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<document id="332F9F989C2D3A5FD5EC204DFACC694F" ID-DOI="10.1093/isd/ixab003" ID-ISSN="2399-3421" ID-Zenodo-Dep="5384533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299435056" checkinUser="valdenar" docAuthor="Means, Jackson C., Hennen, Derek A., Tanabe, Tsutomu &amp; Marek, Paul E." docDate="2021" docId="332087BBFF8CB77644BFFC7B612EA51F" docLanguage="en" docName="ixab003.pdf" docOrigin="Insect Systematics and Diversity 5 (2), No. 1" docSource="http://dx.doi.org/10.1093/isd/ixab003" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Cleptoria Chamberlin 1939" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="CF19FFC3FF83B779442DFFF36515A159" masterDocTitle="Phylogenetic Systematics of the Millipede Family Xystodesmidae" masterLastPageNumber="26" masterPageNumber="1" pageNumber="16" updateTime="1699219996674" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3BF72AB3F3C51208DF1C9AD1E70A9AFE">Phylogenetic Systematics of the Millipede Family Xystodesmidae</mods:title>
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<mods:namePart id="AA527B25644B399126ECA6CAF5ED5B8B">Means, Jackson C.</mods:namePart>
<mods:affiliation id="5BA94AA7465023CCF87E2F062F8BFD10">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
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<mods:namePart id="2BE7E3824B0ABFA71E8C2BE7A749C7ED">Hennen, Derek A.</mods:namePart>
<mods:affiliation id="1298142F8E98FBA2AACAB6A7E5EF7AF7">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
</mods:name>
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<mods:namePart id="4999E3572D19B30B95279D523F3F1D3E">Tanabe, Tsutomu</mods:namePart>
<mods:affiliation id="4BAD9B19B31B9EBEDEBA06606606DFF4">Faculty of Advanced Science and Technology, Kumamoto</mods:affiliation>
</mods:name>
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<mods:namePart id="49223255FA8EC9595108B2B9C86D042A">Marek, Paul E.</mods:namePart>
<mods:affiliation id="2883CD48E7077067D9EAA67723B99B15">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
<mods:nameIdentifier id="3477D57E010F7C238B7E967647EAE5BD" type="email">paulemarek@gmail.com</mods:nameIdentifier>
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<mods:date id="99B065FAEBE69DB0CC136D73A5BE932D">2021</mods:date>
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<subSubSection id="F3936526FF8CB77644BFFC7B6773A2C5" box="[146,614,904,924]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="BB3636ADFF8CB77644BFFC7B6773A2C5" blockId="15.[146,614,904,924]" box="[146,614,904,924]" pageId="15" pageNumber="16">
<heading id="E07E81C1FF8CB77644BFFC7B6773A2C5" box="[146,614,904,924]" fontSize="8" level="2" pageId="15" pageNumber="16" reason="9">
<taxonomicName id="7C894D2EFF8CB77644BFFC7B65FCA2C5" ID-CoL="cf22f430-92f5-4472-8a1f-a41105577b18" authorityName="Chamberlin" authorityYear="1939" box="[146,233,904,924]" class="Diplopoda" family="Xystodesmidae" genus="Cleptoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF8CB77644BFFC7B65FCA2C5" box="[146,233,904,924]" italics="true" pageId="15" pageNumber="16">Cleptoria</emphasis>
</taxonomicName>
n. sp. Hagoods
<bibRefCitation id="DF184B5CFF8CB77645A5FC7A6773A2C5" author="Means, J. C. &amp; P. E. Marek" box="[392,614,904,924]" pageId="15" pageNumber="16" pagination="3854" refId="ref23310" refString="Means, J. C., and P. E. Marek. 2017. Is geography an accurate predictor of evolutionary history in the millipede family Xystodesmidae? PeerJ 5: e 3854." type="journal article" year="2017">Means and Marek 2017</bibRefCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8CB77644BFFC3A678DA5B0" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="BB3636ADFF8CB77644BFFC3A678DA5B0" blockId="15.[146,771,966,1257]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB77644BFFC3A644BA285" bold="true" box="[146,350,969,988]" italics="true" pageId="15" pageNumber="16">Material examined:</emphasis>
Type specimens.
<specimenCount id="AD8FFD24FF8CB776461CFC356754A285" box="[561,577,966,988]" pageId="15" pageNumber="16" type="male"></specimenCount>
<typeStatus id="6432880FFF8CB776467AFC3A67BCA284" box="[599,681,969,989]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
(
<collectionCode id="DD98AE68FF8CB77646EBFC3A6617A284" box="[710,770,969,989]" pageId="15" pageNumber="16">VTEC</collectionCode>
MPE00291),
<specimenCount id="AD8FFD24FF8CB7764534FC156450A2A3" box="[281,325,996,1018]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
<typeStatus id="6432880FFF8CB7764578FC1464B2A2A3" box="[341,423,999,1018]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
(
<collectionCode id="DD98AE68FF8CB7764593FC15671DA2A3" box="[446,520,998,1018]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="15" pageNumber="16">VMNH</collectionCode>
SPC000389), and
<specimenCount id="AD8FFD24FF8CB77646FBFC156617A2A3" box="[726,770,996,1018]" pageId="15" pageNumber="16" type="female">2 ♀</specimenCount>
<typeStatus id="6432880FFF8CB77644BFFBF665F8A541" box="[146,237,1029,1048]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
(
<collectionCode id="DD98AE68FF8CB7764528FBF76454A541" box="[261,321,1028,1048]" pageId="15" pageNumber="16">VTEC</collectionCode>
SPC000390, 394).
<typeStatus id="6432880FFF8CB776460DFBF7676DA541" box="[544,632,1028,1048]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
from South Carolina, Barnwell County, Toby Creek, near Hagoods Mill, SC-64 (
<geoCoordinate id="DEBD506AFF8CB77644B5FBB36411A50D" box="[152,260,1088,1108]" degrees="33.2119" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="5" value="33.2119">33.2119°N</geoCoordinate>
, -
<geoCoordinate id="DEBD506AFF8CB7764538FBB36494A50D" box="[277,385,1088,1108]" degrees="81.3201" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="5" value="-81.3201">81.3201°W</geoCoordinate>
, Elev:
<quantity id="7C719B48FF8CB77645EFFBB364E0A50D" box="[450,501,1088,1108]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="15" pageNumber="16" unit="m" value="46.0">46 m</quantity>
),
<date id="CF37106DFF8CB776462AFBB367D7A50D" box="[519,706,1088,1108]" pageId="15" pageNumber="16" value="2003-11-23">23 November 2003</date>
, (Coll: D. Beamer).
<typeStatus id="6432880FFF8CB776453FFBAD6478A528" box="[274,365,1118,1137]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
from South Carolina, Barnwell County, Hagoods Mill, SC-64, 351 rd m W jct w/
<collectingCountry id="C39E763DFF8CB776463EFB886770A5D6" box="[531,613,1147,1167]" name="Lebanon" pageId="15" pageNumber="16">Lebanon</collectingCountry>
Rd., nr Toby Ck (
<geoCoordinate id="DEBD506AFF8CB77644B5FB6A6405A5F4" box="[152,272,1177,1197]" degrees="33.21206" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="33.21206">33.21206°N</geoCoordinate>
, -
<geoCoordinate id="DEBD506AFF8CB776450DFB6A6499A5F4" box="[288,396,1177,1197]" degrees="81.3198" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="5" value="-81.3198">81.3198°W</geoCoordinate>
, Elev:
<quantity id="7C719B48FF8CB77645E7FB6A64E9A5F4" box="[458,508,1177,1197]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="15" pageNumber="16" unit="m" value="46.0">46 m</quantity>
),
<date id="CF37106DFF8CB7764621FB6A67BAA5F4" box="[524,687,1177,1197]" pageId="15" pageNumber="16" value="2004-10-17">17 October 2004</date>
, beneath leaf litter (
<taxonomicName id="7C894D2EFF8CB77644D9FB44646FA592" authority=", Acer" authorityName="Acer" box="[244,378,1207,1227]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="89FDEABFFF8CB77644D9FB44646FA592" box="[244,378,1207,1227]" italics="true" pageId="15" pageNumber="16">Quercus, Acer</emphasis>
</taxonomicName>
) (Coll: P. Marek, D. Beamer, M. Beamer). Material examined listed in Supp Table 2 (online only).
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8CB77644BFFAE66437A65C" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="BB3636ADFF8CB77644BFFAE66437A65C" blockId="15.[146,771,1301,1797]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB77644BFFAE665E0A471" bold="true" box="[146,245,1301,1320]" italics="true" pageId="15" pageNumber="16">Diagnosis:</emphasis>
Adult males of
<taxonomicName id="7C894D2EFF8CB77645B2FAE66753A470" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[415,582,1301,1321]" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beameri" status="sp. nov.">
<emphasis id="89FDEABFFF8CB77645B2FAE66753A470" box="[415,582,1301,1321]" italics="true" pageId="15" pageNumber="16">Sigmoria beameri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8CB776467EFAE5679EA470" box="[595,651,1302,1321]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
are distinct from other apheloriine species based on the following combination of characters:
<emphasis id="89FDEABFFF8CB776450DFAA36493A43D" bold="true" box="[288,390,1360,1380]" pageId="15" pageNumber="16">Gonopods.</emphasis>
Gonopodal acropodite broad, curving dorsomedially at an almost 90° angle at anterior bend (
<figureCitation id="23B22A28FF8CB7764689FA9D67E3A4DB" box="[676,758,1390,1410]" captionStart="Fig" captionStartId="16.[146,173,565,582]" captionTargetBox="[314,1274,147,530]" captionTargetId="figure-726@16.[314,1274,147,530]" captionTargetPageId="16" captionText="Fig. 13. Sigmoria beameri n. sp. paratype ♂ left gonopod (VTEC SPC000389). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384562" httpUri="https://zenodo.org/record/5384562/files/figure.png" pageId="15" pageNumber="16">Fig. 13A</figureCitation>
). Basal zone lacking spines or tubercles, prefemoral process absent. Medial flange thick, laminate. Acropodite peak broad, flattened, distally with a wide, laminate lateral flange which is distinctly constricted at its base, setting it off from its ventrally-directed tip, separating it from other species with a birdhead-like acropodal apex (=
<taxonomicName id="7C894D2EFF8CB77644F3F9D26420A76C" authorityName="Chamberlin" authorityYear="1939" box="[222,309,1569,1589]" class="Diplopoda" family="Xystodesmidae" genus="Cleptoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF8CB77644F3F9D26420A76C" box="[222,309,1569,1589]" italics="true" pageId="15" pageNumber="16">Cleptoria</emphasis>
</taxonomicName>
)
<taxonomicName id="7C894D2EFF8CB7764564F9D2678AA76C" authority="(Hoffman, 1967)" baseAuthorityName="Hoffman" baseAuthorityYear="1967" box="[329,671,1569,1589]" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="abbotti">
<emphasis id="89FDEABFFF8CB7764564F9D264FEA76C" box="[329,491,1569,1589]" italics="true" pageId="15" pageNumber="16">Sigmoria abbotti</emphasis>
(Hoffman, 1967)
</taxonomicName>
,
<taxonomicName id="7C894D2EFF8CB776469DF9D264D7A70A" authority="(Hoffman, 1967)" baseAuthorityName="Hoffman" baseAuthorityYear="1967" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bipraesidens">
<emphasis id="89FDEABFFF8CB776469DF9D26416A70A" italics="true" pageId="15" pageNumber="16">Sigmoria bipraesidens</emphasis>
(Hoffman, 1967)
</taxonomicName>
,
<taxonomicName id="7C894D2EFF8CB77645F5F9CC65DFA729" authority="(Chamberlin, 1939)" baseAuthorityName="Chamberlin" baseAuthorityYear="1939" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="macra">
<emphasis id="89FDEABFFF8CB77645F5F9CC6760A70B" box="[472,629,1599,1619]" italics="true" pageId="15" pageNumber="16">Sigmoria macra</emphasis>
(Chamberlin, 1939)
</taxonomicName>
,
<taxonomicName id="7C894D2EFF8CB77644FAF9AF6713A729" authority="(Bollman, 1889)" baseAuthorityName="Bollman" baseAuthorityYear="1889" box="[215,518,1628,1648]" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rileyi">
<emphasis id="89FDEABFFF8CB77644FAF9AF6474A729" box="[215,353,1628,1648]" italics="true" pageId="15" pageNumber="16">Sigmoria rileyi</emphasis>
(Bollman, 1889)
</taxonomicName>
,
<taxonomicName id="7C894D2EFF8CB776463EF9AF65D6A7D7" authority="Shelley, 1986" authorityName="Shelley" authorityYear="1986" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="89FDEABFFF8CB776463EF9AF67A0A729" box="[531,693,1628,1648]" italics="true" pageId="15" pageNumber="16">Sigmoria robusta</emphasis>
Shelley, 1986
</taxonomicName>
,
<taxonomicName id="7C894D2EFF8CB77644FFF9896737A7D7" authority="(Loomis, 1944)" baseAuthorityName="Loomis" baseAuthorityYear="1944" box="[210,546,1658,1678]" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="shelfordi">
<emphasis id="89FDEABFFF8CB77644FFF9896494A7D7" box="[210,385,1658,1678]" italics="true" pageId="15" pageNumber="16">Sigmoria shelfordi</emphasis>
(Loomis, 1944)
</taxonomicName>
, and
<taxonomicName id="7C894D2EFF8CB7764672F989640DA7F5" authority="(Shelley, 1981)" baseAuthorityName="Shelley" baseAuthorityYear="1981" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="89FDEABFFF8CB7764672F9896617A7D4" box="[607,770,1658,1678]" italics="true" pageId="15" pageNumber="16">Sigmoria arcuata</emphasis>
(Shelley, 1981)
</taxonomicName>
. Acropodite without a cingulum, as in
<taxonomicName id="7C894D2EFF8CB7764652F96B67CCA7F5" authorityName="Chamberlin" authorityYear="1939" box="[639,729,1688,1708]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF8CB7764652F96B67CCA7F5" box="[639,729,1688,1708]" italics="true" pageId="15" pageNumber="16">Brachoria</emphasis>
</taxonomicName>
and
<taxonomicName id="7C894D2EFF8CB77644BFF9456401A793" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[146,276,1718,1738]" class="Diplopoda" family="Xystodesmidae" genus="Appalachioria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF8CB77644BFF9456401A793" box="[146,276,1718,1738]" italics="true" pageId="15" pageNumber="16">Appalachioria</emphasis>
</taxonomicName>
.
<emphasis id="89FDEABFFF8CB776450FF9456449A793" bold="true" box="[290,348,1718,1738]" pageId="15" pageNumber="16">Color.</emphasis>
Tergites with light-red paranotal spots (
<figureCitation id="23B22A28FF8CB77646CDF94565B8A7BE" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="15" pageNumber="16">Fig. 5R</figureCitation>
). Dark brown background. Collum with light red anterior and lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8CB77644BFF8C160B1A3AB" pageId="15" pageNumber="16" type="description">
<paragraph id="BB3636ADFF8CB77644BFF8C16417A63A" blockId="15.[146,770,1839,1951]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB77644BFF8C16411A61C" bold="true" box="[146,260,1842,1861]" italics="true" pageId="15" pageNumber="16">Description:</emphasis>
Supp Table 3 (online only). Based on
<typeStatus id="6432880FFF8CB77646AEF8C167CFA61F" box="[643,730,1842,1862]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
(
<specimenCount id="AD8FFD24FF8CB77646C6F8DC67EEA61C" box="[747,763,1839,1861]" pageId="15" pageNumber="16" type="male"></specimenCount>
) MPE00291.
</paragraph>
<paragraph id="BB3636ADFF8CB776449FF89D646AA6C6" blockId="15.[146,770,1839,1951]" pageId="15" pageNumber="16">Measurements (mm): BL = 47.0, CW = 6.9, IW = 4.5, ISW = 1.5, B11W = 9.4, B11H = 5.8.</paragraph>
<paragraph id="BB3636ADFF8CB776471FFD2C60B1A3AB" blockId="15.[818,1444,734,754]" box="[818,1444,734,754]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB776471FFD2C6684A3AB" bold="true" box="[818,913,735,754]" italics="true" pageId="15" pageNumber="16">Variation:</emphasis>
No significant variation from the
<typeStatus id="6432880FFF8CB77640E6FD2D6009A3AB" box="[1227,1308,734,754]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
was observed.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8CB776471FFCED60B6A209" pageId="15" pageNumber="16" type="biology_ecology">
<paragraph id="BB3636ADFF8CB776471FFCED60B6A209" blockId="15.[818,1443,798,848]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB776471FFCED613AA26B" box="[818,1071,798,818]" italics="true" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB776471FFCED6697A268" bold="true" box="[818,898,798,817]" italics="true" pageId="15" pageNumber="16">Ecology:</emphasis>
<taxonomicName id="7C894D2EFF8CB77647A6FCED613AA26B" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[907,1071,798,818]" class="Diplopoda" family="Xystodesmidae" genus="Sigmoria" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beameri" status="sp. nov.">Sigmoria beameri</taxonomicName>
</emphasis>
<taxonomicNameLabel id="92CE57C4FF8CB7764014FCEC617BA26B" box="[1081,1134,799,818]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
was collected from the riparian area around Toby Creek and was found under oak and maple leaves.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8CB776471FFC8F669CA292" pageId="15" pageNumber="16" type="distribution">
<paragraph id="BB3636ADFF8CB776471FFC8F669CA292" blockId="15.[818,1442,892,971]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB776471FFC8F66BEA2D6" bold="true" box="[818,939,892,911]" italics="true" pageId="15" pageNumber="16">Distribution:</emphasis>
Only known from the
<typeStatus id="6432880FFF8CB77640A0FC8E61A1A2C9" box="[1165,1204,893,912]" pageId="15" pageNumber="16">type</typeStatus>
locality, a small wooded area around Toby Creek, located in the Coastal Plain region of
<collectingRegion id="794DF84FFF8CB7764141FC6A6693A292" country="United States of America" name="South Carolina" pageId="15" pageNumber="16">South Carolina</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8CB776471FFC04612EA51F" pageId="15" pageNumber="16" type="etymology">
<paragraph id="BB3636ADFF8CB776471FFC04612EA51F" blockId="15.[818,1442,1015,1094]" pageId="15" pageNumber="16">
<emphasis id="89FDEABFFF8CB776471FFC046688A553" bold="true" box="[818,925,1015,1034]" italics="true" pageId="15" pageNumber="16">Etymology:</emphasis>
This species is named for its discoverer, Dr. David Beamer of Nash Community College. The specific name is a genitive noun derived as a patronym.
</paragraph>
</subSubSection>
</treatment>
</document>