadzhikistan and new genus Pseudoprobaticus gen. n. (Coleoptera, Tenebrionidae). Annales Zoologici 51: 113 - 117." type="journal article" year="2001">Nabozhenko, 2001</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="445BC0A9FFFFFF9F9056D7E90F93FD70" box="[1286,1433,696,719]" country="Turkey" name="Van" pageId="4" pageNumber="415">Van Province</collectingRegion>
, which was transferred to the subgenus
<taxonomicName id="419F75C8FFFFFF9F9194D789097BFCB0" authority="Reitter, 1922" authorityName="Reitter" authorityYear="1922" class="Insecta" family="Tenebrionidae" genus="Helopondrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B4EBD259FFFFFF9F9194D7890F46FD50" box="[1220,1356,728,751]" italics="true" pageId="4" pageNumber="415">Helopondrus</emphasis>
Reitter, 1922
</taxonomicName>
on the basis of preliminary analysis of COI markers (KΕΚΙΝ et al. 2017b). On the other hand, males of
<taxonomicName id="419F75C8FFFFFF9F90D4D6490981FCF0" authorityName="Nabozhenko" authorityYear="2001" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B4EBD259FFFFFF9F90D4D6490981FCF0" italics="true" pageId="4" pageNumber="415">N. kaszabi</emphasis>
</taxonomicName>
are unknown, and we cannot support our molecular data by the structure of male genitalia.
<taxonomicName id="419F75C8FFFFFF9F91B4D6090F96FCD0" authority="Nabozhenko &amp; Keskin &amp; Papadopoulou, 2020" authorityName="Nabozhenko &amp; Keskin &amp; Papadopoulou" authorityYear="2020" box="[1252,1436,856,879]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="becvari" status="sp. nov.">
<emphasis id="B4EBD259FFFFFF9F91B4D6090F96FCD0" box="[1252,1436,856,879]" italics="true" pageId="4" pageNumber="415">Nalassus becvari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFD86F22FFFFFF9F966CD6290987FC30" box="[828,909,888,911]" pageId="4" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="419F75C8FFFFFF9F9144D6290E8BFC30" authorityName="Nabozhenko" authorityYear="2001" box="[1044,1153,888,911]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="species" species="kaszabi">
<emphasis id="B4EBD259FFFFFF9F9144D6290E8BFC30" box="[1044,1153,888,911]" italics="true" pageId="4" pageNumber="415">N. kaszabi</emphasis>
</taxonomicName>
in narrowly flattened sides along lateral margins of pronotum and epipleura not reaching elytral sutural angles. The new species differs from all other Turkish species of
<taxonomicName id="419F75C8FFFFFF9F9165D6890E99FC50" authorityName="Mulsant" authorityYear="1854" box="[1077,1171,984,1007]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B4EBD259FFFFFF9F9165D6890E99FC50" box="[1077,1171,984,1007]" italics="true" pageId="4" pageNumber="415">Nalassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFD86F22FFFFFF9F91C8D6890EC7FC50" box="[1176,1229,984,1007]" pageId="4" pageNumber="415" sensu="stricto">s. str.</taxonomicNameLabel>
in the ventral aspect of eye, which bears a weak posterior ventral impression (rather than a distinct groove of other species). This new species can be distinguished from other Turkish
<taxonomicName id="419F75C8FFFFFF9F906CD1690F96FBF0" authorityName="Mulsant" authorityYear="1854" box="[1340,1436,1080,1103]" class="Insecta" family="Tenebrionidae" genus="Nalassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="415" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B4EBD259FFFFFF9F906CD1690F96FBF0" box="[1340,1436,1080,1103]" italics="true" pageId="4" pageNumber="415">Nalassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFD86F22FFFFFF9F966CD109097EFBD0" box="[828,884,1112,1135]" pageId="4" pageNumber="415" sensu="stricto">s. str.</taxonomicNameLabel>
using the key below.
</paragraph>
</subSubSection>
<subSubSection id="CE855DC0FFFFFF9F966CD1290E4FFB50" pageId="4" pageNumber="415" type="etymology">
<paragraph id="86200E4BFFFFFF9F966CD1290E4FFB50" blockId="4.[828,1437,184,1263]" pageId="4" pageNumber="415">
<emphasis id="B4EBD259FFFFFF9F966CD12909B6FB30" bold="true" box="[828,956,1144,1167]" pageId="4" pageNumber="415">Etymology.</emphasis>
The species is named in honour of our colleague, Stanislav Bečvář, who transferred to us many interesting darkling beetles for study. The name is a noun in singular genitive case.
</paragraph>
</subSubSection>
</treatment>
</document>