290 lines
31 KiB
XML
290 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.417.7733" ID-GBIF-Dataset="6ed372e2-700c-4945-b3a0-fe5872ef38db" ID-PMC="PMC4109464" ID-Pensoft-Pub="1313-2970-417-71" ID-PubMed="25061356" ID-ZBK="4675ED7211FA4D42836CBD36B77FC296" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-417-71" ModsDocOrigin="ZooKeys 417" ModsDocTitle="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution" checkinTime="1451245732228" checkinUser="pensoft" docAuthor="Sokolov, Igor M., Reddell, James R. & Kavanaugh, David H." docDate="2014" docId="6D969D31E8F51CB86212546C7EECFD6F" docLanguage="en" docName="ZooKeys 417: 71-101" docOrigin="ZooKeys 417" docSource="http://dx.doi.org/10.3897/zookeys.417.7733" docTitle="Anillinus forthoodensis Sokolov & Reddell, sp. n." docType="treatment" docUuid="A5899D9D-93A5-45A0-BC9E-E13A8A7EBED6" docUuidSource="ZooBank" docVersion="4" lastPageNumber="87" masterDocId="FF97FFE46B074A506D13FF9B6152FFF2" masterDocTitle="Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution" masterLastPageNumber="101" masterPageNumber="71" pageNumber="84" updateTime="1668158702436" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sokolov, Igor M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reddell, James R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>417</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>71</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.417.7733</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.417.7733</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-417-71</mods:identifier>
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<mods:identifier type="ZBK">4675ED7211FA4D42836CBD36B77FC296</mods:identifier>
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<mods:identifier type="ZooBank">4675ED7211FA4D42836CBD36B77FC296</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152053695" LSID="urn:lsid:zoobank.org:act:A5899D9D-93A5-45A0-BC9E-E13A8A7EBED6" httpUri="http://treatment.plazi.org/id/6D969D31E8F51CB86212546C7EECFD6F" lastPageId="16" lastPageNumber="87" pageId="13" pageNumber="84">
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<subSubSection pageId="13" pageNumber="84" type="multiple">
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<paragraph pageId="13" pageNumber="84">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="84" type="nomenclature">
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<paragraph pageId="13" pageNumber="84">
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<taxonomicName LSID="http://zoobank.org/A5899D9D-93A5-45A0-BC9E-E13A8A7EBED6" authority="Sokolov & Reddell" class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis Sokolov & Reddell</taxonomicName>
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<taxonomicNameLabel pageId="13" pageNumber="84">sp. n.</taxonomicNameLabel>
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Figs 2B, E,
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<normalizedToken originalValue="H–I">H-I</normalizedToken>
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, 3A, C, 4
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, F, 5F, 6
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<normalizedToken originalValue="N–P">N-P</normalizedToken>
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, 7A, E, 8
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="85" pageId="13" pageNumber="84" type="type material">
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<paragraph pageId="13" pageNumber="84">Type material.</paragraph>
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<paragraph lastPageId="14" lastPageNumber="85" pageId="13" pageNumber="84">
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HOLOTYPE, a male, deposited in CAS, point-mounted, dissected, labeled: \ TX: Bell Co., Talking Crows Cave, Fort Hood, 4.V.2006, J. Fant, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #45.781 \ Holotype
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="forthoodensis">
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<pageBreakToken pageId="14" pageNumber="85" start="start">Anillinus</pageBreakToken>
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forthoodensis
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</taxonomicName>
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Sokolov & Reddell 2014 [red label] \ CAS Type No. 18872. PARATYPES: 4 males and 3 females; two males and one female, in TMM, labeled same as holotype; one male, in CNC, labeled: \ TX: Bell Co., Talking Crows Cave, Fort Hood, 2 June 2005, J. Fant, J. Reddell, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #38.153 \; one female, in TMM, labeled: \ TX: Bell Co., Nolan Creek Cave, Fort Hood, 27.IV.2007, J. Fant, J. Reddell \ Texas Memorial Museum Invertebrate Zool Coll #55.333 \; one male and one female, in TMM, labeled: \ TX: Bell Co., Bell Cave, Fort Hood, 4 March 2010, J. Fant \ Texas Memorial Museum Invertebrate Zool Coll #70.872 \. All paratypes also labeled: \ Paratype
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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Sokolov & Reddell 2014 [yellow label] \.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="85" type="type locality">
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<paragraph pageId="14" pageNumber="85">Type locality.</paragraph>
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<paragraph pageId="14" pageNumber="85">U.S.A., Texas, Bell County, Fort Hood area.</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="85" type="etymology">
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<paragraph pageId="14" pageNumber="85">Etymology.</paragraph>
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<paragraph pageId="14" pageNumber="85">The specific epithet is a Latinized adjective in the masculine form based on Fort Hood, the U. S. military post located in Texas, from the surroundings of which the new species is described.</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="85" type="recognition">
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<paragraph pageId="14" pageNumber="85">Recognition.</paragraph>
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<paragraph pageId="14" pageNumber="85">Adults of this new species are distinguished easily from those of other Texan species of the genus by the following combination of external characters: small size, markedly elongate habitus, distinctly elongate pronotum with shallowly sinuate lateral margins, and truncate apex of elytron.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="87" pageId="14" pageNumber="85" type="description">
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<paragraph pageId="14" pageNumber="85">Description.</paragraph>
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<paragraph pageId="14" pageNumber="85">
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Medium-sized for genus (SBL range 1.65-1.73 mm, mean 1.69
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.040 mm, n=4).
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</paragraph>
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<paragraph pageId="14" pageNumber="85">
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Habitus. Body form (Fig. 5F) subdepressed, subparallel, markedly elongate (WE/SBL 0.33
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.010), head large for genus compared to pronotum (WH/WPm 0.79
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.022), pronotum wide in comparison to elytra (WPm/WE 0.88
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.031).
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</paragraph>
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<paragraph pageId="14" pageNumber="85">Color. Body rufotestaceous, appendages testaceous.</paragraph>
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<paragraph pageId="14" pageNumber="85">Microsculpture. Distinct over all dorsal surfaces of head, pronotum and elytra, with slightly transverse polygonal mesh of more or less scaly appearance on elytra.</paragraph>
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<paragraph pageId="14" pageNumber="85">Head. Labium (Fig. 3A) with mental tooth; mentum and submentum separated by suture. Glossal sclerite with distinct paraglossae laterally and with two setae apically.</paragraph>
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<caption pageId="14" pageNumber="85">
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<paragraph pageId="14" pageNumber="85">
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Figure 3. SEM images of structural features of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
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species.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Head capsule, ventral aspect: A
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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(TEXAS, Bell County, Talking Crows Cave) B
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus acutipennis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="acutipennis">Anillinus acutipennis</taxonomicName>
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(TEXAS, Bell County, Talking Crows Cave)
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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Abdominal ventrites 3-5, males, latero-ventral aspect: C
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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(TEXAS, Bell County, Talking Crows Cave) D
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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(TEXAS, Travis County, Tooth Cave). gsc - glossal sclerite; m - mentum; ms - mental-submental suture; pg - paraglossa; sm - submentum; v3-v5 - abdominal ventrites. Scale bars = 0.1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="14" pageNumber="85">
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Prothorax. Pronotum (Fig. 2E) relatively long (LP/LE 0.43
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.027) and markedly elongate (WPm/LP 1.21
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.024), with lateral margins shallowly sinuate and moderately constricted posteriorly (WPm/WPp 1.29
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.016). Anterior angles indistinct, posterior angles nearly rectangular (90-100°). Width between anterior and posterior angles of approximately equal length (WPa/WPp 1.03
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.012). Basal margin slightly convex.
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</paragraph>
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<paragraph pageId="14" pageNumber="85">
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Elytra (Fig. 2
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<normalizedToken originalValue="H–I">H-I</normalizedToken>
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). Widely depressed along suture, of normal length (LE/SBL 0.56
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.010) and narrow for genus (WE/LE 0.59
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.022), with traces of 4-5 striae. Humeri distinct, rounded, in outline forming obtuse angle with longitudinal axis of body. Lateral margins subparallel, slightly divergent at basal fifth, evenly rounded to apex in apical fourth, without subapical sinuation. Vestiture of elytra short (less than one-third length of discal setae). Apex of elytron truncate (Fig. 2H, 6 specimens out of 7 investigated) or shallowly emarginate (Fig. 2I, 1 female from 7 investigated).
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</paragraph>
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<paragraph pageId="14" pageNumber="85">Legs. Male protarsomere 1 markedly dilated apico-laterally with two rows of adhesive setae ventrally (Fig. 4A). Male hind legs modified: trochanters with many minute bumps scattered across ventral surface (Fig. 4B), metafemora triangularly dilated along posteroventral margin with a small tooth at tip of dilation (Fig. 4B, F), and metatibiae with granulate posterior margin (Fig. 4C).</paragraph>
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<caption pageId="14" pageNumber="85">
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<paragraph pageId="14" pageNumber="85">
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Figure 4. Structural features of legs of males of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
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species.
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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(TEXAS, Bell County, Talking Crows Cave): A left protibia B left metafemur C left metatibia
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<normalizedToken originalValue="D–G">D-G</normalizedToken>
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Left metafemora: D
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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(TEXAS, Travis County, Tooth Cave) E
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wisemanensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="wisemanensis">Anillinus wisemanensis</taxonomicName>
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(TEXAS, Hays County, Wiseman Sink) F
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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(TEXAS, Bell County, Talking Crows Cave) G
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus comalensis" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="comalensis">Anillinus comalensis</taxonomicName>
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(TEXAS, Comal County, 7mi W New Braunfels). ac - antennal cleaner; as - articulo-setae; asp - anterior spur; asr - anterior setal row; cls - clip seta; gr - granulation; mtbr - metatibial brush; mttr - metatrochanter; psp - posterior spur; psr - posterior setal row; sb - setal band; ta1 - tarsomere 1. Scale bars:
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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= 0.1 mm;.
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<normalizedToken originalValue="D–G">D-G</normalizedToken>
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= 0.2 mm.
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</paragraph>
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</caption>
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<paragraph pageId="14" pageNumber="85">Abdomen. Ventrite 5 of male with medial depression (Fig. 3C).</paragraph>
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<paragraph lastPageId="16" lastPageNumber="87" pageId="14" pageNumber="85">
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Male genitalia. Median lobe of aedeagus (Fig. 6N) with short basal lobe, long rectangularly bent shaft, and enlarged apex, broadly rounded at tip. Dorsal margin strongly sclerotized along almost all its length. Ventral margin enlarged along entire
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<pageBreakToken pageId="15" pageNumber="86" start="start">length</pageBreakToken>
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to basal orifice, with numerous poriferous canals. Dorsal sclerite in form of a semicircular blade-like structure with characteristic basal prolongations. Without distinct ventral sclerites. Dorsal membranous field with numerous small spines lo
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<pageBreakToken pageId="16" pageNumber="87" start="start">cated</pageBreakToken>
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dorsally from dorsal sclerite. Enlarged apical area of median lobe with a dark spine-like structure. Right paramere enlarged, long and wide with numerous (>8) long setae (Fig. 6P), with length approximately two-thirds of length of the paramere. Left paramere wide, slightly enlarged apically and basally (Fig. 6O), without long setae.
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</paragraph>
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<paragraph pageId="16" pageNumber="87">Female genitalia. Gonocoxite 2 (Fig. 7A) unguiform (gc2), rather long, with slightly curved blade (bl) and narrowly rounded apex, with nematiform (ns) and two ensiform setae, with the lateral (les) of these thicker than the medial (mes). Laterotergite (lt) with 9-10 setae. Spermatheca with distal part of cornu markedly dilated. Nodulus short, ramus undifferentiated (Fig. 7E).</paragraph>
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<caption pageId="16" pageNumber="87">
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<paragraph pageId="16" pageNumber="87">
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Figure 5. Habitus images of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
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species. A
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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(TEXAS, Travis County, Tooth Cave) B
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus sinuatus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="sinuatus">Anillinus sinuatus</taxonomicName>
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(TEXAS, Bexar County) C
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wiseman" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="wiseman">Anillinus wiseman</taxonomicName>
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ensis (TEXAS, Hays County, Wiseman Sink), holotype; holotype D
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus comalensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="comalensis">Anillinus comalensis</taxonomicName>
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(TEXAS, Comal County, 7mi W New Braunfels), paratype E
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus acutipennis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="acutipennis">Anillinus acutipennis</taxonomicName>
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(TEXAS, Bell County, Hidden Pit Cave), paratype F
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
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(TEXAS, Bell County, Talking Crows Cave), holotype. Scale bar = 0.5 mm.
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</paragraph>
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</caption>
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<caption pageId="16" pageNumber="87">
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<paragraph pageId="16" pageNumber="87">
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Figure 6. Illustrations of male aedeagus of
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
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species.
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
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(TEXAS, Travis County, Tooth Cave): A median lobe, right lateral aspect B left paramere, left lateral aspect C right paramere, right lateral aspect.
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus sinuatus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="sinuatus">Anillinus sinuatus</taxonomicName>
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(TEXAS, Bexar County) D median lobe, right lateral aspect E left paramere, left lateral aspect F right paramere, right lateral aspect.
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<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wisemanensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="wisemanensis">Anillinus wisemanensis</taxonomicName>
|
||
(TEXAS, Hays County, Wiseman Sink) G median lobe, right lateral aspect H left paramere, left lateral aspect I right paramere, right lateral aspect.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wisemanensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="wisemanensis">Anillinus wisemanensis</taxonomicName>
|
||
(TEXAS, Bell County, Talking Crows Cave) J median lobe, right lateral aspect.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus comalensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="comalensis">Anillinus comalensis</taxonomicName>
|
||
(TEXAS, Comal County, 7mi W New Braunfels) K median lobe, right lateral aspect L left paramere, left lateral aspect M right paramere, right lateral aspect.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
|
||
(TEXAS, Bell County, Talking Crows Cave) N median lobe, right lateral aspect O left paramere, left lateral aspect P right paramere, right lateral aspect. aa - apical area; bl - basal lobe; dp - dorsal protuberance; ds - dorsal sclerite; sh - shaft; ss - spine-like structure; vs - ventral sclerite. Scale = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="16" pageNumber="87">
|
||
<paragraph pageId="16" pageNumber="87">
|
||
Figure 7. Illustrations of female genitalia of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
|
||
species. Ovipositor sclerites: A
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
|
||
(TEXAS, Bell County, Talking Crows Cave B
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
|
||
(TEXAS, Travis County, Tooth Cave). Spermatheca: C
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
|
||
(TEXAS, Travis County, Tooth Cave) D
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wisemanensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="wisemanensis">Anillinus wisemanensis</taxonomicName>
|
||
(TEXAS, Hays County, Wiseman Sink) E
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
|
||
(TEXAS, Bell County, Talking Crows Cave) F
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus comalensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="comalensis">Anillinus comalensis</taxonomicName>
|
||
(TEXAS, Comal County, 7mi W New Braunfels) G
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus acutipennis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="acutipennis">Anillinus acutipennis</taxonomicName>
|
||
(TEXAS, Hays County, Wiseman Sink). bl - blade of gonocoxite 2, dpc - distal part of cornu; gc1 - gonoxocite 1; gc2 - gonocoxite 2; les - lateral ensiform seta; lt - laterotergite; mes - medial ensiform seta; n - nodulus; ns - nematiform seta; ppc - proximal part of cornu; sd - spermathecal duct; sg - spermathecal gland; sp - spermatheca. Scale bars: A, B = 0.1 mm; C-G = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="87" type="geographical distribution">
|
||
<paragraph pageId="16" pageNumber="87">Geographical distribution.</paragraph>
|
||
<paragraph pageId="16" pageNumber="87">This species is known only from several caves distributed in the Fort Hood area, Bell County, Texas (Fig. 8, white quadrangle), Lampasas Cut Plain.</paragraph>
|
||
<caption pageId="16" pageNumber="87">
|
||
<paragraph pageId="16" pageNumber="87">
|
||
Figure 8. Locality records for
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
|
||
species:
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus acutipennis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="acutipennis">Anillinus acutipennis</taxonomicName>
|
||
- white stars;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
|
||
- black triangle;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus comalensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="comalensis">Anillinus comalensis</taxonomicName>
|
||
- white diamond;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus depressus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="depressus">Anillinus depressus</taxonomicName>
|
||
- black diamond;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
|
||
- white quadrangle;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus lescheni" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="lescheni">Anillinus lescheni</taxonomicName>
|
||
- black circle
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus sinuatus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="sinuatus">Anillinus sinuatus</taxonomicName>
|
||
- black quadrangle;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus wisemanensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="wisemanensis">Anillinus wisemanensis</taxonomicName>
|
||
- white triangles;
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
|
||
spp. - white circles (1 - Bell and Coryell Counties; 2 - Williamson County; 3 - Bexar County). Violet color - range of the western
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="genus">Anillinus</taxonomicName>
|
||
species in Arkansas and Oklahoma (
|
||
<bibRefCitation author="Sokolov, IM" journalOrPublisher="The Coleopterists Bulletin" pageId="27" pageNumber="98" pagination="185 - 233" title="Review of Anillinus, with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Carabidae: Trechinae: Bembidiini)." url="10.1649/611" volume="58" year="2004">Sokolov et al. 2004</bibRefCitation>
|
||
). Light blue color - recharge zone of the Edwards Aquifer, dark blue color - the Balcones Escarpment, topographic expression of the Balcones Fault Zone (Woodruff and Abbot 1986). Contour of the Cross Timbers ecological region (highlighted) were taken from Wikipedia: http://en.wikipedia.org/wiki/Cross_Timbers. Elevation scale bar is given in meters.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="87" type="way of life">
|
||
<paragraph pageId="16" pageNumber="87">Way of life.</paragraph>
|
||
<paragraph pageId="16" pageNumber="87">This species has been found only in caves. Specimens were taken in darkness on the underside of rocks shallowly embedded in soil.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="87" type="relationships">
|
||
<paragraph pageId="16" pageNumber="87">Relationships.</paragraph>
|
||
<paragraph pageId="16" pageNumber="87">
|
||
The medially depressed abdominal ventrite 5, the enlargement of the ventral margin of the median lobe and the presence of small spines on the dorsal membranous field in the internal sac of both
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus forthoodensis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="forthoodensis">Anillinus forthoodensis</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Anillinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anillinus affabilis" order="Coleoptera" pageId="16" pageNumber="87" phylum="Arthropoda" rank="species" species="affabilis">Anillinus affabilis</taxonomicName>
|
||
males suggest that they are the closest relatives among the Texan anillines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |