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<document id="9C9753C9376652A6D59027739564DEC4" ID-DOI="10.5281/zenodo.4504448" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504448" ID-ZooBank="37CF3BF7-BE06-4D4C-B0F5-F5E648108351" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612466381302" checkinUser="carolina" docAuthor="Larson, Helen K., Hadiaty, Renny K. &amp; Hubert, Nicolas" docDate="2017" docId="EA3EC41AE225FFF6FC02F964FACDF9D8" docLanguage="en" docName="RafflesBZool.65.175-180.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Pseudogobiopsis lumbantobing Larson &amp; Hadiaty &amp; Hubert 2017, n. sp." docType="treatment" docUuid="B1C88B97-C30A-40A3-9AC8-709E0A22A155" docUuidSource="ZooBank" docVersion="5" lastPageNumber="179" masterDocId="1607BC62E225FFF2FFB1FFECFF9CFFC1" masterDocTitle="A new species of the gobiid fish g enus Pseudogobiopsis (Teleostei, Gobiidae, Gobionellinae) from Indonesia" masterLastPageNumber="180" masterPageNumber="175" pageNumber="175" updateTime="1698866215998" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="394D46D80536D4AE577DE3B5D15BF55C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BB57FA5EB5A8528458AB95AC35A570CD">
<mods:title id="4C2A96013D8979FD363D64F756F3DD22">A new species of the gobiid fish g enus Pseudogobiopsis (Teleostei, Gobiidae, Gobionellinae) from Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="6FE0B7071759D95C9116426E286CC3A3" type="personal">
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</mods:role>
<mods:namePart id="8C7B8A044069C801CBB4F98CFAFC9531">Larson, Helen K.</mods:namePart>
</mods:name>
<mods:name id="7F1574143068621F14BE6D0C713934BA" type="personal">
<mods:role id="437C4E659A49C406471AA2B0E5DF2F95">
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</mods:role>
<mods:namePart id="3EFAA33F33856D03F5FFDB99829AD473">Hadiaty, Renny K.</mods:namePart>
</mods:name>
<mods:name id="F1E3CD93CB6261E437D9C6371CED05BE" type="personal">
<mods:role id="093D40E230F96E11A51EB1C449621B9D">
<mods:roleTerm id="537F00EAB377C6AED8DAD1F12E22E15C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFC11F0902ABEE6AF1457C45F78E935D">Hubert, Nicolas</mods:namePart>
</mods:name>
<mods:typeOfResource id="22A7A0475ACB62DB6AF6CBC581FBBD15">text</mods:typeOfResource>
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<mods:titleInfo id="47E24F9B88474DE0FC57C53D002DBAFC">
<mods:title id="D3ACC6768AB7FDCBEA71AD513E791DE0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="9978425BD5F42A781985ABDC52DFC900">
<mods:date id="28547265F2F455B8BE1BDDB80DDAE10D">2017</mods:date>
<mods:detail id="7984237F1402FBFD5C097E88C8C7A3A8" type="pubDate">
<mods:number id="E11B1858486FC7B93A9B2941995C3D54">2017-06-01</mods:number>
</mods:detail>
<mods:detail id="54322B9DA91450902887EF5535DA35CE" type="volume">
<mods:number id="5DE0A1CF66BF02712F5CFDE9AAB07DE5">65</mods:number>
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<mods:start id="55C79C427C5B139B9F6FD47DEF104373">175</mods:start>
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<mods:classification id="C1E180F9B31D3B76102625BE314697D7">journal article</mods:classification>
<mods:identifier id="29FF2FFFB5B90DA75B596FD2B8CDA74F" type="DOI">10.5281/zenodo.4504448</mods:identifier>
<mods:identifier id="F983DE87439FB478F09E376E57BE7641" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A4DBCBBD0436D4AB1EB2F6885F4A45D6" type="Zenodo-Dep">4504448</mods:identifier>
<mods:identifier id="D55414E95063083F3F42824053B0F0DD" type="ZooBank">37CF3BF7-BE06-4D4C-B0F5-F5E648108351</mods:identifier>
</mods:mods>
<treatment id="EA3EC41AE225FFF6FC02F964FACDF9D8" LSID="urn:lsid:zoobank.org:act:B1C88B97-C30A-40A3-9AC8-709E0A22A155" httpUri="http://treatment.plazi.org/id/EA3EC41AE225FFF6FC02F964FACDF9D8" lastPageId="4" lastPageNumber="179" pageId="0" pageNumber="175">
<subSubSection id="2A8D2687E225FFF2FC02F964FA0EF91C" pageId="0" pageNumber="175" type="nomenclature">
<paragraph id="6228750CE225FFF2FC02F964FACEF960" blockId="0.[947,1362,1672,1697]" box="[947,1362,1672,1697]" pageId="0" pageNumber="175">
<heading id="3960C260E225FFF2FC02F964FACEF960" bold="true" box="[947,1362,1672,1697]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="175" reason="2">
<emphasis id="50E3A91EE225FFF2FC02F964FACEF960" bold="true" box="[947,1362,1672,1697]" pageId="0" pageNumber="175">
<taxonomicName id="A5970E8FE225FFF2FC02F964FA90F961" authority="Larson &amp; Hadiaty &amp; Hubert, 2017" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[947,1292,1672,1696]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="175" phylum="Chordata" rank="species" species="lumbantobing" status="sp. nov.">
<emphasis id="50E3A91EE225FFF2FC02F964FA90F961" bold="true" box="[947,1292,1672,1696]" italics="true" pageId="0" pageNumber="175">Pseudogobiopsis lumbantobing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BD01465E225FFF2FAA4F965FACEF960" box="[1301,1362,1673,1697]" pageId="0" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6228750CE225FFF2FCF2F92BFA0EF91C" blockId="0.[835,1426,1735,1757]" box="[835,1426,1735,1757]" pageId="0" pageNumber="175">
Gen. &amp; spec. undet. –
<bibRefCitation id="060608FDE225FFF2FB92F92BFB48F91D" author="Roberts TR" box="[1059,1236,1735,1757]" pageId="0" pageNumber="175" pagination="1 - 94" refId="ref4872" refString="Roberts TR (1993) The freshwater fishes of Java, as observed by Kuhl and van Hasselt in 1820 - 23. Zoologische Verhandelingen, 285: 1 - 94." type="journal article" year="1993">Roberts, 1993: 44</bibRefCitation>
, Fig. 51 (Tjisekat).
</paragraph>
</subSubSection>
<subSubSection id="2A8D2687E225FFF3FCF2F8E9FD6CFB14" lastPageId="1" lastPageNumber="176" pageId="0" pageNumber="175" type="materials_examined">
<paragraph id="6228750CE225FFF3FCF2F8E9FE1FFBD4" blockId="0.[835,1475,1797,2109]" lastBlockId="1.[113,753,189,1046]" lastPageId="1" lastPageNumber="176" pageId="0" pageNumber="175">
<emphasis id="50E3A91EE225FFF2FCF2F8E9FBBEF8DC" bold="true" box="[835,1058,1797,1821]" pageId="0" pageNumber="175">Material examined.</emphasis>
<materialsCitation id="D2FF7F51E225FFF2FB98F8E9FABDF89C" accessionNumber="MZB 23017" collectingDate="2006-07-13" collectionCode="MZB" collectorName="Sungei Kluet &amp; Sumatra &amp; Indonesia &amp; D. Lumbantobing" pageId="0" pageNumber="175" specimenCount="1" typeStatus="holotype">
<typeStatus id="BD2CCBAEE225FFF2FB98F8E9FB2AF8DC" box="[1065,1206,1797,1821]" pageId="0" pageNumber="175" type="holotype">HOLOTYPE</typeStatus>
–
<collectionCode id="0486EDC9E225FFF2FB60F8EAFA97F8DC" box="[1233,1291,1798,1821]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="175" type="Museum">MZB</collectionCode>
23017,
<quantity id="A56FD8E9E225FFF2FAD2F8E9FA5EF8DC" box="[1379,1474,1797,1821]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="0" pageNumber="175" unit="mm" value="29.5">29.5 mm</quantity>
SL male,
<collectorName id="CF6210DAE225FFF2FC1BF8C9FBA5F8FC" box="[938,1081,1829,1853]" pageId="0" pageNumber="175">Sungei Kluet</collectorName>
, northwest
<collectorName id="CF6210DAE225FFF2FB02F8C9FA93F8FC" box="[1203,1295,1829,1853]" pageId="0" pageNumber="175">Sumatra</collectorName>
,
<collectorName id="CF6210DAE225FFF2FAA8F8C9FA1FF8FC" box="[1305,1411,1829,1853]" pageId="0" pageNumber="175">Indonesia</collectorName>
, BDS 17,
<collectorName id="CF6210DAE225FFF2FCDBF8AAFBB9F89C" box="[874,1061,1861,1885]" pageId="0" pageNumber="175">D. Lumbantobing</collectorName>
and party,
<date id="162953CCE225FFF2FB29F8A9FABDF89C" box="[1176,1313,1861,1885]" pageId="0" pageNumber="175" value="2006-07-13">
<collectingDate id="066DAA24E225FFF2FB29F8A9FABDF89C" box="[1176,1313,1861,1885]" pageId="0" pageNumber="175" value="2006-07-13">13 July 2006</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="D2FF7F51E225FFF2FA9BF8A9FCE1F81C" accessionNumber="MZB 23018" collectingDate="2006-07-23" collectionCode="MZB" collectorName="D. Lumbantobing" country="Indonesia" county="Batu Garirgis River" location="Kabupaten Tapanuli Tengah" municipality="Barus" pageId="0" pageNumber="175" specimenCode="14" specimenCount="1" stateProvince="Sumatera Barat" typeStatus="paratype">
<typeStatus id="BD2CCBAEE225FFF2FA9BF8A9FA5EF89C" box="[1322,1474,1861,1885]" pageId="0" pageNumber="175" type="paratype">PARATYPES</typeStatus>
–
<collectingCountry id="1A80359CE225FFF2FCEAF889FC6EF8BC" box="[859,1010,1893,1917]" name="Indonesia" pageId="0" pageNumber="175">INDONESIA</collectingCountry>
:
<collectionCode id="0486EDC9E225FFF2FC4EF88AFBA5F8BC" box="[1023,1081,1894,1917]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="175" type="Museum">MZB</collectionCode>
23018,
<specimenCode id="3231DD77E225FFF2FB26F889FB2EF8BC" box="[1175,1202,1893,1917]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="175" type="Museum">14</specimenCode>
(16–33.5),
<collectingCounty id="8B490D80E225FFF2FA9CF88AFC1DF85C" pageId="0" pageNumber="175">Batu Garirgis River</collectingCounty>
(near
<collectingMunicipality id="824CEF76E225FFF2FC63F869FB84F85C" box="[978,1048,1925,1949]" pageId="0" pageNumber="175">Barus</collectingMunicipality>
),
<location id="674823D7E225FFF2FB9EF869FAE6F85C" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E225FFF2FB9EF869FAE6F85C" box="[1071,1402,1925,1949]" country="Indonesia" county="Batu Garirgis River" municipality="Barus" name="Kabupaten Tapanuli Tengah" pageId="0" pageNumber="175" stateProvince="Sumatera Barat">Kabupaten Tapanuli Tengah</location>
,
<collectingRegion id="A053BBEEE225FFF2FA38F86AFC3EF87C" country="Indonesia" name="Sumatera Barat" pageId="0" pageNumber="175">West Sumatra</collectingRegion>
, BDS 35,
<collectorName id="CF6210DAE225FFF2FBAFF84AFB7AF87C" box="[1054,1254,1957,1981]" pageId="0" pageNumber="175">D. Lumbantobing</collectorName>
and party,
<date id="162953CCE225FFF2FADFF849FCE1F81C" pageId="0" pageNumber="175" value="2006-07-23">
<collectingDate id="066DAA24E225FFF2FADFF849FCE1F81C" pageId="0" pageNumber="175" value="2006-07-23">23 July 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E225FFF2FC3DF829FA14F7DC" collectingDate="2006-07-23" collectionCode="USNM" collectorName="D. Lumbantobing" country="Indonesia" county="Batu Garirgis River" location="Kabupaten Tapanuli Tengah" municipality="Barus" pageId="0" pageNumber="175" specimenCode="USNM 432514, 14" specimenCount="1" stateProvince="Sumatera Barat" typeStatus="paratype">
<specimenCode id="3231DD77E225FFF2FC3DF829FBA6F81C" box="[908,1082,1989,2013]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="175" type="Museum">USNM 432514</specimenCode>
,
<specimenCode id="3231DD77E225FFF2FBF8F829FBF8F81C" box="[1097,1124,1989,2013]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="0" pageNumber="175" type="Museum">14</specimenCode>
(16–33.5),
<collectingCounty id="8B490D80E225FFF2FB53F82AFA5FF81C" box="[1250,1475,1989,2013]" pageId="0" pageNumber="175">Batu Garirgis River</collectingCounty>
(near
<collectingMunicipality id="824CEF76E225FFF2FC35F809FC5AF83C" box="[900,966,2021,2045]" pageId="0" pageNumber="175">Barus</collectingMunicipality>
),
<location id="674823D7E225FFF2FC6AF809FA88F83C" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E225FFF2FC6AF809FA88F83C" box="[987,1300,2021,2045]" country="Indonesia" county="Batu Garirgis River" municipality="Barus" name="Kabupaten Tapanuli Tengah" pageId="0" pageNumber="175" stateProvince="Sumatera Barat">Kabupaten Tapanuli Tengah</location>
,
<collectingRegion id="A053BBEEE225FFF2FA93F80AFA23F83C" box="[1314,1471,2021,2045]" country="Indonesia" name="Sumatera Barat" pageId="0" pageNumber="175">West Sumatra</collectingRegion>
, BDS 35,
<collectorName id="CF6210DAE225FFF2FC1CF7EAFBECF7DC" box="[941,1136,2053,2077]" pageId="0" pageNumber="175">D. Lumbantobing</collectorName>
and party,
<date id="162953CCE225FFF2FB42F7E9FA14F7DC" box="[1267,1416,2053,2077]" pageId="0" pageNumber="175" value="2006-07-23">
<collectingDate id="066DAA24E225FFF2FB42F7E9FA14F7DC" box="[1267,1416,2053,2077]" pageId="0" pageNumber="175" value="2006-07-23">23 July 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E225FFF3FA26F7E9FE6DFF34" collectingDate="2013-12-07" collectionCode="QM" collectorName="N. Hubert &amp; F. Busson &amp; S. Sauri" country="Indonesia" lastPageId="1" lastPageNumber="176" location="Kabupaten Pandeglang" municipality="Cibeber" pageId="0" pageNumber="175" specimenCount="1" stateProvince="Banten" typeStatus="paratype">
<collectionCode id="0486EDC9E225FFF2FA26F7E9FA5EF7DC" box="[1431,1474,2053,2077]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="0" pageNumber="175" type="Museum">QM</collectionCode>
I.39794 (ex MZB-BIF 1437–1438), 2(27–28.5),
<collectingMunicipality id="824CEF76E225FFF2FAD4F7C9FA22F7FC" box="[1381,1470,2085,2109]" pageId="0" pageNumber="175">Cibeber</collectingMunicipality>
,
<location id="674823D7E224FFF3FFC0FF51FEFBFF14" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FFC0FF51FEFBFF14" box="[113,359,189,213]" country="Indonesia" municipality="Cibeber" name="Kabupaten Pandeglang" pageId="1" pageNumber="176" stateProvince="Banten">Kabupaten Pandeglang</location>
,
<collectingRegion id="A053BBEEE224FFF3FEC0FF52FDBEFF14" box="[369,546,189,213]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten Province</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FD9AFF52FDC1FF14" box="[555,605,190,213]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="176">Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FDD7FF52FD4DFF14" box="[614,721,189,213]" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FD6BFF52FF5EFF34" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<collectorName id="CF6210DAE224FFF3FF7CFF31FEB9FF34" box="[205,293,221,245]" pageId="1" pageNumber="176">S. Sauri</collectorName>
,
<date id="162953CCE224FFF3FE9EFF32FE6DFF34" box="[303,497,221,245]" pageId="1" pageNumber="176" value="2013-12-07">
<collectingDate id="066DAA24E224FFF3FE9EFF32FE6DFF34" box="[303,497,221,245]" pageId="1" pageNumber="176" value="2013-12-07">7 December 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FE4AFF32FE25FE94" collectingDate="2013-12-07" collectionCode="MNHN" collectorName="N. Hubert &amp; F. Busson &amp; S. Sauri" country="Indonesia" location="Kabupaten Pandeglang" municipality="Cibeber" pageId="1" pageNumber="176" specimenCode="MNHN 2016-0195, 96" specimenCount="1" stateProvince="Banten" typeStatus="paratype">
<specimenCode id="3231DD77E224FFF3FE4AFF32FD5BFF34" box="[507,711,221,245]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="176">MNHN 2016-0195</specimenCode>
–
<specimenCode id="3231DD77E224FFF3FD64FF31FD73FF34" box="[725,751,221,245]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="176">96</specimenCode>
(ex MZB-BIF 1439–1440), 2(26–27),
<collectingMunicipality id="824CEF76E224FFF3FDA6FF11FDF2FED4" box="[535,622,253,277]" pageId="1" pageNumber="176">Cibeber</collectingMunicipality>
,
<location id="674823D7E224FFF3FDCDFF11FF6CFEF4" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FDCDFF11FF6CFEF4" country="Indonesia" municipality="Cibeber" name="Kabupaten Pandeglang" pageId="1" pageNumber="176" stateProvince="Banten">Kabupaten Pandeglang</location>
,
<collectingRegion id="A053BBEEE224FFF3FF4CFEF2FE28FEF4" box="[253,436,285,309]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten Province</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FE71FEF2FE6EFEF4" box="[448,498,286,309]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="176">Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FE4EFEF2FDF3FEF4" box="[511,623,285,309]" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FDCDFEF2FD77FEF4" box="[636,747,286,309]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<collectorName id="CF6210DAE224FFF3FFC0FED1FF48FE94" box="[113,212,317,341]" pageId="1" pageNumber="176">S. Sauri</collectorName>
,
<date id="162953CCE224FFF3FF55FED2FE25FE94" box="[228,441,317,341]" pageId="1" pageNumber="176" value="2013-12-07">
<collectingDate id="066DAA24E224FFF3FF55FED2FE25FE94" box="[228,441,317,341]" pageId="1" pageNumber="176" value="2013-12-07">7 December 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FE7BFED2FDEAFE54" collectingDate="2013-12-07" collectorName="N. Hubert &amp; F. Busson &amp; S. Sauri" country="Indonesia" location="Kabupaten Pandeglang" municipality="Cikareo" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Banten" typeStatus="paratype">
MZB-BIF 1559, 1(27.5),
<collectingMunicipality id="824CEF76E224FFF3FFC3FEB1FF55FEB4" box="[114,201,349,373]" pageId="1" pageNumber="176">Cikareo</collectingMunicipality>
,
<location id="674823D7E224FFF3FF64FEB1FE52FEB4" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FF64FEB1FE52FEB4" box="[213,462,349,373]" country="Indonesia" municipality="Cikareo" name="Kabupaten Pandeglang" pageId="1" pageNumber="176" stateProvince="Banten">Kabupaten Pandeglang</location>
,
<collectingRegion id="A053BBEEE224FFF3FE6BFEB2FD12FEB4" box="[474,654,349,373]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten Province</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FD28FEB2FD57FEB4" box="[665,715,350,373]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="176">Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FD67FEB2FF22FE54" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FF7AFE92FEA7FE54" box="[203,315,382,405]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<collectorName id="CF6210DAE224FFF3FEF8FE91FE3EFE54" box="[329,418,381,405]" pageId="1" pageNumber="176">S. Sauri</collectorName>
,
<date id="162953CCE224FFF3FE01FE92FDEAFE54" box="[432,630,381,405]" pageId="1" pageNumber="176" value="2013-12-07">
<collectingDate id="066DAA24E224FFF3FE01FE92FDEAFE54" box="[432,630,381,405]" pageId="1" pageNumber="176" value="2013-12-07">7 December 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FD32FE92FD35FE14" collectingDate="2015-05-20" collectorName="N. Hubert &amp; F. Busson &amp; S. Sauri" country="Indonesia" location="Cicamara" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Banten" typeStatus="paratype">
MZB-BIF 4138–4140, 3(18–21),
<location id="674823D7E224FFF3FED9FE71FE4DFE74" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FED9FE71FE4DFE74" box="[360,465,413,437]" country="Indonesia" name="Cicamara" pageId="1" pageNumber="176" stateProvince="Banten">Cicamara</location>
,
<collectingRegion id="A053BBEEE224FFF3FE6CFE72FDB7FE74" box="[477,555,414,437]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FD86FE72FD70FE74" box="[567,748,413,437]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten Province</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FFC0FE52FF3FFE14" box="[113,163,446,469]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="176">Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FF01FE52FE83FE14" box="[176,287,445,469]" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FE9AFE52FE06FE14" box="[299,410,446,469]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<collectorName id="CF6210DAE224FFF3FE16FE51FE60FE14" box="[423,508,445,469]" pageId="1" pageNumber="176">S. Sauri</collectorName>
,
<date id="162953CCE224FFF3FDA5FE51FD35FE14" box="[532,681,445,470]" pageId="1" pageNumber="176" value="2015-05-20">
<collectingDate id="066DAA24E224FFF3FDA5FE51FD35FE14" box="[532,681,445,470]" pageId="1" pageNumber="176" value="2015-05-20">20 May 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FD07FE51FEF8FDF4" collectingDate="2015-05-20" collectorName="N. Hubert &amp; F. Busson &amp; S. Sauri" country="Indonesia" location="Cicamara" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Banten" typeStatus="paratype">
AMS I.47145-001, (ex MZB-BIF 4141), 1(20.5),
<location id="674823D7E224FFF3FDCAFE31FD76FE34" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FDCAFE31FD76FE34" box="[635,746,477,501]" country="Indonesia" name="Cicamara" pageId="1" pageNumber="176" stateProvince="Banten">Cicamara</location>
,
<collectingRegion id="A053BBEEE224FFF3FFC3FE12FF5DFDD4" box="[114,193,510,533]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FF61FE12FE16FDD4" box="[208,394,509,533]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten Province</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FE29FE12FE57FDD4" box="[408,459,510,533]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="176">Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FE68FE12FDD0FDD4" box="[473,588,509,533]" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FDEBFE12FD50FDD4" box="[602,716,510,533]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<collectorName id="CF6210DAE224FFF3FD6AFE11FF30FDF4" pageId="1" pageNumber="176">S. Sauri</collectorName>
,
<date id="162953CCE224FFF3FF79FDF1FEF8FDF4" box="[200,356,541,566]" pageId="1" pageNumber="176" value="2015-05-20">
<collectingDate id="066DAA24E224FFF3FF79FDF1FEF8FDF4" box="[200,356,541,566]" pageId="1" pageNumber="176" value="2015-05-20">20 May 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FEC5FDF2FDDBFDB4" collectingDate="2015-05-20" collectionCode="MNHN" collectorName="N. Hubert &amp; F. Busson &amp; S. Sauri" country="Indonesia" location="Cicamara" pageId="1" pageNumber="176" specimenCode="MNHN 2016-0197" specimenCount="1" stateProvince="Banten" typeStatus="paratype">
<specimenCode id="3231DD77E224FFF3FEC5FDF2FDD5FDF4" box="[372,585,541,565]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="176">MNHN 2016-0197</specimenCode>
(ex MZB-BIF 4138), 1(20.5),
<location id="674823D7E224FFF3FEA7FDD1FEE3FD94" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FEA7FDD1FEE3FD94" box="[278,383,573,597]" country="Indonesia" name="Cicamara" pageId="1" pageNumber="176" stateProvince="Banten">Cicamara</location>
,
<collectingRegion id="A053BBEEE224FFF3FE39FDD2FE4AFD94" box="[392,470,574,597]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FE6EFDD2FD0EFD94" box="[479,658,573,597]" country="Indonesia" name="Banten" pageId="1" pageNumber="176">Banten Province</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FD2AFDD2FD51FD94" box="[667,717,574,597]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="176">Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FD67FDD2FF22FDB4" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FF7BFDB2FEA5FDB4" box="[202,313,606,629]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<collectorName id="CF6210DAE224FFF3FEF4FDB1FE07FDB4" box="[325,411,605,629]" pageId="1" pageNumber="176">S. Sauri</collectorName>
,
<date id="162953CCE224FFF3FE03FDB1FDDBFDB4" box="[434,583,605,630]" pageId="1" pageNumber="176" value="2015-05-20">
<collectingDate id="066DAA24E224FFF3FE03FDB1FDDBFDB4" box="[434,583,605,630]" pageId="1" pageNumber="176" value="2015-05-20">20 May 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FDE3FDB1FF7FFD14" collectingDate="2015-05-23" collectorName="H. Darhuddin &amp; N. Hubert &amp; F. Busson" country="Indonesia" location="Barat" municipality="Wai Bambang" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Lampung" typeStatus="paratype">
AMS I.47144- 001, (ex MZB-BIF 4229), 1(33),
<collectingMunicipality id="824CEF76E224FFF3FE6EFD91FDE1FD54" box="[479,637,637,661]" pageId="1" pageNumber="176">Wai Bambang</collectingMunicipality>
,
<collectingRegion id="A053BBEEE224FFF3FD38FD92FD73FD57" box="[649,751,638,662]" country="Indonesia" name="Lampung" pageId="1" pageNumber="176">Lampung</collectingRegion>
<location id="674823D7E224FFF3FFC0FD72FF32FD74" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FFC0FD72FF32FD74" box="[113,174,670,693]" country="Indonesia" municipality="Wai Bambang" name="Barat" pageId="1" pageNumber="176" stateProvince="Lampung">Barat</location>
,
<collectingRegion id="A053BBEEE224FFF3FF0CFD71FE86FD74" box="[189,282,669,693]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="176">Sumatra</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FE98FD72FE58FD74" box="[297,452,669,693]" pageId="1" pageNumber="176">H. Darhuddin</collectorName>
,
<collectorName id="CF6210DAE224FFF3FE63FD72FDD9FD74" box="[466,581,669,693]" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FDE2FD72FD5AFD74" box="[595,710,670,693]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<date id="162953CCE224FFF3FD65FD71FF7FFD14" pageId="1" pageNumber="176" value="2015-05-23">
<collectingDate id="066DAA24E224FFF3FD65FD71FF7FFD14" pageId="1" pageNumber="176" value="2015-05-23">23 May 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FF5EFD51FECAFCD4" accessionNumber="ZRC 54925" collectingDate="2015-11-22" collectionCode="ZRC" country="Indonesia" location="Sungai Tumbuan" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Bengkulu" typeStatus="paratype">
<collectionCode id="0486EDC9E224FFF3FF5EFD51FEBFFD14" box="[239,291,701,725]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="176">ZRC</collectionCode>
54925 (ex MZB-BIF 4856–57), 2(27–33),
<location id="674823D7E224FFF3FFC0FD31FEAEFD34" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FFC0FD31FEAEFD34" box="[113,306,733,757]" country="Indonesia" name="Sungai Tumbuan" pageId="1" pageNumber="176" stateProvince="Bengkulu">Sungai Tumbuan</location>
,
<collectingRegion id="A053BBEEE224FFF3FEF0FD31FE31FD34" box="[321,429,733,757]" country="Indonesia" name="Bengkulu" pageId="1" pageNumber="176">Bengkulu</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FE0DFD31FD1CFD34" box="[444,640,733,757]" country="Indonesia" name="Sumatera Selatan" pageId="1" pageNumber="176">Sumatera Selatan</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FD3FFD31FD77FD34" box="[654,747,733,757]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="176">Sumatra</collectingRegion>
,
<date id="162953CCE224FFF3FFC3FD11FECAFCD4" box="[114,342,765,789]" pageId="1" pageNumber="176" value="2015-11-22">
<collectingDate id="066DAA24E224FFF3FFC3FD11FECAFCD4" box="[114,342,765,789]" pageId="1" pageNumber="176" value="2015-11-22">22 November 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FED7FD12FED2FC94" collectingDate="2015-11-22" country="Indonesia" location="Sungai Tumbuan" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Bengkulu" typeStatus="paratype">
MZB-BIF 4858, 60-61, 3(24–33),
<location id="674823D7E224FFF3FFC3FCF1FEAFFCF4" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FFC3FCF1FEAFFCF4" box="[114,307,797,821]" country="Indonesia" name="Sungai Tumbuan" pageId="1" pageNumber="176" stateProvince="Bengkulu">Sungai Tumbuan</location>
,
<collectingRegion id="A053BBEEE224FFF3FEF0FCF1FE31FCF4" box="[321,429,797,821]" country="Indonesia" name="Bengkulu" pageId="1" pageNumber="176">Bengkulu</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FE0DFCF1FD1DFCF4" box="[444,641,797,821]" country="Indonesia" name="Sumatera Selatan" pageId="1" pageNumber="176">Sumatera Selatan</collectingRegion>
,
<collectingRegion id="A053BBEEE224FFF3FD3EFCF1FD70FCF4" box="[655,748,797,821]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="176">Sumatra</collectingRegion>
,
<date id="162953CCE224FFF3FFC3FCD1FED2FC94" box="[114,334,829,853]" pageId="1" pageNumber="176" value="2015-11-22">
<collectingDate id="066DAA24E224FFF3FFC3FCD1FED2FC94" box="[114,334,829,853]" pageId="1" pageNumber="176" value="2015-11-22">22 November 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FEEDFCD1FE96FC54" accessionNumber="ZRC 55524" collectionCode="ZRC" collectorName="Kabupaten &amp; Singkil" country="Indonesia" location="Trumon" municipality="Subulussalam-Singkil" pageId="1" pageNumber="176" specimenCode="3" specimenCount="1" stateProvince="Aceh" typeStatus="paratype">
<collectionCode id="0486EDC9E224FFF3FEEDFCD1FE0CFC94" box="[348,400,829,853]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="176">ZRC</collectionCode>
55524,
<specimenCode id="3231DD77E224FFF3FE41FCD1FE60FC94" box="[496,508,829,853]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="176">3</specimenCode>
(28.0–31), hill stream along road
<collectingMunicipality id="824CEF76E224FFF3FF56FCB1FE48FCB4" box="[231,468,861,885]" pageId="1" pageNumber="176">Subulussalam-Singkil</collectingMunicipality>
,
<location id="674823D7E224FFF3FE6CFCB2FDAAFCB4" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FE6CFCB2FDAAFCB4" box="[477,566,862,885]" country="Indonesia" municipality="Subulussalam-Singkil" name="Trumon" pageId="1" pageNumber="176" stateProvince="Aceh">Trumon</location>
,
<collectorName id="CF6210DAE224FFF3FD8EFCB1FD2FFCB4" box="[575,691,861,885]" pageId="1" pageNumber="176">Kabupaten</collectorName>
<collectingRegion id="A053BBEEE224FFF3FD09FCB1FD6CFCB4" box="[696,752,861,885]" country="Indonesia" name="Aceh" pageId="1" pageNumber="176">Aceh</collectingRegion>
<collectorName id="CF6210DAE224FFF3FFC0FC91FF5DFC54" box="[113,193,893,917]" pageId="1" pageNumber="176">Singkil</collectorName>
,
<collectingRegion id="A053BBEEE224FFF3FF7EFC91FE96FC54" box="[207,266,893,917]" country="Indonesia" name="Aceh" pageId="1" pageNumber="176">Aceh</collectingRegion>
</materialsCitation>
,
<materialsCitation id="D2FF7F51E224FFF3FEA6FC92FD32FC54" box="[279,686,893,918]" collectingDate="2009-04-18" collectionCode="T" country="Indonesia" location="Sim" pageId="1" pageNumber="176" specimenCount="1" typeStatus="paratype">
<collectionCode id="0486EDC9E224FFF3FEA6FC92FEB6FC54" box="[279,298,894,917]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="176">T</collectionCode>
.
<location id="674823D7E224FFF3FE89FC91FEFFFC54" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FE89FC91FEFFFC54" box="[312,355,893,917]" country="Indonesia" name="Sim" pageId="1" pageNumber="176">Sim</location>
and company,
<date id="162953CCE224FFF3FDBEFC91FD32FC54" box="[527,686,893,917]" pageId="1" pageNumber="176" value="2009-04-18">
<collectingDate id="066DAA24E224FFF3FDBEFC91FD32FC54" box="[527,686,893,917]" pageId="1" pageNumber="176" value="2009-04-18">18 April 2009</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FD0DFC91FDF3FC34" accessionNumber="ZRC 55525" collectionCode="ZRC" collectorName="Kecamatan Gosong Telaga &amp; Kabupaten &amp; Singkil" country="Indonesia" county="Singkil-Subulussalam" location="Desa Kampung Baru" municipality="Singkil town" pageId="1" pageNumber="176" specimenCode="7" specimenCount="1" stateProvince="Aceh" typeStatus="paratype">
<collectionCode id="0486EDC9E224FFF3FD0DFC91FD6CFC54" box="[700,752,893,917]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="176">ZRC</collectionCode>
55525,
<specimenCode id="3231DD77E224FFF3FF72FC71FF4DFC74" box="[195,209,925,949]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="176">7</specimenCode>
(13–25), stream along road
<collectingCounty id="8B490D80E224FFF3FE4EFC71FD70FC74" box="[511,748,925,949]" pageId="1" pageNumber="176">Singkil-Subulussalam</collectingCounty>
, outskirts of
<collectingMunicipality id="824CEF76E224FFF3FF43FC51FEE2FC14" box="[242,382,957,981]" pageId="1" pageNumber="176">Singkil town</collectingMunicipality>
,
<location id="674823D7E224FFF3FE38FC52FDF0FC14" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FE38FC52FDF0FC14" box="[393,620,958,982]" country="Indonesia" county="Singkil-Subulussalam" municipality="Singkil town" name="Desa Kampung Baru" pageId="1" pageNumber="176" stateProvince="Aceh">Desa Kampung Baru</location>
,
<collectorName id="CF6210DAE224FFF3FDC6FC52FE8AFC34" pageId="1" pageNumber="176">Kecamatan Gosong Telaga</collectorName>
,
<collectorName id="CF6210DAE224FFF3FE90FC31FE09FC34" box="[289,405,989,1013]" pageId="1" pageNumber="176">Kabupaten</collectorName>
<collectingRegion id="A053BBEEE224FFF3FE2AFC31FE4FFC34" box="[411,467,989,1013]" country="Indonesia" name="Aceh" pageId="1" pageNumber="176">Aceh</collectingRegion>
<collectorName id="CF6210DAE224FFF3FE6BFC31FDB6FC34" box="[474,554,989,1013]" pageId="1" pageNumber="176">Singkil</collectorName>
,
<collectingRegion id="A053BBEEE224FFF3FD85FC31FDF3FC34" box="[564,623,989,1013]" country="Indonesia" name="Aceh" pageId="1" pageNumber="176">Aceh</collectingRegion>
</materialsCitation>
,
<materialsCitation id="D2FF7F51E224FFF3FDCBFC32FE1CFBD4" collectingDate="2009-04-19" collectionCode="T" country="Indonesia" location="Sim" pageId="1" pageNumber="176" specimenCount="1" typeStatus="paratype">
<collectionCode id="0486EDC9E224FFF3FDCBFC32FD11FC34" box="[634,653,990,1013]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="176">T</collectionCode>
.
<location id="674823D7E224FFF3FD26FC31FD5EFC34" LSID="urn:lsid:plazi:treatment:EA3EC41AE225FFF6FC02F964FACDF9D8:674823D7E224FFF3FD26FC31FD5EFC34" box="[663,706,989,1013]" country="Indonesia" name="Sim" pageId="1" pageNumber="176">Sim</location>
and company,
<date id="162953CCE224FFF3FF53FC11FE1CFBD4" box="[226,384,1021,1045]" pageId="1" pageNumber="176" value="2009-04-19">
<collectingDate id="066DAA24E224FFF3FF53FC11FE1CFBD4" box="[226,384,1021,1045]" pageId="1" pageNumber="176" value="2009-04-19">19 April 2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="6228750CE224FFF3FFC2FBD1FD6CFB14" blockId="1.[113,752,1085,1237]" pageId="1" pageNumber="176">
<emphasis id="50E3A91EE224FFF3FFC2FBD1FED2FB94" bold="true" box="[115,334,1085,1109]" pageId="1" pageNumber="176">Material examined</emphasis>
<emphasis id="50E3A91EE224FFF3FEE4FBD1FDB4FB94" box="[341,552,1085,1109]" italics="true" pageId="1" pageNumber="176">(but no data taken)</emphasis>
.
<materialsCitation id="D2FF7F51E224FFF3FD86FBD2FEC3FB74" collectingDate="2013-12-07" collectorName="Cibeber &amp; Kabupaten Pandeglang &amp; N. Hubert &amp; F. Busson" location="Banten Province, Java" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Banten Province, Java">
MZB-BIF 1441– 1442, 2,
<collectorName id="CF6210DAE224FFF3FF61FBB1FEBBFBB4" box="[208,295,1117,1141]" pageId="1" pageNumber="176">Cibeber</collectorName>
,
<collectorName id="CF6210DAE224FFF3FE83FBB1FDB0FBB4" box="[306,556,1117,1141]" pageId="1" pageNumber="176">Kabupaten Pandeglang</collectorName>
,
<collectingRegion id="A053BBEEE224FFF3FD86FBB2FF3FFB54" isEnumeration="true" pageId="1" pageNumber="176">Banten Province, Java</collectingRegion>
,
<collectorName id="CF6210DAE224FFF3FF03FB92FEBEFB54" box="[178,290,1149,1173]" pageId="1" pageNumber="176">N. Hubert</collectorName>
,
<collectorName id="CF6210DAE224FFF3FE81FB92FE3DFB54" box="[304,417,1150,1173]" pageId="1" pageNumber="176">F. Busson</collectorName>
,
<date id="162953CCE224FFF3FE1EFB92FDEFFB54" box="[431,627,1149,1173]" pageId="1" pageNumber="176" value="2013-12-07">
<collectingDate id="066DAA24E224FFF3FE1EFB92FDEFFB54" box="[431,627,1149,1173]" pageId="1" pageNumber="176" value="2013-12-07">7 December 2013</collectingDate>
</date>
(both with most of body missing)
</materialsCitation>
;
<materialsCitation id="D2FF7F51E224FFF3FED9FB72FD70FB14" collectingDate="2015-11-22" collectorName="Sungai Tumbuan" location="Bengkulu, Sumatera Selatan, Sumatra" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Bengkulu, Sumatera Selatan, Sumatra">
MZB-BIF 4859, 1,
<collectorName id="CF6210DAE224FFF3FD85FB71FD70FB74" box="[564,748,1181,1205]" pageId="1" pageNumber="176">Sungai Tumbuan</collectorName>
,
<collectingRegion id="A053BBEEE224FFF3FFC0FB51FD91FB14" box="[113,525,1213,1237]" isEnumeration="true" pageId="1" pageNumber="176">Bengkulu, Sumatera Selatan, Sumatra</collectingRegion>
,
<date id="162953CCE224FFF3FDA9FB51FD70FB14" box="[536,748,1213,1237]" pageId="1" pageNumber="176" value="2015-11-22">
<collectingDate id="066DAA24E224FFF3FDA9FB51FD70FB14" box="[536,748,1213,1237]" pageId="1" pageNumber="176" value="2015-11-22">22 November 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2A8D2687E224FFF3FFC0FB11FE8BF974" pageId="1" pageNumber="176" type="diagnosis">
<paragraph id="6228750CE224FFF3FFC0FB11FE8BF974" blockId="1.[112,752,1277,1717]" pageId="1" pageNumber="176">
<emphasis id="50E3A91EE224FFF3FFC0FB11FF77FAD4" bold="true" box="[113,235,1277,1301]" pageId="1" pageNumber="176">Diagnosis.</emphasis>
A moderately slender
<taxonomicName id="A5970E8FE224FFF3FE46FB11FD2FFAD4" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[503,691,1277,1301]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="176" phylum="Chordata" rank="genus">
<emphasis id="50E3A91EE224FFF3FE46FB11FD2FFAD4" box="[503,691,1277,1301]" italics="true" pageId="1" pageNumber="176">Pseudogobiopsis</emphasis>
</taxonomicName>
with second dorsal-fin rays always I,6; anal-fin rays always I,6; pectoral-fin rays 17–18; longitudinal scales 21–24; TRB 6–8; predorsal scales 5–6, large, reaching up to close behind eyes; jaws enlarged in male; three preopercular pores present, posterior portion of oculoscapular canal present, but no canal or pores over opercle; most scales on body ctenoid; first spine of dorsal fin longest in both sexes, greatly elongate in males; five elongate dark blotches along midside of body, with five indistinct dusky short saddles crossing dorsum, black spot behind anus and four internal black blotches along midventral line, commencing at anal fin origin, black spot on upper part of pectoral fin base and chin with blackish mental frenum.
</paragraph>
</subSubSection>
<subSubSection id="2A8D2687E224FFF6FC9CFF51FDACFAB8" lastPageId="4" lastPageNumber="179" pageId="1" pageNumber="176" type="description">
<paragraph id="6228750CE224FFF3FC9CFF51FBB4FED4" blockId="1.[813,1452,189,277]" pageId="1" pageNumber="176">
<emphasis id="50E3A91EE224FFF3FC9CFF51FC24FF14" bold="true" box="[813,952,189,213]" pageId="1" pageNumber="176">Description.</emphasis>
Based on
<specimenCount id="7491BE85E224FFF3FB9DFF51FB5CFF14" box="[1068,1216,189,213]" pageId="1" pageNumber="176" type="generic">18 specimens</specimenCount>
,
<quantity id="A56FD8E9E224FFF3FB7DFF51FAC8FF14" box="[1228,1364,189,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.575" metricValueMax="3.35" metricValueMin="1.8" pageId="1" pageNumber="176" unit="mm" value="25.75" valueMax="33.5" valueMin="18.0">18–33.5 mm</quantity>
SL. An asterisk indicates counts of
<typeStatus id="BD2CCBAEE224FFF3FBEFFF31FB27FF34" box="[1118,1211,221,245]" pageId="1" pageNumber="176" type="holotype">holotype</typeStatus>
(
<figureCitation id="FAAC6989E224FFF3FB7CFF31FA93FF34" box="[1229,1295,221,245]" captionStart="Fig" captionStartId="1.[113,148,2092,2114]" captionTargetBox="[113,1451,1787,2077]" captionTargetId="figure-772@1.[113,1451,1787,2077]" captionTargetPageId="1" captionText="Fig. 1. Pseudogobiopsis lumbantobing, photo of holotype male, MZB 23017. Photograph by Sandra Raredon." figureDoi="http://doi.org/10.5281/zenodo.4504450" httpUri="https://zenodo.org/record/4504450/files/figure.png" pageId="1" pageNumber="176">Fig. 1</figureCitation>
). See
<tableCitation id="2F1540B7E224FFF3FAE9FF31FA37FF34" box="[1368,1451,221,245]" captionStart="Table 1" captionStartId="2.[135,190,189,211]" captionText="Table 1. Morphometrics of Pseudogobiopsis lumbantobing n. sp." pageId="1" pageNumber="176">Table 1</tableCitation>
for morphometric data.
</paragraph>
<paragraph id="6228750CE224FFF3FC9CFED1FB2EFE34" blockId="1.[813,1452,317,501]" pageId="1" pageNumber="176">First dorsal-fin VI*; second dorsal-fin I,6*; anal-fin I,6*, pectoral-fin rays 16–18* (modally 18), segmented caudal-fin rays 16–17*; caudal-fin ray pattern 9/8; branched caudal-fin rays 11–14* (modally 7/7); longitudinal scale count 21–24* (modally 23*); TRB 6–8 (modally 7*); predorsal scale count 5*–6; circumpeduncular scales 12*.</paragraph>
<paragraph id="6228750CE224FFF3FC9CFDF1FB88FB74" blockId="1.[811,1453,541,1205]" pageId="1" pageNumber="176">Body compressed posteriorly, rounded anteriorly. Body depth at anal-fin origin 17.0–20.8% (mean 19.1%) of SL. Head length 25.6–29.6% (mean 27.6%) of SL; wider than deep, but not flattened or depressed even in large males. Head depth at posterior preopercular margin 57.9–69.1% (mean 62.1%) of HL; head width at posterior preopercular margin 62.9–74.5% (mean 69.1%) of HL; cheeks may be inflated in adult males. Mouth terminal, slightly oblique; jaws enlarged in males, reaching back to below mid-eye; females with jaws reaching to below front half of eye. Lips smooth, no fleshy fimbriae on inner edge of upper lip; lower lip free at sides, fused across anterior edge, with slightly raised fleshy symphysis. Upper jaw 24.5–52.5% (mean 34.9% in females, 45.3% in males) of HL. Eyes dorsolateral, forming part of dorsal profile, 26.3–36.4% of HL. Snout short, rounded and fleshy, curving anteriorly to slightly overhang upper lip in large specimens, 20.0–30.9% (mean 26.1%) of HL. Interorbital very narrow, 8.4–16.0% (mean 11.3%) of HL. Caudal peduncle long, compressed, length 30.0–36.7% (mean 32.3%) of SL. Caudal peduncle depth 11.9–14.4% (mean 13.4%) of SL.</paragraph>
<paragraph id="6228750CE224FFF3FC9CFB31FABCF914" blockId="1.[812,1452,1245,1749]" pageId="1" pageNumber="176">First dorsal fin triangular, first spine always longest in males, spine elongate and filamentous, reaching well past insertion of second dorsal fin when depressed; females without elongate first dorsal fin spines, usually first spine longest; fin barely reaching second dorsal fin origin when depressed. First dorsal-fin spine 33.6–49.5% (mean 39.6%) in males, 16.7–19.6% (mean 18.2%) in females, of SL. Second dorsal-fin spine longest in one female, length 17.3% of SL. Second dorsal and anal fins short-based, posteriormost fin rays longest, pointed, rays falling well short of caudal fin base when depressed. Pectoral fin slender, pointed, central rays longest, 24.8–29.3% (mean 27.4%) of SL; fin rays branched but for two or three uppermost. Pelvic fins slender to oval, reaching to or just past anus, 23.0–28.3% (mean 25.7%) of SL. Caudal fin relatively short, rounded to slightly pointed posteriorly, 26.3–31.9% (mean 28.8%) of SL.</paragraph>
<caption id="36E82584E224FFF3FFC0F7C0FB34F783" ID-DOI="http://doi.org/10.5281/zenodo.4504450" ID-Zenodo-Dep="4504450" box="[113,1192,2092,2114]" httpUri="https://zenodo.org/record/4504450/files/figure.png" pageId="1" pageNumber="176" startId="1.[113,148,2092,2114]" targetBox="[113,1451,1787,2077]" targetPageId="1">
<paragraph id="6228750CE224FFF3FFC0F7C0FB34F783" blockId="1.[113,1192,2092,2114]" box="[113,1192,2092,2114]" pageId="1" pageNumber="176">
Fig. 1.
<taxonomicName id="A5970E8FE224FFF3FF0AF7C0FE75F780" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[187,489,2092,2113]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="176" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE224FFF3FF0AF7C0FE75F780" box="[187,489,2092,2113]" italics="true" pageId="1" pageNumber="176">Pseudogobiopsis lumbantobing</emphasis>
</taxonomicName>
, photo of holotype male, MZB 23017. Photograph by Sandra Raredon.
</paragraph>
</caption>
<caption id="36E82584E227FFF0FF36FF51FC9AFF12" box="[135,774,189,211]" pageId="2" pageNumber="177">
<paragraph id="6228750CE227FFF0FF36FF51FC9AFF12" blockId="2.[135,774,189,211]" box="[135,774,189,211]" pageId="2" pageNumber="177">
Table 1. Morphometrics of
<taxonomicName id="A5970E8FE227FFF0FE28FF51FD54FF13" authority="Larson &amp; Hadiaty &amp; Hubert, 2017" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[409,712,189,210]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="177" phylum="Chordata" rank="species" species="lumbantobing" status="sp. nov.">
<emphasis id="50E3A91EE227FFF0FE28FF51FD54FF13" box="[409,712,189,210]" italics="true" pageId="2" pageNumber="177">Pseudogobiopsis lumbantobing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4BD01465E227FFF0FD61FF52FC9AFF12" box="[720,774,190,211]" pageId="2" pageNumber="177" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="6228750CE227FFF0FD9FFEE9FA05FC9F" pageId="2" pageNumber="177">
<table id="109787ACE227000DFF27FEE8FA00FC9E" box="[150,1436,260,863]" gridcols="6" gridrows="18" pageId="2" pageNumber="177">
<tr id="DCA7774EE227000DFF27FEE8FA00FEDB" box="[150,1436,260,282]" gridrow="0" pageId="2" pageNumber="177" rowspan-0="1">
<th id="9F761E32E227000DFD9FFEE8FD17FEDB" box="[558,651,260,282]" gridcol="1" gridrow="0" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FD9FFEE9FD17FEDB" bold="true" box="[558,651,261,282]" pageId="2" pageNumber="177">Holotype</emphasis>
</th>
<th id="9F761E32E227000DFD08FEE8FC0DFEDB" box="[697,913,260,282]" gridcol="2" gridrow="0" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FD08FEE9FC0DFEDB" bold="true" box="[697,913,260,282]" pageId="2" pageNumber="177">Males Range (n= 10)</emphasis>
</th>
<th id="9F761E32E227000DFC61FEE8FB97FEDB" box="[976,1035,260,282]" gridcol="3" gridrow="0" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FC61FEE9FB97FEDB" bold="true" box="[976,1035,261,282]" pageId="2" pageNumber="177">Mean</emphasis>
</th>
<th id="9F761E32E227000DFBF5FEE8FABBFEDB" box="[1092,1319,260,282]" gridcol="4" gridrow="0" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FBF5FEE9FABBFEDB" bold="true" box="[1092,1319,260,282]" pageId="2" pageNumber="177">Females Range (n= 8)</emphasis>
</th>
<th id="9F761E32E227000DFAD0FEE8FA00FEDB" box="[1377,1436,260,282]" gridcol="5" gridrow="0" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FAD0FEE9FA00FEDB" bold="true" box="[1377,1436,261,282]" pageId="2" pageNumber="177">Mean</emphasis>
</th>
</tr>
<tr id="DCA7774EE227000DFF27FED7FA00FE90" box="[150,1436,315,337]" gridrow="1" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FED7FA00FE90" box="[150,1436,315,337]" colspan="6" colspanRight="5" gridcol="0" gridrow="1" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FF27FED7FE53FE91" bold="true" box="[150,463,315,337]" pageId="2" pageNumber="177">Percentage of standard length</emphasis>
</th>
</tr>
<tr id="DCA7774EE227000DFF27FEB7FA00FEB0" box="[150,1436,347,369]" gridrow="2" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FEB7FE07FEB0" box="[150,411,347,369]" gridcol="0" gridrow="2" pageId="2" pageNumber="177">Head length</th>
<td id="9F761E32E227000DFD9FFEB7FD17FEB0" box="[558,651,347,369]" gridcol="1" gridrow="2" pageId="2" pageNumber="177">27.5</td>
<td id="9F761E32E227000DFD08FEB7FC0DFEB0" box="[697,913,347,369]" gridcol="2" gridrow="2" pageId="2" pageNumber="177">25.6–29.6</td>
<td id="9F761E32E227000DFC61FEB7FB97FEB0" box="[976,1035,347,369]" gridcol="3" gridrow="2" pageId="2" pageNumber="177">28.1</td>
<td id="9F761E32E227000DFBF5FEB7FABBFEB0" box="[1092,1319,347,369]" gridcol="4" gridrow="2" pageId="2" pageNumber="177">25.9–29.5</td>
<td id="9F761E32E227000DFAD0FEB7FA00FEB0" box="[1377,1436,347,369]" gridcol="5" gridrow="2" pageId="2" pageNumber="177">27.1</td>
</tr>
<tr id="DCA7774EE227000DFF27FE97FA00FE50" box="[150,1436,379,401]" gridrow="3" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FE97FE07FE50" box="[150,411,379,401]" gridcol="0" gridrow="3" pageId="2" pageNumber="177">Body depth</th>
<td id="9F761E32E227000DFD9FFE97FD17FE50" box="[558,651,379,401]" gridcol="1" gridrow="3" pageId="2" pageNumber="177">18.6</td>
<td id="9F761E32E227000DFD08FE97FC0DFE50" box="[697,913,379,401]" gridcol="2" gridrow="3" pageId="2" pageNumber="177">17.0–20.0</td>
<td id="9F761E32E227000DFC61FE97FB97FE50" box="[976,1035,379,401]" gridcol="3" gridrow="3" pageId="2" pageNumber="177">18.6</td>
<td id="9F761E32E227000DFBF5FE97FABBFE50" box="[1092,1319,379,401]" gridcol="4" gridrow="3" pageId="2" pageNumber="177">18.1–20.8</td>
<td id="9F761E32E227000DFAD0FE97FA00FE50" box="[1377,1436,379,401]" gridcol="5" gridrow="3" pageId="2" pageNumber="177">19.5</td>
</tr>
<tr id="DCA7774EE227000DFF27FE76FA00FE71" box="[150,1436,410,432]" gridrow="4" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FE76FE07FE71" box="[150,411,410,432]" gridcol="0" gridrow="4" pageId="2" pageNumber="177">Body width</th>
<td id="9F761E32E227000DFD9FFE76FD17FE71" box="[558,651,410,432]" gridcol="1" gridrow="4" pageId="2" pageNumber="177">–</td>
<td id="9F761E32E227000DFD08FE76FC0DFE71" box="[697,913,410,432]" gridcol="2" gridrow="4" pageId="2" pageNumber="177">14.1–16.1</td>
<td id="9F761E32E227000DFC61FE76FB97FE71" box="[976,1035,410,432]" gridcol="3" gridrow="4" pageId="2" pageNumber="177">15.2</td>
<td id="9F761E32E227000DFBF5FE76FABBFE71" box="[1092,1319,410,432]" gridcol="4" gridrow="4" pageId="2" pageNumber="177">15.0–17.2</td>
<td id="9F761E32E227000DFAD0FE76FA00FE71" box="[1377,1436,410,432]" gridcol="5" gridrow="4" pageId="2" pageNumber="177">15.8</td>
</tr>
<tr id="DCA7774EE227000DFF27FE56FA00FE11" box="[150,1436,442,464]" gridrow="5" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FE56FE07FE11" box="[150,411,442,464]" gridcol="0" gridrow="5" pageId="2" pageNumber="177">Length of caudal peduncle</th>
<td id="9F761E32E227000DFD9FFE56FD17FE11" box="[558,651,442,464]" gridcol="1" gridrow="5" pageId="2" pageNumber="177">30.2</td>
<td id="9F761E32E227000DFD08FE56FC0DFE11" box="[697,913,442,464]" gridcol="2" gridrow="5" pageId="2" pageNumber="177">30.0–34.7</td>
<td id="9F761E32E227000DFC61FE56FB97FE11" box="[976,1035,442,464]" gridcol="3" gridrow="5" pageId="2" pageNumber="177">31.6</td>
<td id="9F761E32E227000DFBF5FE56FABBFE11" box="[1092,1319,442,464]" gridcol="4" gridrow="5" pageId="2" pageNumber="177">31.3–36.7</td>
<td id="9F761E32E227000DFAD0FE56FA00FE11" box="[1377,1436,442,464]" gridcol="5" gridrow="5" pageId="2" pageNumber="177">33.0</td>
</tr>
<tr id="DCA7774EE227000DFF27FE36FA00FE31" box="[150,1436,474,496]" gridrow="6" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FE36FE07FE31" box="[150,411,474,496]" gridcol="0" gridrow="6" pageId="2" pageNumber="177">Depth of caudal peduncle</th>
<td id="9F761E32E227000DFD9FFE36FD17FE31" box="[558,651,474,496]" gridcol="1" gridrow="6" pageId="2" pageNumber="177">13.2</td>
<td id="9F761E32E227000DFD08FE36FC0DFE31" box="[697,913,474,496]" gridcol="2" gridrow="6" pageId="2" pageNumber="177">13.0–14.4</td>
<td id="9F761E32E227000DFC61FE36FB97FE31" box="[976,1035,474,496]" gridcol="3" gridrow="6" pageId="2" pageNumber="177">13.7</td>
<td id="9F761E32E227000DFBF5FE36FABBFE31" box="[1092,1319,474,496]" gridcol="4" gridrow="6" pageId="2" pageNumber="177">11.9–13.9</td>
<td id="9F761E32E227000DFAD0FE36FA00FE31" box="[1377,1436,474,496]" gridcol="5" gridrow="6" pageId="2" pageNumber="177">13.1</td>
</tr>
<tr id="DCA7774EE227000DFF27FE16FA00FDD1" box="[150,1436,506,528]" gridrow="7" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FE16FE07FDD1" box="[150,411,506,528]" gridcol="0" gridrow="7" pageId="2" pageNumber="177">Length of pectoral fin</th>
<td id="9F761E32E227000DFD9FFE16FD17FDD1" box="[558,651,506,528]" gridcol="1" gridrow="7" pageId="2" pageNumber="177">25.8</td>
<td id="9F761E32E227000DFD08FE16FC0DFDD1" box="[697,913,506,528]" gridcol="2" gridrow="7" pageId="2" pageNumber="177">25.8–29.3</td>
<td id="9F761E32E227000DFC61FE16FB97FDD1" box="[976,1035,506,528]" gridcol="3" gridrow="7" pageId="2" pageNumber="177">27.4</td>
<td id="9F761E32E227000DFBF5FE16FABBFDD1" box="[1092,1319,506,528]" gridcol="4" gridrow="7" pageId="2" pageNumber="177">24.8–29.0</td>
<td id="9F761E32E227000DFAD0FE16FA00FDD1" box="[1377,1436,506,528]" gridcol="5" gridrow="7" pageId="2" pageNumber="177">27.2</td>
</tr>
<tr id="DCA7774EE227000DFF27FDF5FA00FDF1" box="[150,1436,537,560]" gridrow="8" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FDF5FE07FDF1" box="[150,411,537,560]" gridcol="0" gridrow="8" pageId="2" pageNumber="177">Length of pelvic fin</th>
<td id="9F761E32E227000DFD9FFDF5FD17FDF1" box="[558,651,537,560]" gridcol="1" gridrow="8" pageId="2" pageNumber="177">25.1</td>
<td id="9F761E32E227000DFD08FDF5FC0DFDF1" box="[697,913,537,560]" gridcol="2" gridrow="8" pageId="2" pageNumber="177">24.8–27.8</td>
<td id="9F761E32E227000DFC61FDF5FB97FDF1" box="[976,1035,537,560]" gridcol="3" gridrow="8" pageId="2" pageNumber="177">26.1</td>
<td id="9F761E32E227000DFBF5FDF5FABBFDF1" box="[1092,1319,537,560]" gridcol="4" gridrow="8" pageId="2" pageNumber="177">23.0–28.3</td>
<td id="9F761E32E227000DFAD0FDF5FA00FDF1" box="[1377,1436,537,560]" gridcol="5" gridrow="8" pageId="2" pageNumber="177">25.2</td>
</tr>
<tr id="DCA7774EE227000DFF27FDD5FA00FD90" box="[150,1436,569,593]" gridrow="9" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FDD5FE07FD90" box="[150,411,569,593]" gridcol="0" gridrow="9" pageId="2" pageNumber="177">Length of caudal fin</th>
<td id="9F761E32E227000DFD9FFDD5FD17FD90" box="[558,651,569,593]" gridcol="1" gridrow="9" pageId="2" pageNumber="177">28.8</td>
<td id="9F761E32E227000DFD08FDD5FC0DFD90" box="[697,913,569,593]" gridcol="2" gridrow="9" pageId="2" pageNumber="177">28.5–31.9</td>
<td id="9F761E32E227000DFC61FDD5FB97FD90" box="[976,1035,569,593]" gridcol="3" gridrow="9" pageId="2" pageNumber="177">29.7</td>
<td id="9F761E32E227000DFBF5FDD5FABBFD90" box="[1092,1319,569,593]" gridcol="4" gridrow="9" pageId="2" pageNumber="177">26.3–29.4</td>
<td id="9F761E32E227000DFAD0FDD5FA00FD90" box="[1377,1436,569,593]" gridcol="5" gridrow="9" pageId="2" pageNumber="177">27.6</td>
</tr>
<tr id="DCA7774EE227000DFF27FDB0FA00FDB3" box="[150,1436,604,626]" gridrow="10" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FDB0FE07FDB3" box="[150,411,604,626]" gridcol="0" gridrow="10" pageId="2" pageNumber="177">Longest D1 spine</th>
<td id="9F761E32E227000DFD9FFDB0FD17FDB3" box="[558,651,604,626]" gridcol="1" gridrow="10" pageId="2" pageNumber="177">33.6</td>
<td id="9F761E32E227000DFD08FDB0FC0DFDB3" box="[697,913,604,626]" gridcol="2" gridrow="10" pageId="2" pageNumber="177">33.6–49.5</td>
<td id="9F761E32E227000DFC61FDB0FB97FDB3" box="[976,1035,604,626]" gridcol="3" gridrow="10" pageId="2" pageNumber="177">39.6</td>
<td id="9F761E32E227000DFBF5FDB0FABBFDB3" box="[1092,1319,604,626]" gridcol="4" gridrow="10" pageId="2" pageNumber="177">16.7–19.6</td>
<td id="9F761E32E227000DFAD0FDB0FA00FDB3" box="[1377,1436,604,626]" gridcol="5" gridrow="10" pageId="2" pageNumber="177">18.2</td>
</tr>
<tr id="DCA7774EE227000DFF27FD66FA00FD61" box="[150,1436,650,672]" gridrow="11" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FD66FA00FD61" box="[150,1436,650,672]" colspan="6" colspanRight="5" gridcol="0" gridrow="11" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FF27FD66FE38FD5E" bold="true" box="[150,420,650,672]" pageId="2" pageNumber="177">Percentage of head length</emphasis>
</th>
</tr>
<tr id="DCA7774EE227000DFF27FD46FA00FD01" box="[150,1436,682,704]" gridrow="12" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FD46FE07FD01" box="[150,411,682,704]" gridcol="0" gridrow="12" pageId="2" pageNumber="177">Head depth</th>
<td id="9F761E32E227000DFD9FFD46FD17FD01" box="[558,651,682,704]" gridcol="1" gridrow="12" pageId="2" pageNumber="177">59.3</td>
<td id="9F761E32E227000DFD08FD46FC0DFD01" box="[697,913,682,704]" gridcol="2" gridrow="12" pageId="2" pageNumber="177">57.6–65.3</td>
<td id="9F761E32E227000DFC61FD46FB97FD01" box="[976,1035,682,704]" gridcol="3" gridrow="12" pageId="2" pageNumber="177">60.5</td>
<td id="9F761E32E227000DFBF5FD46FABBFD01" box="[1092,1319,682,704]" gridcol="4" gridrow="12" pageId="2" pageNumber="177">58.1–69.1</td>
<td id="9F761E32E227000DFAD0FD46FA00FD01" box="[1377,1436,682,704]" gridcol="5" gridrow="12" pageId="2" pageNumber="177">63.3</td>
</tr>
<tr id="DCA7774EE227000DFF27FD26FA00FD21" box="[150,1436,714,736]" gridrow="13" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FD26FE07FD21" box="[150,411,714,736]" gridcol="0" gridrow="13" pageId="2" pageNumber="177">Head width</th>
<td id="9F761E32E227000DFD9FFD26FD17FD21" box="[558,651,714,736]" gridcol="1" gridrow="13" pageId="2" pageNumber="177">67.9</td>
<td id="9F761E32E227000DFD08FD26FC0DFD21" box="[697,913,714,736]" gridcol="2" gridrow="13" pageId="2" pageNumber="177">62.9–72.4</td>
<td id="9F761E32E227000DFC61FD26FB97FD21" box="[976,1035,714,736]" gridcol="3" gridrow="13" pageId="2" pageNumber="177">67.7</td>
<td id="9F761E32E227000DFBF5FD26FABBFD21" box="[1092,1319,714,736]" gridcol="4" gridrow="13" pageId="2" pageNumber="177">64.3–74.5</td>
<td id="9F761E32E227000DFAD0FD26FA00FD21" box="[1377,1436,714,736]" gridcol="5" gridrow="13" pageId="2" pageNumber="177">70.1</td>
</tr>
<tr id="DCA7774EE227000DFF27FD06FA00FCC1" box="[150,1436,746,768]" gridrow="14" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FD06FE07FCC1" box="[150,411,746,768]" gridcol="0" gridrow="14" pageId="2" pageNumber="177">Snout length</th>
<td id="9F761E32E227000DFD9FFD06FD17FCC1" box="[558,651,746,768]" gridcol="1" gridrow="14" pageId="2" pageNumber="177">27.2</td>
<td id="9F761E32E227000DFD08FD06FC0DFCC1" box="[697,913,746,768]" gridcol="2" gridrow="14" pageId="2" pageNumber="177">24.6–30.9</td>
<td id="9F761E32E227000DFC61FD06FB97FCC1" box="[976,1035,746,768]" gridcol="3" gridrow="14" pageId="2" pageNumber="177">27.8</td>
<td id="9F761E32E227000DFBF5FD06FABBFCC1" box="[1092,1319,746,768]" gridcol="4" gridrow="14" pageId="2" pageNumber="177">20.0–25.7</td>
<td id="9F761E32E227000DFAD0FD06FA00FCC1" box="[1377,1436,746,768]" gridcol="5" gridrow="14" pageId="2" pageNumber="177">24.1</td>
</tr>
<tr id="DCA7774EE227000DFF27FCE6FA00FCE1" box="[150,1436,778,800]" gridrow="15" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FCE6FE07FCE1" box="[150,411,778,800]" gridcol="0" gridrow="15" pageId="2" pageNumber="177">Eye width</th>
<td id="9F761E32E227000DFD9FFCE6FD17FCE1" box="[558,651,778,800]" gridcol="1" gridrow="15" pageId="2" pageNumber="177">29.6</td>
<td id="9F761E32E227000DFD08FCE6FC0DFCE1" box="[697,913,778,800]" gridcol="2" gridrow="15" pageId="2" pageNumber="177">26.3–31.9</td>
<td id="9F761E32E227000DFC61FCE6FB97FCE1" box="[976,1035,778,800]" gridcol="3" gridrow="15" pageId="2" pageNumber="177">29.2</td>
<td id="9F761E32E227000DFBF5FCE6FABBFCE1" box="[1092,1319,778,800]" gridcol="4" gridrow="15" pageId="2" pageNumber="177">30.0–36.4</td>
<td id="9F761E32E227000DFAD0FCE6FA00FCE1" box="[1377,1436,778,800]" gridcol="5" gridrow="15" pageId="2" pageNumber="177">32.2</td>
</tr>
<tr id="DCA7774EE227000DFF27FCC5FA00FCFE" box="[150,1436,809,831]" gridrow="16" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FCC5FE07FCFE" box="[150,411,809,831]" gridcol="0" gridrow="16" pageId="2" pageNumber="177">Jaw length</th>
<td id="9F761E32E227000DFD9FFCC5FD17FCFE" box="[558,651,809,831]" gridcol="1" gridrow="16" pageId="2" pageNumber="177">44.4</td>
<td id="9F761E32E227000DFD08FCC5FC0DFCFE" box="[697,913,809,831]" gridcol="2" gridrow="16" pageId="2" pageNumber="177">38.6–52.4</td>
<td id="9F761E32E227000DFC61FCC5FB97FCFE" box="[976,1035,809,831]" gridcol="3" gridrow="16" pageId="2" pageNumber="177">45.3</td>
<td id="9F761E32E227000DFBF5FCC5FABBFCFE" box="[1092,1319,809,831]" gridcol="4" gridrow="16" pageId="2" pageNumber="177">24.5–40.4</td>
<td id="9F761E32E227000DFAD0FCC5FA00FCFE" box="[1377,1436,809,831]" gridcol="5" gridrow="16" pageId="2" pageNumber="177">34.9</td>
</tr>
<tr id="DCA7774EE227000DFF27FCA5FA00FC9E" box="[150,1436,841,863]" gridrow="17" pageId="2" pageNumber="177">
<th id="9F761E32E227000DFF27FCA5FE07FC9E" box="[150,411,841,863]" gridcol="0" gridrow="17" pageId="2" pageNumber="177">Interorbital width</th>
<td id="9F761E32E227000DFD9FFCA5FD17FC9E" box="[558,651,841,863]" gridcol="1" gridrow="17" pageId="2" pageNumber="177">16.0</td>
<td id="9F761E32E227000DFD08FCA5FC0DFC9E" box="[697,913,841,863]" gridcol="2" gridrow="17" pageId="2" pageNumber="177">8.4–16.0</td>
<td id="9F761E32E227000DFC61FCA5FB97FC9E" box="[976,1035,841,863]" gridcol="3" gridrow="17" pageId="2" pageNumber="177">11.1</td>
<td id="9F761E32E227000DFBF5FCA5FABBFC9E" box="[1092,1319,841,863]" gridcol="4" gridrow="17" pageId="2" pageNumber="177">8.6–15.1</td>
<td id="9F761E32E227000DFAD0FCA5FA00FC9E" box="[1377,1436,841,863]" gridcol="5" gridrow="17" pageId="2" pageNumber="177">12.09</td>
</tr>
</table>
</paragraph>
<paragraph id="6228750CE227FFF0FF39FC42FD2EFAE7" blockId="2.[136,777,942,1318]" pageId="2" pageNumber="177">No distinct mental frenum, but chin smooth, with rounded to almost triangular fleshy pad behind symphysis (most developed in males). Anterior nostril placed just above preorbital edge, in very short tube. Posterior nostril rounded, placed just above front centre margin of eye. Gill opening moderate, usually extending forward to just under opercle. Inner edge of pectoral girdle smooth with no ridge or flange. Tongue short, tip usually blunt (rarely slightly concave to rounded). Teeth in both jaws small, curved and sharp, in 2–4 rows across front and only one row at side; outermost row teeth at front of jaws largest, curved and pointed (teeth larger in males and usually in more rows at front).</paragraph>
<caption id="36E82584E227FFF0FCF5FACDFC72FA4C" ID-DOI="http://doi.org/10.5281/zenodo.4504452" ID-Zenodo-Dep="4504452" httpUri="https://zenodo.org/record/4504452/files/figure.png" pageId="2" pageNumber="177" startId="2.[836,873,1313,1335]" targetBox="[835,1475,943,1298]" targetPageId="2">
<paragraph id="6228750CE227FFF0FCF5FACDFC72FA4C" blockId="2.[834,1474,1313,1421]" pageId="2" pageNumber="177">
Fig. 2. Sensory pores and papilla pattern in
<emphasis id="50E3A91EE227FFF0FAA9FACDFC53FA92" italics="true" pageId="2" pageNumber="177">
<taxonomicName id="A5970E8FE227FFF0FAA9FACDFC50FA92" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="177" phylum="Chordata" rank="species" species="lumbantobing">Pseudogobiopsis lumbantobing</taxonomicName>
.
</emphasis>
Composite drawing based on holotype and two other male specimens (MZB-BIF 4229 and MZB-BIF 4856). Drawing by Helen Larson.
</paragraph>
</caption>
<paragraph id="6228750CE227FFF0FF39FAA2FDF7F9A7" blockId="2.[136,777,1358,1638]" pageId="2" pageNumber="177">Predorsal scales moderately large, evenly sized, reaching to close behind eyes; anteriormost scale broader than those posterior to it. Operculum with 2–3 rows of large cycloid scales. Cheek naked. Pectoral-fin base covered with few large cycloid scales. Prepelvic area covered with cycloid scales. Belly covered with cycloid or ctenoid scales, may have some ctenoid scales near anus. Ctenoid scales on side of body extending forward to pectoral-fin base. Anteriormost scales on body larger than those posteriorly.</paragraph>
<paragraph id="6228750CE227FFF0FF39F962FDFAF907" blockId="2.[136,775,1678,1734]" pageId="2" pageNumber="177">Genital papilla in male elongate, flattened, narrowing toward tip; in female, papilla rounded and bulbous.</paragraph>
<paragraph id="6228750CE227FFF0FF39F902FEC7F844" blockId="2.[136,775,1774,1926]" pageId="2" pageNumber="177">
Head pores present (
<figureCitation id="FAAC6989E227FFF0FEC5F902FE27F8C7" box="[372,443,1774,1798]" captionStart="Fig" captionStartId="2.[836,873,1313,1335]" captionTargetBox="[835,1475,943,1298]" captionTargetId="figure-8@2.[814,1509,917,1318]" captionTargetPageId="2" captionText="Fig. 2. Sensory pores and papilla pattern in Pseudogobiopsis lumbantobing. Composite drawing based on holotype and two other male specimens (MZB-BIF 4229 and MZB-BIF 4856). Drawing by Helen Larson." figureDoi="http://doi.org/10.5281/zenodo.4504452" httpUri="https://zenodo.org/record/4504452/files/figure.png" pageId="2" pageNumber="177">Fig. 2</figureCitation>
). Pair of nasal pores, pair of anterior interorbital pores, a median posterior interorbital pore, postorbital and infraorbital pore behind each eye, oculoscapular canal and pore over preoperculum and three preopercular pores.
</paragraph>
<paragraph id="6228750CE227FFF0FF39F842FB14F9DD" blockId="2.[136,776,1966,2086]" lastBlockId="2.[835,1474,1476,1564]" pageId="2" pageNumber="177">
Sensory papillae pattern longitudinal, as in
<figureCitation id="FAAC6989E227FFF0FDDEF842FD28F807" box="[623,692,1966,1990]" captionStart="Fig" captionStartId="2.[836,873,1313,1335]" captionTargetBox="[835,1475,943,1298]" captionTargetId="figure-8@2.[814,1509,917,1318]" captionTargetPageId="2" captionText="Fig. 2. Sensory pores and papilla pattern in Pseudogobiopsis lumbantobing. Composite drawing based on holotype and two other male specimens (MZB-BIF 4229 and MZB-BIF 4856). Drawing by Helen Larson." figureDoi="http://doi.org/10.5281/zenodo.4504452" httpUri="https://zenodo.org/record/4504452/files/figure.png" pageId="2" pageNumber="177">Fig. 2</figureCitation>
. Cheek rows
<emphasis id="50E3A91EE227FFF0FF73F822FF75F824" box="[194,233,1998,2021]" italics="true" pageId="2" pageNumber="177">a, c</emphasis>
and
<emphasis id="50E3A91EE227FFF0FEADF822FEA9F824" box="[284,309,1998,2021]" italics="true" pageId="2" pageNumber="177">cp</emphasis>
composed of large, widely spaced papillae, and rows
<emphasis id="50E3A91EE227FFF0FF49F802FE99F7C7" box="[248,261,2030,2054]" italics="true" pageId="2" pageNumber="177">b</emphasis>
and
<emphasis id="50E3A91EE227FFF0FEF0F802FED2F7C7" box="[321,334,2030,2054]" italics="true" pageId="2" pageNumber="177">d</emphasis>
of very small, closely spaced papillae. Two to three
<emphasis id="50E3A91EE227FFF0FE90F7E2FEB7F7E4" box="[289,299,2062,2085]" italics="true" pageId="2" pageNumber="177">s</emphasis>
rows on snout, of one papilla each (upper row by nostril may have two papillae); middle row often absent. Mandibular
<emphasis id="50E3A91EE227FFF0FBADFA08FBBFFA3D" box="[1052,1059,1508,1532]" italics="true" pageId="2" pageNumber="177">f</emphasis>
row of two papillae, one on each side of fleshy frenum/pad on chin.
</paragraph>
<paragraph id="6228750CE227FFF1FCF2F9A8FEF8FB0A" blockId="2.[834,1476,1604,2108]" lastBlockId="3.[113,754,788,1228]" lastPageId="3" lastPageNumber="178" pageId="2" pageNumber="177">
<emphasis id="50E3A91EE227FFF0FCF2F9A8FB07F99D" bold="true" box="[835,1179,1604,1628]" pageId="2" pageNumber="177">Colouration of live material.</emphasis>
Based on photographs of captive and fresh dead specimens (
<figureCitation id="FAAC6989E227FFF0FB63F988FADDF9BD" box="[1234,1345,1636,1660]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[113,150,653,675]" captionStartId-1="3.[813,850,653,675]" captionStartId-2="3.[813,850,1156,1178]" captionStartId-3="4.[135,170,627,649]" captionTargetBox-0="[113,1452,189,638]" captionTargetBox-2="[813,1452,759,1141]" captionTargetBox-3="[135,1474,188,612]" captionTargetId-2="figure-621@3.[813,1452,759,1141]" captionTargetPageId-0="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="Fig. 3. Captive specimen of Pseudogobiopsis lumbantobing, supposedly imported from Myanmar. Photograph by Christophe Maillet." captionText-1="Fig. 4. Captive specimen of Pseudogobiopsis lumbantobing. Photograph by Thomas Ackerman." captionText-2="Fig. 5. Captive specimen of Pseudogobiopsis lumbantobing, imported via Singapore. Photograph by Emma Turner." captionText-3="Fig. 6. Freshly dead male (upper) and female (lower) specimens of Pseudogobiopsis lumbantobing (MZB-BIF 4856 and MZB-BIF 5859). Photograph by Nicolas Hubert." figureDoi-0="http://doi.org/10.5281/zenodo.4504454" figureDoi-2="http://doi.org/10.5281/zenodo.4504456" figureDoi-3="http://doi.org/10.5281/zenodo.4504460" httpUri-0="https://zenodo.org/record/4504454/files/figure.png" httpUri-2="https://zenodo.org/record/4504456/files/figure.png" httpUri-3="https://zenodo.org/record/4504460/files/figure.png" pageId="2" pageNumber="177">Figs. 3–6</figureCitation>
). Head and body translucent pearly grey, ventrally whitish, body and nape scattered with small rounded orange spots; in some individuals, spots smaller posteriorly and more brownishorange on dorsum and nape; scales on dorsal third of body with very thin dark brown margins. Side of head with scattered small orange spots; spots on snout brownish-orange. Upper part of cheek, mid-opercle and most of pectoral-fin base with fine opalescent speckles. Underside of head white with distinct black patch on chin and tip of lower jaw. Some photographs show a diffuse blackish area along anterior part of pectoral-fin base. First dorsal fin transparent, with first spine whitish and a few brown to black spots along edge, nearly half of fin occupied by black to dark brown vertical blotches extending out to fin margin (blotches smaller and more brownish in females); one photograph of a male shows the fin mostly translucent pinkish-orange to reddish, grey along base, with greyish blotches on outer part of fin membrane. Second dorsal fin transparent with about 3–6 rows of dusky yellow to pale brown spots on spines, fin margin transparent to pinkish. Caudal fin translucent, with 5 or 6 irregular vertical bands or rows of irregular spots of pale pinkish brown, bands breaking up into spots ventrally and fading posteriorly; one adult male has most of caudal fin pinkish-orange with whitish centre, and rows of brownish spots. Anal fin whitish to pink; one adult male displaying pinkish-orange fin rays with whitish membrane. Pectoral fin transparent, whitish near base. Pelvic fins translucent whitish to transparent.
</paragraph>
<caption id="36E82584E226FFF1FFC0FD61FF22FD1C" ID-DOI="http://doi.org/10.5281/zenodo.4504454" ID-Zenodo-Dep="4504454" httpUri="https://zenodo.org/record/4504454/files/figure.png" pageId="3" pageNumber="178" startId="3.[113,150,653,675]" targetBox="[113,1452,189,638]" targetPageId="3">
<paragraph id="6228750CE226FFF1FFC0FD61FF22FD1C" blockId="3.[113,752,653,733]" pageId="3" pageNumber="178">
Fig. 3. Captive specimen of
<taxonomicName id="A5970E8FE226FFF1FE15FD61FD76FD63" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[420,746,653,674]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE226FFF1FE15FD61FD76FD63" box="[420,746,653,674]" italics="true" pageId="3" pageNumber="178">Pseudogobiopsis lumbantobing</emphasis>
</taxonomicName>
, supposedly imported from Myanmar. Photograph by Christophe Maillet.
</paragraph>
</caption>
<caption id="36E82584E226FFF1FC9CFD61FB1FFD01" pageId="3" pageNumber="178">
<paragraph id="6228750CE226FFF1FC9CFD61FB1FFD01" blockId="3.[813,1452,653,704]" pageId="3" pageNumber="178">
Fig. 4. Captive specimen of
<emphasis id="50E3A91EE226FFF1FBD1FD61FA30FD63" box="[1120,1452,653,674]" italics="true" pageId="3" pageNumber="178">
<taxonomicName id="A5970E8FE226FFF1FBD1FD61FA34FD63" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[1120,1448,653,674]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="lumbantobing">Pseudogobiopsis lumbantobing</taxonomicName>
.
</emphasis>
Photograph by Thomas Ackerman.
</paragraph>
</caption>
<caption id="36E82584E226FFF1FC9CFB68FAD8FB77" ID-DOI="http://doi.org/10.5281/zenodo.4504456" ID-Zenodo-Dep="4504456" httpUri="https://zenodo.org/record/4504456/files/figure.png" pageId="3" pageNumber="178" startId="3.[813,850,1156,1178]" targetBox="[813,1452,759,1141]" targetPageId="3">
<paragraph id="6228750CE226FFF1FC9CFB68FAD8FB77" blockId="3.[813,1451,1156,1207]" pageId="3" pageNumber="178">
Fig. 5. Captive specimen of
<taxonomicName id="A5970E8FE226FFF1FBD1FB68FA39FB58" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[1120,1445,1156,1177]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE226FFF1FBD1FB68FA39FB58" box="[1120,1445,1156,1177]" italics="true" pageId="3" pageNumber="178">Pseudogobiopsis lumbantobing</emphasis>
</taxonomicName>
, imported via Singapore. Photograph by Emma Turner.
</paragraph>
</caption>
<paragraph id="6228750CE226FFF1FFC0FB18FE65F92D" blockId="3.[112,754,1268,1772]" pageId="3" pageNumber="178">
<emphasis id="50E3A91EE226FFF1FFC0FB18FE6EFACD" bold="true" box="[113,498,1268,1292]" pageId="3" pageNumber="178">Colouration of preserved material.</emphasis>
In males, head and body whitish to pale speckled yellowish, white on ventral third, with five elongate, rectangular, diffuse dark grey blotches along midside of body, posteriormost blotch just before hypural crease and may be darker than others; body with scattered small grey-margined pale spots, becoming darker and arranged more regularly on lower part of body (
<figureCitation id="FAAC6989E226FFF1FD0DFA58FFE2FA2D" captionStart="Fig" captionStartId="1.[113,148,2092,2114]" captionTargetBox="[113,1451,1787,2077]" captionTargetId="figure-772@1.[113,1451,1787,2077]" captionTargetPageId="1" captionText="Fig. 1. Pseudogobiopsis lumbantobing, photo of holotype male, MZB 23017. Photograph by Sandra Raredon." figureDoi="http://doi.org/10.5281/zenodo.4504450" httpUri="https://zenodo.org/record/4504450/files/figure.png" pageId="3" pageNumber="178">Figs. 1</figureCitation>
,
<figureCitation id="FAAC6989E226FFF1FF3CFA38FF01FA2D" box="[141,157,1492,1516]" captionStart="Fig" captionStartId="4.[135,171,1110,1132]" captionTargetBox="[135,774,762,1095]" captionTargetId="figure-431@4.[135,774,762,1095]" captionTargetPageId="4" captionText="Fig. 7. Preserved specimens of Pseudogobiopsis lumbantobing, ZRC 55524 (both males, upper 31.4 mm SL, lower 28.0 mm SL; not to scale). Photograph by Tan Heok Hui." figureDoi="http://doi.org/10.5281/zenodo.4504462" httpUri="https://zenodo.org/record/4504462/files/figure.png" pageId="3" pageNumber="178">7</figureCitation>
). Five diffuse dusky grey saddles usually visible. Nape and dorsum often with scale margins greyish, giving netlike appearance. Pectoral fin base with dense black irregular blotch on upper portion, blotch may partly extend upward onto body and under opercle toward pectoral girdle. Ventral midline of caudal peduncle with four evenly spaced, partly internal, narrow black blotches present. Belly with partly internal black spot just behind the anus in both sexes. Male urogenital papilla pale to dark grey.
</paragraph>
<paragraph id="6228750CE226FFF1FFC0F8F8FB37F987" blockId="3.[113,753,1812,2124]" lastBlockId="3.[813,1452,1294,1606]" pageId="3" pageNumber="178">
Head dusky grey dorsally with short dark grey triangular to linear mark extending from front of eye to almost straight back to end on cheek well above corner of mouth; second dark grey broad bar extending from rear part of eye obliquely across cheek. Small blackish blotch around and including posterior nostril by eye. Opercle with large rounded grey blotch or horizontal bar across centre. Upper lip plain dusky grey, lower lip paler in most specimens.
<collectingRegion id="A053BBEEE226FFF1FD8FF818FDEEF7CD" box="[574,626,2036,2060]" country="Myanmar" name="Chin" pageId="3" pageNumber="178">Chin</collectingRegion>
with black blotch on mental frenum; black pigment extending up to centre of lower lip. Breast and belly whitish to faintly greyish. First dorsal fin in males grey to blackish with two to three rows of black spots on spines and fin membranes (spots on membrane joining to form irregular blackish rows), first dorsal-fin spine mostly blackish. Second dorsal fin dusky grey, with 5–7 rows of blackish spots forming rows, becoming darker posteriorly. Anal fin plain dark grey, blackish posteriorly. Caudal fin translucent to dusky with 7–12 rows of small dark grey spots or fine irregular dusky bars. Pectoral fin translucent to dusky grey, darker ventrally. Pelvic fins and frenum dusky grey.
</paragraph>
<paragraph id="6228750CE226FFF1FC9CF982FC5DF887" blockId="3.[813,1452,1646,1862]" pageId="3" pageNumber="178">Females similar to males but paler and markings less pronounced. Black blotch on pectoral fin base smaller, less distinct; first dorsal fin transparent with large black blotch on rear third of fin; mental frenum paler, never black; pelvic fins transparent; caudal fin with 3–5 rows of brownish grey spots coalescing to form irregular oblique bands; urogenital papilla white.</paragraph>
<paragraph id="6228750CE226FFF6FC9CF882FDACFAB8" blockId="3.[811,1453,1902,2086]" lastBlockId="4.[136,776,1248,1401]" lastPageId="4" lastPageNumber="179" pageId="3" pageNumber="178">
<emphasis id="50E3A91EE226FFF1FC9CF882FC57F847" bold="true" box="[813,971,1902,1926]" pageId="3" pageNumber="178">Comparisons.</emphasis>
This species is most similar to
<taxonomicName id="A5970E8FE226FFF1FA9AF883FA39F847" baseAuthorityName="Bleeker" baseAuthorityYear="1875" box="[1323,1445,1902,1926]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="oligactis">
<emphasis id="50E3A91EE226FFF1FA9AF883FA39F847" box="[1323,1445,1902,1926]" italics="true" pageId="3" pageNumber="178">P. oligactis</emphasis>
</taxonomicName>
, differing in that
<taxonomicName id="A5970E8FE226FFF1FC50F863FB08F867" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[993,1172,1934,1958]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE226FFF1FC50F863FB08F867" box="[993,1172,1934,1958]" italics="true" pageId="3" pageNumber="178">P. lumbantobing</emphasis>
</taxonomicName>
has a pale body scattered with small orange spots when live; has a grey to blackish blotch on the chin; a black blotch on the upper part of pectoral fin base; and the snout is rounded over the anterior part of the upper lip (snout inflated in males) versus
<taxonomicName id="A5970E8FE226FFF1FA9FF7E3FA30F7E7" baseAuthorityName="Bleeker" baseAuthorityYear="1875" box="[1326,1452,2062,2086]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="178" phylum="Chordata" rank="species" species="oligactis">
<emphasis id="50E3A91EE226FFF1FA9FF7E3FA30F7E7" box="[1326,1452,2062,2086]" italics="true" pageId="3" pageNumber="178">P. oligactis</emphasis>
</taxonomicName>
which has a greyish body with internal grey blotches when live; the lips are plain dusky grey without a black blotch on the chin; the pectoral-fin base is grey but without a distinct black blotch; and the snout is pointed and flat. See
<bibRefCitation id="060608FDE221FFF6FD0CFAADFF53FAB8" author="Larson HK" pageId="4" pageNumber="179" pagination="127 - 181" refId="ref4829" refString="Larson HK (2009) Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae: Gobionellinae), with descriptions of three new species. Raffles Bulletin of Zoology, 57 (1): 127 - 181." type="journal article" year="2009">Larson (2009)</bibRefCitation>
for comparison with congeners.
</paragraph>
</subSubSection>
<caption id="36E82584E221FFF6FF36FD9FFE62FD02" ID-DOI="http://doi.org/10.5281/zenodo.4504460" ID-Zenodo-Dep="4504460" httpUri="https://zenodo.org/record/4504460/files/figure.png" pageId="4" pageNumber="179" startId="4.[135,170,627,649]" targetBox="[135,1474,188,612]" targetPageId="4">
<paragraph id="6228750CE221FFF6FF36FD9FFE62FD02" blockId="4.[135,774,627,707]" pageId="4" pageNumber="179">
Fig. 6. Freshly dead male (upper) and female (lower) specimens of
<taxonomicName id="A5970E8FE221FFF6FF10FD7CFE53FD64" authorityName="Larson &amp; Hadiaty &amp; Hubert" authorityYear="2017" box="[161,463,656,677]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="179" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE221FFF6FF10FD7CFE53FD64" box="[161,463,656,677]" italics="true" pageId="4" pageNumber="179">Pseudogobiopsis lumbantobing</emphasis>
</taxonomicName>
(MZB-BIF 4856 and MZB-BIF 5859). Photograph by Nicolas Hubert.
</paragraph>
</caption>
<caption id="36E82584E221FFF6FCF2FD9FFA5FFD48" box="[835,1475,627,649]" pageId="4" pageNumber="179">
<paragraph id="6228750CE221FFF6FCF2FD9FFA5FFD48" blockId="4.[835,1475,627,649]" box="[835,1475,627,649]" pageId="4" pageNumber="179">Fig. 8. Sungei Kluet, Sumatra. Photograph by Daniel Lumbantobing.</paragraph>
</caption>
<caption id="36E82584E221FFF6FF36FBBAFDA4FB67" ID-DOI="http://doi.org/10.5281/zenodo.4504462" ID-Zenodo-Dep="4504462" httpUri="https://zenodo.org/record/4504462/files/figure.png" pageId="4" pageNumber="179" startId="4.[135,171,1110,1132]" targetBox="[135,774,762,1095]" targetPageId="4">
<paragraph id="6228750CE221FFF6FF36FBBAFDA4FB67" blockId="4.[135,774,1110,1190]" pageId="4" pageNumber="179">
Fig. 7. Preserved specimens of
<taxonomicName id="A5970E8FE221FFF6FE78FBBAFF2AFB49" authority=", ZRC" authorityName="ZRC" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="179" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE221FFF6FE78FBBAFC9CFBAA" box="[457,768,1110,1131]" italics="true" pageId="4" pageNumber="179">Pseudogobiopsis lumbantobing</emphasis>
, ZRC
</taxonomicName>
55524 (both males, upper 31.4 mm SL, lower 28.0 mm SL; not to scale). Photograph by Tan Heok Hui.
</paragraph>
</caption>
<caption id="36E82584E221FFF6FCF2FB9AFBE9FB68" ID-DOI="http://doi.org/10.5281/zenodo.4504464" ID-Zenodo-Dep="4504464" httpUri="https://zenodo.org/record/4504464/files/figure.png" pageId="4" pageNumber="179" startId="4.[835,870,1142,1164]" targetBox="[835,1474,704,1127]" targetPageId="4">
<paragraph id="6228750CE221FFF6FCF2FB9AFBE9FB68" blockId="4.[835,1475,1142,1193]" pageId="4" pageNumber="179">Fig. 9. River at Cibeber, Kabupaten Pandeglang, Banten Province. Photograph by Nicolas Hubert.</paragraph>
</caption>
<subSubSection id="2A8D2687E221FFF6FF39FA4CFD7BF9D9" pageId="4" pageNumber="179" type="distribution">
<paragraph id="6228750CE221FFF6FF39FA4CFD7BF9D9" blockId="4.[136,776,1440,1561]" pageId="4" pageNumber="179">
<emphasis id="50E3A91EE221FFF6FF39FA4CFEBAFA79" bold="true" box="[136,294,1440,1464]" pageId="4" pageNumber="179">Distribution.</emphasis>
Known only from freshwater drainages of western
<collectingRegion id="A053BBEEE221FFF6FEBAFA2DFEFBFA18" box="[267,359,1473,1497]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="179">Sumatra</collectingRegion>
and western
<collectingRegion id="A053BBEEE221FFF6FDB6FA2DFDA7FA19" box="[519,571,1473,1496]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="179">Java</collectingRegion>
,
<collectingCountry id="1A80359CE221FFF6FDFBFA2CFD26FA19" box="[586,698,1472,1496]" name="Indonesia" pageId="4" pageNumber="179">Indonesia</collectingCountry>
. Some specimens in the aquarium trade are said to be imported from
<collectingCountry id="1A80359CE221FFF6FF74F9EDFEB1F9D8" box="[197,301,1537,1561]" name="Myanmar" pageId="4" pageNumber="179">Myanmar</collectingCountry>
(
<figureCitation id="FAAC6989E221FFF6FE8CF9ECFEE3F9D8" box="[317,383,1536,1561]" captionStart="Fig" captionStartId="3.[113,150,653,675]" captionTargetBox="[113,1452,189,638]" captionTargetPageId="3" captionText="Fig. 3. Captive specimen of Pseudogobiopsis lumbantobing, supposedly imported from Myanmar. Photograph by Christophe Maillet." figureDoi="http://doi.org/10.5281/zenodo.4504454" httpUri="https://zenodo.org/record/4504454/files/figure.png" pageId="4" pageNumber="179">Fig. 3</figureCitation>
), but this is yet to be confirmed.
</paragraph>
</subSubSection>
<subSubSection id="2A8D2687E221FFF6FF39F9ADFE94F978" pageId="4" pageNumber="179" type="biology_ecology">
<paragraph id="6228750CE221FFF6FF39F9ADFE94F978" blockId="4.[135,774,1600,1721]" pageId="4" pageNumber="179">
<emphasis id="50E3A91EE221FFF6FF39F9ADFF75F998" bold="true" box="[136,233,1601,1625]" pageId="4" pageNumber="179">Ecology.</emphasis>
Freshwater, found in rivers and streams at altitudes of
<quantity id="A56FD8E9E221FFF6FF19F98CFE9EF9B9" box="[168,258,1632,1656]" metricMagnitude="1" metricUnit="m" metricValue="1.35" metricValueMax="2.2" metricValueMin="0.5" pageId="4" pageNumber="179" unit="m" value="13.5" valueMax="22.0" valueMin="5.0">5–22 m</quantity>
, with a substratum of sand, gravel, rock and boulders; algae and aquatic macrophytes may be present (
<figureCitation id="FAAC6989E221FFF6FF21F94CFF41F978" box="[144,221,1696,1721]" captionStart="Fig" captionStartId="4.[835,869,627,649]" captionText="Fig.8. Sungei Kluet, Sumatra.Photograph by Daniel Lumbantobing." pageId="4" pageNumber="179">Figs. 8</figureCitation>
,
<figureCitation id="FAAC6989E221FFF6FF5AF94DFF67F978" box="[235,251,1697,1721]" captionStart="Fig" captionStartId="4.[835,870,1142,1164]" captionTargetBox="[835,1474,704,1127]" captionTargetId="figure-471@4.[835,1474,704,1127]" captionTargetPageId="4" captionText="Fig. 9. River at Cibeber, Kabupaten Pandeglang, Banten Province. Photograph by Nicolas Hubert." figureDoi="http://doi.org/10.5281/zenodo.4504464" httpUri="https://zenodo.org/record/4504464/files/figure.png" pageId="4" pageNumber="179">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2A8D2687E221FFF6FF39F90DFD45F859" pageId="4" pageNumber="179" type="etymology">
<paragraph id="6228750CE221FFF6FF39F90DFD45F859" blockId="4.[135,775,1760,1944]" pageId="4" pageNumber="179">
<emphasis id="50E3A91EE221FFF6FF39F90DFE96F938" bold="true" box="[136,266,1761,1785]" pageId="4" pageNumber="179">Etymology.</emphasis>
This species is named for Daniel Lumbantobing of
<collectingRegion id="A053BBEEE221FFF6FF16F8ECFF6AF8D9" box="[167,246,1792,1816]" country="Indonesia" name="Jakarta Raya" pageId="4" pageNumber="179">Jakarta</collectingRegion>
, who collected the first specimens and showed them to HKL in 2012, which solved the mystery of the orange-spotted goby of which aquarists had been sending her photos. Daniel is an ichthyologist who specialises in freshwater fishes. Name used as a name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="2A8D2687E221FFF6FF39F82DFACDF9D8" pageId="4" pageNumber="179" type="discussion">
<paragraph id="6228750CE221FFF6FF39F82DFAB6FA59" blockId="4.[134,775,1984,2073]" lastBlockId="4.[835,1475,1248,1433]" pageId="4" pageNumber="179">
<emphasis id="50E3A91EE221FFF6FF39F82DFF69F818" bold="true" box="[136,245,1985,2009]" pageId="4" pageNumber="179">Remarks.</emphasis>
The identity of a Kuhl and van Hasselt watercolour painting, done during 1820–1823, of a goby from
<collectingRegion id="A053BBEEE221FFF6FD7FF80DFC9DF839" box="[718,769,2017,2040]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="179">Java</collectingRegion>
, illustrated in
<bibRefCitation id="060608FDE221FFF6FEA8F7ECFE22F7D8" author="Roberts TR" box="[281,446,2048,2073]" pageId="4" pageNumber="179" pagination="1 - 94" refId="ref4872" refString="Roberts TR (1993) The freshwater fishes of Java, as observed by Kuhl and van Hasselt in 1820 - 23. Zoologische Verhandelingen, 285: 1 - 94." type="journal article" year="1993">Roberts (1993)</bibRefCitation>
, has been a mystery for some time (
<bibRefCitation id="060608FDE221FFF6FC31FB0DFBDBFB39" author="Larson HK" box="[896,1095,1248,1273]" pageId="4" pageNumber="179" pagination="127 - 181" refId="ref4829" refString="Larson HK (2009) Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae: Gobionellinae), with descriptions of three new species. Raffles Bulletin of Zoology, 57 (1): 127 - 181." type="journal article" year="2009">Larson, 2009: 162</bibRefCitation>
). Now that specimens are available, we conclude from the body and fin shape and colour pattern, including the dark grey chin (
<bibRefCitation id="060608FDE221FFF6FB3BFACCFABBFAF8" author="Roberts TR" box="[1162,1319,1312,1337]" pageId="4" pageNumber="179" pagination="1 - 94" refId="ref4872" refString="Roberts TR (1993) The freshwater fishes of Java, as observed by Kuhl and van Hasselt in 1820 - 23. Zoologische Verhandelingen, 285: 1 - 94." type="journal article" year="1993">Roberts, 1993</bibRefCitation>
: Fig. 51), that the figure is of
<taxonomicName id="A5970E8FE221FFF6FC59FAADFB07FA99" box="[1000,1179,1344,1368]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="179" phylum="Chordata" rank="species" species="lumbantobing">
<emphasis id="50E3A91EE221FFF6FC59FAADFB07FA99" box="[1000,1179,1344,1368]" italics="true" pageId="4" pageNumber="179">P. lumbantobing</emphasis>
</taxonomicName>
. It was hoped to reproduce the painting here, but access to the Naturalis Library is not possible until 2018 (undergoing renovation).
</paragraph>
<paragraph id="6228750CE221FFF6FCF2FA2DFACDF9D8" blockId="4.[835,1474,1472,1561]" pageId="4" pageNumber="179">
It should also be noted that in
<bibRefCitation id="060608FDE221FFF6FB27FA2DFAB7FA18" author="Larson HK" box="[1174,1323,1473,1497]" pageId="4" pageNumber="179" pagination="127 - 181" refId="ref4829" refString="Larson HK (2009) Review of the gobiid fish genera Eugnathogobius and Pseudogobiopsis (Gobioidei: Gobiidae: Gobionellinae), with descriptions of three new species. Raffles Bulletin of Zoology, 57 (1): 127 - 181." type="journal article" year="2009">Larson (2009</bibRefCitation>
: Plate B) that the photograph of
<taxonomicName id="A5970E8FE221FFF6FBB9FA0DFB1DFA39" baseAuthorityName="Bleeker" baseAuthorityYear="1875" box="[1032,1153,1504,1528]" class="Actinopterygii" family="Gobiidae" genus="Pseudogobiopsis" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="179" phylum="Chordata" rank="species" species="oligactis">
<emphasis id="50E3A91EE221FFF6FBB9FA0DFB1DFA39" box="[1032,1153,1504,1528]" italics="true" pageId="4" pageNumber="179">P. oligactis</emphasis>
</taxonomicName>
in Plate 2B was inadvertently attributed to Gerhard Ott; it is by Rainer Hoyer.
</paragraph>
</subSubSection>
</treatment>
</document>
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