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<document id="6536FCC4A28C5C4AAAA317081CDF605A" ID-DOI="10.21248/contrib.entomol.57.2.355-366" ID-ISSN="0005-805X" ID-Zenodo-Dep="5364133" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630242637808" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2007" docId="8F7487ACF1225D067B0929DBFC8193A9" docLanguage="en" docName="ContribEntomol.2007.57.2.355-366.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 57 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1709" docStyle="DocumentStyle:DA3631FDA31ED51D1CF99ECA6F54F23F.3:ContribEntomol.2005-2012.journal_article" docStyleId="DA3631FDA31ED51D1CF99ECA6F54F23F" docStyleName="ContribEntomol.2005-2012.journal_article" docStyleVersion="3" docTitle="Proteinus hamatus Assing 2007, sp. n." docType="treatment" docVersion="3" lastPageNumber="365" masterDocId="734DFFD4F1285D0C7A00296BFF96970E" masterDocTitle="Proteinus crenulatus - a complex of five species (Coleoptera: Staphylinidae: Proteininae)" masterLastPageNumber="366" masterPageNumber="355" pageNumber="365" updateTime="1699196385817" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="575065AB046B93816F4C2776A28B0BBF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FB5F30DB23DCD7D94996A4B84B916939">
<mods:title id="8D39BE6D410D63D88702D693C85DBA55">Proteinus crenulatus - a complex of five species (Coleoptera: Staphylinidae: Proteininae)</mods:title>
</mods:titleInfo>
<mods:name id="288CA188C0B84832ECAC7FF42F64CC52" type="personal">
<mods:role id="96CE41B3406A613FFF4123548D01A0E9">
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<mods:namePart id="5187466FC75482C5388D48622699E327">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="13AD999CF5623F9CF4EA046B71A61B08">text</mods:typeOfResource>
<mods:relatedItem id="8B43F5F12F8259352B894E0DC1699F28" type="host">
<mods:titleInfo id="CEAA67D5AFE888D66F85D8BC67A02B19">
<mods:title id="6A4C8F444C0D2E7339857FB1C82FEC83">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="B60E914071B03E25F990A9CA43BD94A3">
<mods:date id="22747407B328F3FD87653ED830F0F8BA">2007</mods:date>
<mods:detail id="C3794D8A5A7B2FE35CF95B090A753D40" type="pubDate">
<mods:number id="644067D3220D118F1FE22D29D3376212">2007-12-21</mods:number>
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<mods:detail id="DC32834B80EDAADAFFEE74D9B57DE7EB" type="volume">
<mods:number id="332CAB1BF01664EB5A11E6120CC631CE">57</mods:number>
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<mods:number id="0FFDDA9EC61303C5414CC13F39BF16C6">2</mods:number>
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<mods:url id="F6758CE73CF2E0492BD1C5256A7168A5">https://www.contributions-to-entomology.org/article/view/1709</mods:url>
</mods:location>
<mods:classification id="9849502209348DBE8560C172E6058B58">journal article</mods:classification>
<mods:identifier id="ED4A6DD5F44A6ADE93C153EF3C117D34" type="DOI">10.21248/contrib.entomol.57.2.355-366</mods:identifier>
<mods:identifier id="708081C653EC219A29CD7BA1AC3B7702" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="E56CC9BD4FB989E6D6C161E67832B58E" type="Zenodo-Dep">5364133</mods:identifier>
</mods:mods>
<treatment id="8F7487ACF1225D067B0929DBFC8193A9" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9" httpUri="http://treatment.plazi.org/id/8F7487ACF1225D067B0929DBFC8193A9" lastPageNumber="365" pageId="10" pageNumber="365">
<subSubSection id="4FC76531F1225D067B0929DBFD6897C9" box="[265,766,175,200]" pageId="10" pageNumber="365" type="nomenclature">
<paragraph id="076236BAF1225D067B0929DBFD6897C9" blockId="10.[265,766,175,200]" box="[265,766,175,200]" pageId="10" pageNumber="365">
<emphasis id="35A9EAA8F1225D067B0929DBFDB697C6" bold="true" box="[265,544,176,200]" pageId="10" pageNumber="365">
<taxonomicName id="C0DD4D39F1225D067B0929DBFE4B97CC" ID-CoL="4N36Y" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[265,477,176,194]" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="35A9EAA8F1225D067B0929DBFE4B97CC" bold="true" box="[265,477,176,194]" italics="true" pageId="10" pageNumber="365">Proteinus hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2E9A57D3F1225D067BE429DDFDB697C6" box="[484,544,182,200]" pageId="10" pageNumber="365" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="9FE62A3FF1225D06783029C4FD3397C9" box="[560,677,175,199]" captionStart="Figs 12-21" captionStartId="8.[194,230,1432,1454]" captionTargetBox="[205,1166,210,1400]" captionTargetId="figure-18@8.[195,1170,204,1406]" captionTargetPageId="8" captionText="Figs 12-21: Proteinus abditusabditus sp. n. (12), P.angulatusangulatus sp. n. (13-17), P. tronquetitronqueti sp. n. (18-19), and P. hamatus sp. n. (20-21): apical part of aedeagus in lateral view (12, 17); aedeagus in lateral view (13-15, 18-21) of males from Greece (13; holotype), Sicily (14), Basilicata (15), Corsica (18-19; 18: holotype), northern Spain (20; holotype), and southwestern France (21); apical part of aedeagus in dorsal view (16). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364141" httpUri="https://zenodo.org/record/5364141/files/figure.png" pageId="10" pageNumber="365">Figs 20-21</figureCitation>
,
<figureCitation id="9FE62A3FF1225D0678B129DBFD6197C9" box="[689,759,175,199]" captionStart="Map 2" captionStartId="6.[190,234,1604,1625]" captionTargetBox="[189,1172,851,1598]" captionTargetId="figure-310@6.[189,1172,851,1598]" captionTargetPageId="6" captionText="Map 2: Distributions of Proteinus abditus sp. n. (open squares), P. angulatus sp. n. (filled squares), P. hamatustus sp. n. (open circles), and P. tronqueti sp. n. (filled circles) in Europe." figureDoi="http://doi.org/10.5281/zenodo.5364139" httpUri="https://zenodo.org/record/5364139/files/figure.png" pageId="10" pageNumber="365">Map 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4FC76531F1225D067ABD2987FC8F96F1" pageId="10" pageNumber="365" type="materials_examined">
<paragraph id="076236BAF1225D067ABD2987FEC5960A" blockId="10.[189,339,236,260]" box="[189,339,236,260]" pageId="10" pageNumber="365">
<emphasis id="35A9EAA8F1225D067ABD2987FEC5960A" bold="true" box="[189,339,236,260]" pageId="10" pageNumber="365">Type material:</emphasis>
</paragraph>
<paragraph id="076236BAF1225D067ABE2879FC8F96F1" blockId="10.[189,1180,274,511]" pageId="10" pageNumber="365">
<materialsCitation id="B7B53CE7F1225D067ABE2879FF689666" collectingDate="2003-10-15" collectorName="V. Assing" county="Pena Negra" determinerName="V. Assing" elevation="1975" latitude="42.05" location="Castilla-Leon" longLatPrecision="1152" longitude="-2.7666667" municipality="Sierra de Urbion" pageId="10" pageNumber="365" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="D8668818F1225D067ABE2879FE889624" box="[190,286,274,298]" pageId="10" pageNumber="365" type="holotype">Holotype</typeStatus>
<specimenCount id="11DBFD33F1225D067B29287EFEA0962B" box="[297,310,277,293]" pageId="10" pageNumber="365" type="male">♂</specimenCount>
: &quot;E - No. 13,
<location id="02026061F1225D067BD02879FDCB9624" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9:02026061F1225D067BD02879FDCB9624" box="[464,605,274,298]" county="Pena Negra" latitude="42.05" longLatPrecision="1152" longitude="-2.7666667" municipality="Sierra de Urbion" name="Castilla-Leon" pageId="10" pageNumber="365">Castilla-Leon</location>
,
<collectingMunicipality id="E706ACC0F1225D0678682879FC819624" box="[616,791,274,298]" pageId="10" pageNumber="365">Sierra de Urbión</collectingMunicipality>
,
<collectingCounty id="EE034E36F1225D0679212878FC0C9625" box="[801,922,275,299]" pageId="10" pageNumber="365">Peña Negra</collectingCounty>
,
<quantity id="C0259B5FF1225D0679A52879FBA09624" box="[933,1078,274,298]" metricMagnitude="3" metricUnit="m" metricValue="1.975" metricValueMax="2.0" metricValueMin="1.95" pageId="10" pageNumber="365" unit="m" value="1975.0" valueMax="2000.0" valueMin="1950.0">
<elevation id="8CF0D189F1225D0679A52879FBA09624" box="[933,1078,274,298]" metricMagnitude="3" metricUnit="m" metricValue="1.975" metricValueMax="2.0" metricValueMin="1.95" pageId="10" pageNumber="365" unit="m" value="1975.0" valueMax="2000.0" valueMin="1950.0">1950-2000 m</elevation>
</quantity>
,
<geoCoordinate id="62E9507DF1225D067E422879FB019624" box="[1090,1175,274,298]" degrees="42" direction="north" minutes="03" orientation="latitude" pageId="10" pageNumber="365" precision="925" value="42.05">42°03N</geoCoordinate>
,
<geoCoordinate id="62E9507DF1225D067ABE285AFE859647" box="[190,275,305,329]" degrees="02" direction="west" minutes="46" orientation="longitude" pageId="10" pageNumber="365" precision="925" value="-2.7666667">02°46W</geoCoordinate>
,
<date id="7363107AF1225D067B22285AFE059647" box="[290,403,305,329]" pageId="10" pageNumber="365" value="2003-10-15">
<collectingDate id="6327E992F1225D067B22285AFE059647" box="[290,403,305,329]" pageId="10" pageNumber="365" value="2003-10-15">15.X.2003</collectingDate>
</date>
,
<collectorName id="AA28536CF1225D067BA0285AFD959647" box="[416,515,305,329]" pageId="10" pageNumber="365">V. Assing</collectorName>
/
<typeStatus id="D8668818F1225D067820285AFD1B9647" box="[544,653,305,329]" pageId="10" pageNumber="365" type="holotype">Holotypus</typeStatus>
<specimenCount id="11DBFD33F1225D06789A285FFD31964A" box="[666,679,308,324]" pageId="10" pageNumber="365" type="male">♂</specimenCount>
<taxonomicName id="C0DD4D39F1225D0678B6285AFCF9964D" authority="Assing, 2007" authorityName="Assing" authorityYear="2007" box="[694,879,305,323]" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="hamatushamatus" status="sp. nov.">
<emphasis id="35A9EAA8F1225D0678B6285AFCF9964D" box="[694,879,305,323]" italics="true" pageId="10" pageNumber="365">Proteinus hamatushamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2E9A57D3F1225D067979285AFC229646" box="[889,948,305,328]" pageId="10" pageNumber="365" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="F2BEDA7EF1225D0679EF285AFBC79647" box="[1007,1105,305,329]" pageId="10" pageNumber="365">V. Assing</determinerName>
2007&quot; (cAss)
</materialsCitation>
.
<materialsCitation id="B7B53CE7F1225D067B09283BFCF1968B" collectingDate="2003-10-17" collectorName="P. Wunderle" county="Trigaza" elevation="1950" latitude="42.256943" location="No." longLatPrecision="19" longitude="-3.2494445" municipality="Castilla-Leon" pageId="10" pageNumber="365" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="D8668818F1225D067B09283BFEFB9666" box="[265,365,336,360]" pageId="10" pageNumber="365" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="11DBFD33F1225D067B79283BFE3A966C" box="[377,428,336,360]" pageId="10" pageNumber="365" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="11DBFD33F1225D067BBE283AFE4D966C" box="[446,475,336,357]" pageId="10" pageNumber="365" type="female">1 ♀</specimenCount>
: &quot;E -
<location id="02026061F1225D067823283BFDDD9669" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9:02026061F1225D067823283BFDDD9669" box="[547,587,336,359]" county="Trigaza" latitude="42.256943" longLatPrecision="19" longitude="-3.2494445" municipality="Castilla-Leon" name="No." pageId="10" pageNumber="365">No.</location>
19,
<collectingMunicipality id="E706ACC0F1225D06787C2824FC9F9669" box="[636,777,335,359]" pageId="10" pageNumber="365">Castilla-Leon</collectingMunicipality>
,
<locationDeviation id="16D02F98F1225D067914283BFC779666" box="[788,993,335,360]" location="Burgos" pageId="10" pageNumber="365">
<quantity id="C0259B5FF1225D067914283BFCCF9669" box="[788,857,335,360]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="365" unit="km" value="40.0">40 km</quantity>
ESE Burgos
</locationDeviation>
,
<collectingCounty id="EE034E36F1225D0679EB2824FBA19669" box="[1003,1079,335,359]" pageId="10" pageNumber="365">Trigaza</collectingCounty>
,
<quantity id="C0259B5FF1225D067E442824FB019669" box="[1092,1175,335,359]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="10" pageNumber="365" unit="m" value="1950.0">
<elevation id="8CF0D189F1225D067E442824FB019669" box="[1092,1175,335,359]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="10" pageNumber="365" unit="m" value="1950.0">1950 m</elevation>
</quantity>
,
<geoCoordinate id="62E9507DF1225D067ABE2804FEA4968A" box="[190,306,367,388]" degrees="42" direction="north" minutes="15" orientation="latitude" pageId="10" pageNumber="365" precision="15" seconds="25" value="42.256943">42°15 '25N</geoCoordinate>
,
<geoCoordinate id="62E9507DF1225D067B402804FE21968B" box="[320,439,367,389]" degrees="03" direction="west" minutes="14" orientation="longitude" pageId="10" pageNumber="365" precision="15" seconds="58" value="-3.2494445">03°14 ' 58W</geoCoordinate>
,
<date id="7363107AF1225D067BC52804FDA0968B" box="[453,566,367,389]" pageId="10" pageNumber="365" value="2003-10-17">
<collectingDate id="6327E992F1225D067BC52804FDA0968B" box="[453,566,367,389]" pageId="10" pageNumber="365" value="2003-10-17">17.X.2003</collectingDate>
</date>
,
<collectorName id="AA28536CF1225D0678442804FD50968B" box="[580,710,367,389]" pageId="10" pageNumber="365">P. Wunderle</collectorName>
&quot; (cWun, cAss)
</materialsCitation>
;
<materialsCitation id="B7B53CE7F1225D0679752804FE0996CC" collectingDate="2003-04-13" collectorName="P. Wunderle" county="Penarroya" elevation="1890" latitude="40.39278" location="No." longLatPrecision="19" longitude="-0.6525" municipality="Sierra de Gudar" pageId="10" pageNumber="365" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="11DBFD33F1225D0679752804FC05968F" box="[885,915,367,389]" pageId="10" pageNumber="365" type="male">1 ♂</specimenCount>
: &quot;E -
<location id="02026061F1225D0679DE2804FB90968A" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9:02026061F1225D0679DE2804FB90968A" box="[990,1030,367,388]" county="Penarroya" latitude="40.39278" longLatPrecision="19" longitude="-0.6525" municipality="Sierra de Gudar" name="No." pageId="10" pageNumber="365">No.</location>
22, 40 km E
<location id="02026061F1225D067ABD28E6FE9796AB" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9:02026061F1225D067ABD28E6FE9796AB" box="[189,257,397,421]" county="Penarroya" latitude="40.39278" longLatPrecision="19" longitude="-0.6525" municipality="Sierra de Gudar" name="Teruel" pageId="10" pageNumber="365">Teruel</location>
,
<collectingMunicipality id="E706ACC0F1225D067B1128E6FE5796AB" box="[273,449,397,421]" pageId="10" pageNumber="365">Sierra de Gúdar</collectingMunicipality>
, path -&gt;
<collectingCounty id="EE034E36F1225D06782F28E6FD0196AB" box="[559,663,397,421]" pageId="10" pageNumber="365">Peñarroya</collectingCounty>
,
<quantity id="C0259B5FF1225D0678A828E6FC9796AA" box="[680,769,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="10" pageNumber="365" unit="m" value="1890.0">
<elevation id="8CF0D189F1225D0678A828E6FC9796AA" box="[680,769,397,421]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="10" pageNumber="365" unit="m" value="1890.0">1890 m</elevation>
</quantity>
,
<geoCoordinate id="62E9507DF1225D06791228E6FC1196AC" box="[786,903,397,421]" degrees="40" direction="north" minutes="23" orientation="latitude" pageId="10" pageNumber="365" precision="15" seconds="34" value="40.39278">40°23 '34N</geoCoordinate>
,
<geoCoordinate id="62E9507DF1225D06799828E6FB9996AC" box="[920,1039,397,418]" degrees="00" direction="west" minutes="39" orientation="longitude" pageId="10" pageNumber="365" precision="15" seconds="09" value="-0.6525">00°39 '09W</geoCoordinate>
,
<date id="7363107AF1225D067E1F28E6FB0196AC" box="[1055,1175,397,418]" pageId="10" pageNumber="365" value="2003-04-13">
<collectingDate id="6327E992F1225D067E1F28E6FB0196AC" box="[1055,1175,397,418]" pageId="10" pageNumber="365" value="2003-04-13">13.IV.2003</collectingDate>
</date>
,
<collectorName id="AA28536CF1225D067ABE28C7FEAA96CC" box="[190,316,428,450]" pageId="10" pageNumber="365">P. Wunderle</collectorName>
&quot; (cWun)
</materialsCitation>
;
<materialsCitation id="B7B53CE7F1225D067BAD28C7FCB396EC" collectingDate="2006-06-19" collectorName="M. Tronquet" country="France" location="Pyrenees-Orientales" municipality="En Brixtot" pageId="10" pageNumber="365" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="11DBFD33F1225D067BAD28C7FE7696B0" box="[429,480,428,450]" pageId="10" pageNumber="365" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="11DBFD33F1225D067BF528C6FDB196B0" box="[501,551,428,449]" pageId="10" pageNumber="365" type="female">3 ♀♀</specimenCount>
: &quot;
<collectingCountry id="7FCA762AF1225D06784428C7FD1896CA" box="[580,654,428,452]" name="France" pageId="10" pageNumber="365">France</collectingCountry>
,
<location id="02026061F1225D06789C28C0FCFD96CD" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9:02026061F1225D06789C28C0FCFD96CD" box="[668,875,427,451]" country="France" municipality="En Brixtot" name="Pyrenees-Orientales" pageId="10" pageNumber="365">Pyrénées-Orientales</location>
, La Preste, entée [sic] grotte
<collectingMunicipality id="E706ACC0F1225D067ABE28A0FEB896EC" box="[190,302,458,482]" pageId="10" pageNumber="365">En Brixtot</collectingMunicipality>
, litière,
<collectorName id="AA28536CF1225D067B8528A0FD9B96EC" box="[389,525,459,482]" pageId="10" pageNumber="365">M. Tronquet</collectorName>
,
<date id="7363107AF1225D06781A28A1FD0396EC" box="[538,661,458,482]" pageId="10" pageNumber="365" value="2006-06-19">
<collectingDate id="6327E992F1225D06781A28A1FD0396EC" box="[538,661,458,482]" pageId="10" pageNumber="365" value="2006-06-19">19.VI.2006</collectingDate>
</date>
&quot; (cTro, cAss)
</materialsCitation>
;
<materialsCitation id="B7B53CE7F1225D06793128A1FC8096F1" collectionCode="MHNG" collectorName="Coll. G. A. Lohse" country="Spain" location="La Granja" pageId="10" pageNumber="365" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="11DBFD33F1225D06793128A1FCD996D3" box="[817,847,458,482]" pageId="10" pageNumber="365" type="male">1 ♂</specimenCount>
: &quot;
<collectingCountry id="7FCA762AF1225D06796B28A1FC3C96EC" box="[875,938,458,482]" name="Spain" pageId="10" pageNumber="365">Spain</collectingCountry>
,
<location id="02026061F1225D0679B728A0FB8996EC" LSID="urn:lsid:plazi:treatment:8F7487ACF1225D067B0929DBFC8193A9:02026061F1225D0679B728A0FB8996EC" box="[951,1055,458,482]" country="Spain" name="La Granja" pageId="10" pageNumber="365">La Granja</location>
/
<taxonomicName id="C0DD4D39F1225D067E3A28A1FEB096F1" authorityName="PANDELLE. The" authorityYear="1867" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="crenulatus">Proteinus crenulatus</taxonomicName>
/
<collectorName id="AA28536CF1225D067B3C2882FE6596F1" box="[316,499,489,511]" pageId="10" pageNumber="365">Coll. G. A. Lohse</collectorName>
MHNG - 1994&quot; (
<collectionCode id="61CCAE7FF1225D0678B42882FC9896F1" box="[692,782,489,511]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="365" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FC76531F1225D067ABE2B75FC10942A" pageId="10" pageNumber="365" type="description">
<paragraph id="076236BAF1225D067ABE2B75FED69538" blockId="10.[190,320,542,566]" box="[190,320,542,566]" pageId="10" pageNumber="365">
<emphasis id="35A9EAA8F1225D067ABE2B75FED69538" bold="true" box="[190,320,542,566]" pageId="10" pageNumber="365">Description:</emphasis>
</paragraph>
<paragraph id="076236BAF1225D067ABE2B2EFCFD9575" blockId="10.[190,1179,580,635]" pageId="10" pageNumber="365">
<quantity id="C0259B5FF1225D067ABE2B2EFEAA9552" box="[190,316,581,605]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="10" pageNumber="365" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2-2.7 mm</quantity>
(abdomen extended); width of pronotum:
<quantity id="C0259B5FF1225D06790E2B2FFC3A9552" box="[782,940,580,604]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="10" pageNumber="365" unit="mm" value="0.675" valueMax="0.75" valueMin="0.6">0.60-0.75 mm</quantity>
. Coloration and other external characters as in
<taxonomicName id="C0DD4D39F1225D067BB72B0FFDA49578" authorityName="PANDELLE. The" authorityYear="1867" box="[439,562,612,630]" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="35A9EAA8F1225D067BB72B0FFDA49578" box="[439,562,612,630]" italics="true" pageId="10" pageNumber="365">P. crenulatus</emphasis>
</taxonomicName>
, distinguished only as follows:
</paragraph>
<paragraph id="076236BAF1225D067ABE2BE1FCE295AF" blockId="10.[190,884,649,673]" box="[190,884,649,673]" pageId="10" pageNumber="365">Pronotum slightly more transverse, 1.65-1.75 times as wide as long.</paragraph>
<paragraph id="076236BAF1225D067AC12BD9FC10942A" blockId="10.[190,1180,687,804]" pageId="10" pageNumber="365">
<specimenCount id="11DBFD33F1225D067AC12BD9FF5895CC" box="[193,206,690,706]" pageId="10" pageNumber="365" type="male">♂</specimenCount>
: secondary sexual characters as in
<taxonomicName id="C0DD4D39F1225D0678622BDBFD7395CC" authorityName="PANDELLE. The" authorityYear="1867" box="[610,741,688,706]" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="35A9EAA8F1225D0678622BDBFD7395CC" box="[610,741,688,706]" italics="true" pageId="10" pageNumber="365">P. crenulatus</emphasis>
</taxonomicName>
. Aedeagus larger than in
<taxonomicName id="C0DD4D39F1225D067E122BDBFB0395CC" authorityName="PANDELLE. The" authorityYear="1867" box="[1042,1173,688,706]" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="35A9EAA8F1225D067E122BDBFB0395CC" box="[1042,1173,688,706]" italics="true" pageId="10" pageNumber="365">P. crenulatus</emphasis>
</taxonomicName>
, approximately
<quantity id="C0259B5FF1225D067B552BA5FE2295E8" box="[341,436,718,742]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="365" unit="mm" value="0.48">0.48 mm</quantity>
long; ventral process apically longer and more abruptly bent; dorsal plate larger and more strongly curved in lateral view; internal sac with very short flagellum and with additional rather large and dark membraneous structure (
<figureCitation id="9FE62A3FF1225D0679092A60FCEC942A" box="[777,890,779,804]" captionStart="Figs 12-21" captionStartId="8.[194,230,1432,1454]" captionTargetBox="[205,1166,210,1400]" captionTargetId="figure-18@8.[195,1170,204,1406]" captionTargetPageId="8" captionText="Figs 12-21: Proteinus abditusabditus sp. n. (12), P.angulatusangulatus sp. n. (13-17), P. tronquetitronqueti sp. n. (18-19), and P. hamatus sp. n. (20-21): apical part of aedeagus in lateral view (12, 17); aedeagus in lateral view (13-15, 18-21) of males from Greece (13; holotype), Sicily (14), Basilicata (15), Corsica (18-19; 18: holotype), northern Spain (20; holotype), and southwestern France (21); apical part of aedeagus in dorsal view (16). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364141" httpUri="https://zenodo.org/record/5364141/files/figure.png" pageId="10" pageNumber="365">Figs 20-21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4FC76531F1225D067ABE2A54FC0794FE" pageId="10" pageNumber="365" type="etymology">
<paragraph id="076236BAF1225D067ABE2A54FE8B9478" blockId="10.[190,1179,831,886]" pageId="10" pageNumber="365">
<emphasis id="35A9EAA8F1225D067ABE2A54FEA39459" bold="true" box="[190,309,831,855]" pageId="10" pageNumber="365">Etymology:</emphasis>
The name (Latin, adjective: hooked, hook-like) refers to the shape of the apex of the aedeagus.
</paragraph>
<paragraph id="076236BAF1225D067ABE2AF8FE1D94A5" blockId="10.[190,395,915,939]" box="[190,395,915,939]" pageId="10" pageNumber="365">
<emphasis id="35A9EAA8F1225D067ABE2AF8FE1D94A5" bold="true" box="[190,395,915,939]" pageId="10" pageNumber="365">Comparative notes:</emphasis>
</paragraph>
<paragraph id="076236BAF1225D067ABE2AD1FC0794FE" blockId="10.[189,1179,953,1008]" pageId="10" pageNumber="365">
From the other species of the
<taxonomicName id="C0DD4D39F1225D067BE42AD1FDCB94C2" authorityName="PANDELLE. The" authorityYear="1867" box="[484,605,954,972]" class="Insecta" family="Staphylinidae" genus="Proteinus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="35A9EAA8F1225D067BE42AD1FDCB94C2" box="[484,605,954,972]" italics="true" pageId="10" pageNumber="365">P. crenulatus</emphasis>
</taxonomicName>
group, this species is distinguished by the larger aedeagus with an abruptly bent apex and internal structures of distinctive shape.
</paragraph>
</subSubSection>
<subSubSection id="4FC76531F1225D067ABE2D65FC8193A9" pageId="10" pageNumber="365" type="distribution">
<paragraph id="076236BAF1225D067ABE2D65FE7F932E" blockId="10.[190,489,1038,1056]" box="[190,489,1038,1056]" pageId="10" pageNumber="365">
<emphasis id="35A9EAA8F1225D067ABE2D65FE7F932E" bold="true" box="[190,489,1038,1056]" pageId="10" pageNumber="365">Distribution and bionomics:</emphasis>
</paragraph>
<paragraph id="076236BAF1225D067ABD2D58FC8193A9" blockId="10.[189,1180,1075,1192]" pageId="10" pageNumber="365">
The known distribution of this species is confined to southwestern
<collectingCountry id="7FCA762AF1225D0679822D5FFC509345" box="[898,966,1076,1099]" name="France" pageId="10" pageNumber="365">France</collectingCountry>
and northern
<collectingCountry id="7FCA762AF1225D067E622D58FB0A9345" box="[1122,1180,1075,1099]" name="Spain" pageId="10" pageNumber="365">Spain</collectingCountry>
(
<figureCitation id="9FE62A3FF1225D067AC62D39FE9B9364" box="[198,269,1106,1130]" captionStart="Map 2" captionStartId="6.[190,234,1604,1625]" captionTargetBox="[189,1172,851,1598]" captionTargetId="figure-310@6.[189,1172,851,1598]" captionTargetPageId="6" captionText="Map 2: Distributions of Proteinus abditus sp. n. (open squares), P. angulatus sp. n. (filled squares), P. hamatustus sp. n. (open circles), and P. tronqueti sp. n. (filled circles) in Europe." figureDoi="http://doi.org/10.5281/zenodo.5364139" httpUri="https://zenodo.org/record/5364139/files/figure.png" pageId="10" pageNumber="365">Map 2</figureCitation>
); the locality &quot;La Granja&quot; is ambiguous and omitted from
<figureCitation id="9FE62A3FF1225D0679792D38FC579364" box="[889,961,1106,1130]" captionStart="Map 2" captionStartId="6.[190,234,1604,1625]" captionTargetBox="[189,1172,851,1598]" captionTargetId="figure-310@6.[189,1172,851,1598]" captionTargetPageId="6" captionText="Map 2: Distributions of Proteinus abditus sp. n. (open squares), P. angulatus sp. n. (filled squares), P. hamatustus sp. n. (open circles), and P. tronqueti sp. n. (filled circles) in Europe." figureDoi="http://doi.org/10.5281/zenodo.5364139" httpUri="https://zenodo.org/record/5364139/files/figure.png" pageId="10" pageNumber="365">Map 2</figureCitation>
. The
<typeStatus id="D8668818F1225D0679FF2D38FBBD9365" box="[1023,1067,1107,1131]" pageId="10" pageNumber="365">type</typeStatus>
specimens were collected in June and October, those taken by Paul Wunderle and myself were sifted from the litter below old pine trees at altitudes of
<quantity id="C0259B5FF1225D0678832DFBFC8593A9" box="[643,787,1168,1192]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="2.0" metricValueMin="1.89" pageId="10" pageNumber="365" unit="m" value="1945.0" valueMax="2000.0" valueMin="1890.0">1890-2000 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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