treatments-xml/data/33/20/87/332087BBFF88B775471FFE9E65DAA301.xml
2024-06-21 12:33:07 +02:00

268 lines
36 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="30B61F6F1218137D028D21E77AE0AA54" ID-DOI="10.1093/isd/ixab003" ID-ISSN="2399-3421" ID-Zenodo-Dep="5384533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299435056" checkinUser="valdenar" docAuthor="Means, Jackson C., Hennen, Derek A., Tanabe, Tsutomu &amp; Marek, Paul E." docDate="2021" docId="332087BBFF88B775471FFE9E65DAA301" docLanguage="en" docName="ixab003.pdf" docOrigin="Insect Systematics and Diversity 5 (2), No. 1" docSource="http://dx.doi.org/10.1093/isd/ixab003" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Brachoria camptera Means, Hennen, Marek 2021, New Species" docType="treatment" docVersion="4" lastPageNumber="13" masterDocId="CF19FFC3FF83B779442DFFF36515A159" masterDocTitle="Phylogenetic Systematics of the Millipede Family Xystodesmidae" masterLastPageNumber="26" masterPageNumber="1" pageNumber="12" updateTime="1699219996674" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="5870C76DB5F1CAB4F212D9228E891D61" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="40798DC1A97563525385EC9734BA8D75">
<mods:title id="593FD7CFE5822837B68C590C8371C0A0">Phylogenetic Systematics of the Millipede Family Xystodesmidae</mods:title>
</mods:titleInfo>
<mods:name id="69F1468274E3C9019CA4CB4F92A9ADB7" type="personal">
<mods:role id="997ED73AB2D8C64A7DE88943C48D45CC">
<mods:roleTerm id="DBADA142611B4D7BBCE33FC4759B8083">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7282547721198545C348333353207A61">Means, Jackson C.</mods:namePart>
<mods:affiliation id="416FC21BCB7FA9FA22A578C139BB7668">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
</mods:name>
<mods:name id="24FE250CB8503F461D6DCEC248D09EBE" type="personal">
<mods:role id="5F0392BA491D0B0B6DF762A1CB1A7229">
<mods:roleTerm id="6C7EFAADEE24836A7A445349AFB6E68A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47AA5E736E5ED82F64AA070A92646ABC">Hennen, Derek A.</mods:namePart>
<mods:affiliation id="2DFE783371C9C62692C57C57BCDF1C7A">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
</mods:name>
<mods:name id="7F0FDA5C2C75F126553794EE1E74B91F" type="personal">
<mods:role id="2E8CAF14902397BF5426FB31F2470A7E">
<mods:roleTerm id="90DBA266997711DF457DA9D3AFECD317">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF012E5F2517B1EC3B604E19ED428759">Tanabe, Tsutomu</mods:namePart>
<mods:affiliation id="EBD9747E382B4E088A54B5550BB673D7">Faculty of Advanced Science and Technology, Kumamoto</mods:affiliation>
</mods:name>
<mods:name id="89D30C42CF1DED9AE00D3658E65C4A8A" type="personal">
<mods:role id="0DAFB2AF1492D544A4EE8FFA39807384">
<mods:roleTerm id="EFF4930C59F771D72B95060C27AAE7B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C23F5494A1B710683149E3B1327CEA0">Marek, Paul E.</mods:namePart>
<mods:affiliation id="A1C87AB795A9ACD7F9AE3EBDFE969248">Department of Entomology, Virginia Tech, Blacksburg, VA, USA,</mods:affiliation>
<mods:nameIdentifier id="C904B610C8916D117F2F18DFCA1A9E5D" type="email">paulemarek@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E12ED7F70E9CE82D1F3939B3EE168ABC">text</mods:typeOfResource>
<mods:relatedItem id="5D975D686CBFDB94BD2B5390C7AE02CA" type="host">
<mods:titleInfo id="19A029CA5BF989161468D30F265E55FF">
<mods:title id="8AA33A8CC1BC65F7C12D1AB98D6A3FED">Insect Systematics and Diversity</mods:title>
</mods:titleInfo>
<mods:part id="BB632E8FF9374DF9BF3E50A88700206A">
<mods:date id="BA092FC0FFFD1B415D1781A67A408EC1">2021</mods:date>
<mods:detail id="86CF699EDFE019FE1B63FDB2B089ED0B" type="pubDate">
<mods:number id="5EF81B941F08ED2851EBB9E899C3630F">2021-03-01</mods:number>
</mods:detail>
<mods:detail id="74A6826756831FCFBC483424EA867255" type="volume">
<mods:number id="C1AD481D0EECCCC041F7CCFEE2492737">5</mods:number>
</mods:detail>
<mods:detail id="A882E613A9E27C7E66A7CCDEE308C538" type="issue">
<mods:number id="83CE693C6E6297D5D0DD51448705AED3">2</mods:number>
</mods:detail>
<mods:detail id="1A5B54A9C54E938B3E4A2F9B8C953397" type="numero">
<mods:number id="2A54F2E552A447E40E68731D4316E143">1</mods:number>
</mods:detail>
<mods:extent id="E41642B9DF39BD1145377BA49F8EEF28" unit="page">
<mods:start id="3A09D06AF5FB13B5893DEB71C2855D1E">1</mods:start>
<mods:end id="60373888AF011715291E8247841069B3">26</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B3A2C5CD620074A5A65BBC5D8E3D1581">
<mods:url id="869697C1E0E28AFE9C2B8C0CA93B83B6">http://dx.doi.org/10.1093/isd/ixab003</mods:url>
</mods:location>
<mods:classification id="FC5F040D6A8F40831A6ECD341FF52A4B">journal article</mods:classification>
<mods:identifier id="343770AEBD2E5826B62121BD8EFEEE72" type="DOI">10.1093/isd/ixab003</mods:identifier>
<mods:identifier id="804DB9878E704921A1F6BEBF030A7C35" type="ISSN">2399-3421</mods:identifier>
<mods:identifier id="4B304FEF37A6B49C1ACD521F07486254" type="Zenodo-Dep">5384533</mods:identifier>
<mods:identifier id="9B12A74892B2327F554A90BF2755BDD2" type="ZooBank">A55797AF-B8E6-4E46-8FD2-53B5C79FCEF8</mods:identifier>
</mods:mods>
<treatment id="332087BBFF88B775471FFE9E65DAA301" ID-DOI="http://doi.org/10.5281/zenodo.5466763" ID-Zenodo-Dep="5466763" LSID="urn:lsid:plazi:treatment:332087BBFF88B775471FFE9E65DAA301" httpUri="http://treatment.plazi.org/id/332087BBFF88B775471FFE9E65DAA301" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="F3936526FF88B772471FFE9E66E7A0AA" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="BB3636ADFF88B772471FFE9E66C3A0FE" blockId="11.[818,1367,365,423]" pageId="11" pageNumber="12">
<heading id="E07E81C1FF88B772471FFE9E66C3A0FE" bold="true" fontSize="10" level="2" pageId="11" pageNumber="12" reason="3">
<emphasis id="89FDEABFFF88B772471FFE9E66C3A0FE" bold="true" pageId="11" pageNumber="12">
<taxonomicName id="7C894D2EFF88B772471FFE9E6046A0DF" ID-CoL="86Y46" authority="Means, Hennen, Marek" authorityName="Means, Hennen, Marek" authorityYear="2021" box="[818,1363,365,390]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="camptera" status="sp. nov.">
<emphasis id="89FDEABFFF88B772471FFE9E613DA0DF" bold="true" box="[818,1064,365,390]" italics="true" pageId="11" pageNumber="12">Brachoria camptera</emphasis>
Means, Hennen, Marek
</taxonomicName>
,
<taxonomicNameLabel id="92CE57C4FF88B772471FFE7E66C3A0FE" box="[818,982,397,423]" pageId="11" pageNumber="12" rank="species">New Species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="BB3636ADFF88B772471FFE4966E7A0AA" blockId="11.[818,1381,442,499]" pageId="11" pageNumber="12">
<heading id="E07E81C1FF88B772471FFE4966E7A0AA" fontSize="9" level="1" pageId="11" pageNumber="12" reason="6">
Vernacular Name: The City Lake Mimic Millipede
<figureCitation id="23B22A28FF88B772471FFE2F66B3A0AA" box="[818,934,476,499]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="11" pageNumber="12">Figs. 5L,M</figureCitation>
and
<figureCitation id="23B22A28FF88B77247F3FE2F66FEA0AA" box="[990,1003,476,499]" captionStart="Fig" captionStartId="12.[145,172,1938,1955]" captionTargetBox="[314,1274,1453,1903]" captionTargetId="figure-555@12.[314,1274,1453,1903]" captionTargetPageId="12" captionText="Fig. 9. Brachoria camptera n. sp. holotype ♂ left gonopod (VTEC MPE01398). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384554" httpUri="https://zenodo.org/record/5384554/files/figure.png" pageId="11" pageNumber="12">9</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF88B772471FFDEC6123A262" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="BB3636ADFF88B772471FFDEC6123A262" blockId="11.[818,1442,541,827]" pageId="11" pageNumber="12">
<emphasis id="89FDEABFFF88B772471FFDEC66EBA36B" bold="true" box="[818,1022,543,562]" italics="true" pageId="11" pageNumber="12">Material examined:</emphasis>
<materialsCitation id="0BE13CF0FF88B7724039FDD36127A262" accessionNumber="MPE01398, MPE01323, MPE01403, MPE01405" collectingDate="2016-05-21" collectionCode="VTEC, VMNH" collectorName="Elev. &amp; Coll &amp; J. C. Means &amp; D. A. Hennen &amp; Material &amp; Suppl Table" county="Putnam County" elevation="303" latitude="36.13143" location="City Lake Natural Area" longLatPrecision="4" longitude="-85.4439" municipality="Cookeville" pageId="11" pageNumber="12" specimenCode="1325, 1399, 1410, 1416, 1413" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Tennessee" typeStatus="holotype">
Type specimens.
<specimenCount id="AD8FFD24FF88B77240FCFDEE61F4A36A" box="[1233,1249,541,563]" pageId="11" pageNumber="12" type="male"></specimenCount>
<typeStatus id="6432880FFF88B77240DAFDEC605CA36A" box="[1271,1353,543,563]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(
<collectionCode id="DD98AE68FF88B772414BFDEC60B7A36A" box="[1382,1442,543,563]" pageId="11" pageNumber="12">VTEC</collectionCode>
MPE01398),
<specimenCount id="AD8FFD24FF88B7724795FDCF66F7A309" box="[952,994,570,592]" pageId="11" pageNumber="12" type="male">6 ♂</specimenCount>
<typeStatus id="6432880FFF88B77247DDFDCE615EA309" box="[1008,1099,573,592]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
(
<collectionCode id="DD98AE68FF88B7724072FDCF618EA309" box="[1119,1179,572,592]" pageId="11" pageNumber="12">VTEC</collectionCode>
MPE01323,
<specimenCode id="EB2F9ED6FF88B7724105FDCF604FA309" box="[1320,1370,572,592]" collectionCode="VTEC" pageId="11" pageNumber="12">1325</specimenCode>
,
<specimenCode id="EB2F9ED6FF88B7724146FDCF6088A309" box="[1387,1437,572,592]" collectionCode="VTEC" pageId="11" pageNumber="12">1399</specimenCode>
;
<collectionCode id="DD98AE68FF88B772471FFDA8666EA337" box="[818,891,603,622]" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="11" pageNumber="12">VMNH</collectionCode>
MPE01403,
<specimenCode id="EB2F9ED6FF88B7724028FDA96122A337" box="[1029,1079,602,622]" collectionCode="VMNH" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="11" pageNumber="12">1410</specimenCode>
,
<specimenCode id="EB2F9ED6FF88B7724065FDA9616FA337" box="[1096,1146,602,622]" collectionCode="VMNH" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="11" pageNumber="12">1416</specimenCode>
), and
<specimenCount id="AD8FFD24FF88B77240EFFDA961FEA337" box="[1218,1259,600,622]" pageId="11" pageNumber="12" type="female">2 ♀</specimenCount>
<typeStatus id="6432880FFF88B77240D5FDA86046A337" box="[1272,1363,603,622]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
(
<collectionCode id="DD98AE68FF88B772414BFDA960B7A337" box="[1382,1442,602,622]" pageId="11" pageNumber="12">VTEC</collectionCode>
MPE01405,
<specimenCode id="EB2F9ED6FF88B7724780FD8466CBA3D2" box="[941,990,631,651]" collectionCode="VTEC" pageId="11" pageNumber="12">1413</specimenCode>
) from
<collectingRegion id="794DF84FFF88B772400BFD8B6191A3D2" box="[1062,1156,632,651]" country="United States of America" name="Tennessee" pageId="11" pageNumber="12">Tennessee</collectingRegion>
,
<collectingCounty id="52574E21FF88B77240BFFD8B603FA3D2" box="[1170,1322,631,651]" pageId="11" pageNumber="12">Putnam County</collectingCounty>
,
<collectingMunicipality id="5B52ACD7FF88B7724115FD84608BA3D2" box="[1336,1438,631,651]" pageId="11" pageNumber="12">Cookeville</collectingMunicipality>
,
<location id="BE566076FF88B772471FFD676119A3F1" LSID="urn:lsid:plazi:treatment:332087BBFF88B775471FFE9E65DAA301:BE566076FF88B772471FFD676119A3F1" box="[818,1036,660,680]" county="Putnam County" latitude="36.13143" longLatPrecision="4" longitude="-85.4439" municipality="Cookeville" name="City Lake Natural Area" pageId="11" pageNumber="12" stateProvince="Tennessee">City Lake Natural Area</location>
, along trail out of parking lot (
<geoCoordinate id="DEBD506AFF88B772410AFD67608BA3F1" box="[1319,1438,660,680]" degrees="36.13143" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="36.13143">36.13143°N</geoCoordinate>
, -
<geoCoordinate id="DEBD506AFF88B7724715FD4166B0A39F" box="[824,933,690,710]" degrees="85.4439" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="-85.4439">85.4439°W</geoCoordinate>
,
<collectorName id="167C537BFF88B772479AFD4166F1A39F" box="[951,996,690,710]" pageId="11" pageNumber="12">Elev.</collectorName>
<quantity id="7C719B48FF88B77247DCFD41612DA39F" box="[1009,1080,690,710]" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="11" pageNumber="12" unit="m" value="303.0">
<elevation id="30A4D19EFF88B77247DCFD41612DA39F" box="[1009,1080,690,710]" metricMagnitude="2" metricUnit="m" metricValue="3.03" pageId="11" pageNumber="12" unit="m" value="303.0">303 m</elevation>
</quantity>
),
<date id="CF37106DFF88B772407CFD4161F1A39F" box="[1105,1252,690,710]" pageId="11" pageNumber="12" value="2016-05-21">
<collectingDate id="DF73E985FF88B772407CFD4161F1A39F" box="[1105,1252,690,710]" pageId="11" pageNumber="12" value="2016-05-21">21 May 2016</collectingDate>
</date>
, ex beech, maple, deciduous woods, tuliptree, oak, umbrella magnolia, hickory, moist litter on hillsides, understory of poison ivy, honeysuckle, mayapple (
<collectorName id="167C537BFF88B7724714FCF96671A247" box="[825,868,778,798]" pageId="11" pageNumber="12">Coll</collectorName>
:
<collectorName id="167C537BFF88B772475CFCF866FDA247" box="[881,1000,778,798]" pageId="11" pageNumber="12">J. C. Means</collectorName>
,
<collectorName id="167C537BFF88B77247DBFCF86194A247" box="[1014,1153,778,798]" pageId="11" pageNumber="12">D. A. Hennen</collectorName>
).
<collectorName id="167C537BFF88B77240B9FCF961F0A247" box="[1172,1253,778,798]" pageId="11" pageNumber="12">Material</collectorName>
examined listed in
<collectorName id="167C537BFF88B772471FFCD4668AA262" box="[818,927,807,827]" pageId="11" pageNumber="12">Suppl Table</collectorName>
2 (online only)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF88B772471FFC946143A582" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="BB3636ADFF88B772471FFC946143A582" blockId="11.[818,1443,871,1243]" pageId="11" pageNumber="12">
<emphasis id="89FDEABFFF88B772471FFC946680A223" bold="true" box="[818,917,871,890]" italics="true" pageId="11" pageNumber="12">Diagnosis:</emphasis>
Adult males of
<taxonomicName id="7C894D2EFF88B7724019FC9461E5A223" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[1076,1264,871,891]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="camptera" status="sp. nov.">
<emphasis id="89FDEABFFF88B7724019FC9461E5A223" box="[1076,1264,871,891]" italics="true" pageId="11" pageNumber="12">Brachoria camptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF88B77240D4FC9B603BA222" box="[1273,1326,872,891]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
are distinct from other apheloriine species based on the following combination of characters:
<emphasis id="89FDEABFFF88B7724792FC516137A2EF" bold="true" box="[959,1058,930,950]" pageId="11" pageNumber="12">Gonopods</emphasis>
. Gonopodal acropodite D-shaped (
<figureCitation id="23B22A28FF88B7724152FC516658A28A" captionStart="Fig" captionStartId="12.[145,172,1938,1955]" captionTargetBox="[314,1274,1453,1903]" captionTargetId="figure-555@12.[314,1274,1453,1903]" captionTargetPageId="12" captionText="Fig. 9. Brachoria camptera n. sp. holotype ♂ left gonopod (VTEC MPE01398). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384554" httpUri="https://zenodo.org/record/5384554/files/figure.png" pageId="11" pageNumber="12">Fig. 9A</figureCitation>
)—not circular as in
<taxonomicName id="7C894D2EFF88B772403AFC4C6160A28A" authorityName="Chamberlin" authorityYear="1921" box="[1047,1141,959,979]" class="Diplopoda" family="Xystodesmidae" genus="Apheloria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="89FDEABFFF88B772403AFC4C6160A28A" box="[1047,1141,959,979]" italics="true" pageId="11" pageNumber="12">Apheloria</emphasis>
</taxonomicName>
or oval as in
<taxonomicName id="7C894D2EFF88B77240D3FC4C60B7A28A" box="[1278,1442,959,979]" class="Diplopoda" family="Xystodesmidae" genus="Rudiloria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="89FDEABFFF88B77240D3FC4C604CA28A" box="[1278,1369,959,979]" italics="true" pageId="11" pageNumber="12">Rudiloria</emphasis>
species
</taxonomicName>
(
<figureCitation id="23B22A28FF88B7724714FC2F6694A2A9" box="[825,897,988,1008]" captionStart="Fig" captionStartId="15.[146,173,651,668]" captionTargetBox="[314,1274,147,616]" captionTargetId="figure-543@15.[314,1274,147,616]" captionTargetPageId="15" captionText="Fig. 12. Rudiloria charityae n. sp. holotype ♂ left gonopod (VTEC MTX0185).(A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384560" httpUri="https://zenodo.org/record/5384560/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
). Acropodite (when viewed posteriorly) narrow, onethird width of tibia on leg pair 9. Acropodite gradually tapered to acuminate apex, not with a blunt apex as in
<taxonomicName id="7C894D2EFF88B7724108FBEB6089A572" authorityName="Keeton" authorityYear="1959" box="[1317,1436,1047,1067]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hubrichti">
<emphasis id="89FDEABFFF88B7724108FBEB6089A572" box="[1317,1436,1047,1067]" italics="true" pageId="11" pageNumber="12">B. hubrichti</emphasis>
</taxonomicName>
. Acropodite distal to anterior bend slightly curved cephalically. Acropodite shaft with cingulum (midlength transverse groove). Prefemur with curved sickle-shaped prefemoral process (
<figureCitation id="23B22A28FF88B772417FFB9C6081A5DA" box="[1362,1428,1135,1155]" captionStart="Fig" captionStartId="12.[145,172,1938,1955]" captionTargetBox="[314,1274,1453,1903]" captionTargetId="figure-555@12.[314,1274,1453,1903]" captionTargetPageId="12" captionText="Fig. 9. Brachoria camptera n. sp. holotype ♂ left gonopod (VTEC MPE01398). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5384554" httpUri="https://zenodo.org/record/5384554/files/figure.png" pageId="11" pageNumber="12">Fig. 9B</figureCitation>
).
<emphasis id="89FDEABFFF88B772471FFB7F667CA5F9" bold="true" box="[818,873,1164,1184]" pageId="11" pageNumber="12">Color</emphasis>
. Tergites black with 2 orange (
<figureCitation id="23B22A28FF88B772408DFB7F61F0A5F9" box="[1184,1253,1164,1184]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="11" pageNumber="12">Fig. 5L</figureCitation>
), or red (
<figureCitation id="23B22A28FF88B772416AFB7F6080A5F9" box="[1351,1429,1164,1184]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="11" pageNumber="12">Fig. 5M</figureCitation>
), paranotal spots, not with purple/purple-gray metatergal stripes as in
<taxonomicName id="7C894D2EFF88B7724766FB3B66A8A582" authorityName="Keeton" authorityYear="1959" box="[843,957,1223,1243]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hubrichti">
<emphasis id="89FDEABFFF88B7724766FB3B66A8A582" box="[843,957,1223,1243]" italics="true" pageId="11" pageNumber="12">B. hubrichti</emphasis>
</taxonomicName>
and
<taxonomicName id="7C894D2EFF88B77247C0FB3B6145A582" authorityName="Chamberlin" authorityYear="1939" box="[1005,1104,1223,1243]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="initialis">
<emphasis id="89FDEABFFF88B77247C0FB3B6145A582" box="[1005,1104,1223,1243]" italics="true" pageId="11" pageNumber="12">B. initialis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF88B775471FFAF465FCA059" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="BB3636ADFF88B772471FFAF466B7A461" blockId="11.[818,1442,1285,1395]" pageId="11" pageNumber="12">
<emphasis id="89FDEABFFF88B772471FFAF466B1A443" bold="true" box="[818,932,1287,1306]" italics="true" pageId="11" pageNumber="12">Description:</emphasis>
Supp Table 3 (online only). Based on
<typeStatus id="6432880FFF88B772410EFAF4606FA442" box="[1315,1402,1287,1307]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
(
<specimenCount id="AD8FFD24FF88B77241A6FAF6608EA442" box="[1419,1435,1285,1307]" pageId="11" pageNumber="12" type="male"></specimenCount>
) MPE01398.
</paragraph>
<paragraph id="BB3636ADFF88B772477FFAB0613EA42A" blockId="11.[818,1442,1285,1395]" pageId="11" pageNumber="12">Measurements (mm): BL = 39.0, CW = 7.6, IW = 5.0, ISW = 1.5, B11W = 10.0, B11H = 5.2.</paragraph>
<caption id="EFF66625FF88B77244BFF8616068A6FA" ID-DOI="http://doi.org/10.5281/zenodo.5384549" ID-Zenodo-Dep="5384549" box="[146,1405,1936,1955]" httpUri="https://zenodo.org/record/5384549/files/figure.png" pageId="11" pageNumber="12" startId="11.[146,173,1938,1955]" targetBox="[314,1274,1458,1903]" targetPageId="11">
<paragraph id="BB3636ADFF88B77244BFF8616068A6FA" blockId="11.[146,1405,1936,1955]" box="[146,1405,1936,1955]" pageId="11" pageNumber="12">
<emphasis id="89FDEABFFF88B77244BFF86165D6A6FA" bold="true" box="[146,195,1938,1955]" pageId="11" pageNumber="12">Fig. 8.</emphasis>
<taxonomicName id="7C894D2EFF88B77244E6F861649DA6FA" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[203,392,1938,1955]" class="Diplopoda" family="Xystodesmidae" genus="Appalachioria" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brownae" status="sp. nov.">
<emphasis id="89FDEABFFF88B77244E6F861649DA6FA" box="[203,392,1938,1955]" italics="true" pageId="11" pageNumber="12">Appalachioria brownae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF88B77245A0F86164AEA6FA" box="[397,443,1938,1955]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
holotype ♂ left gonopod (VTEC MPE01676). (A) Anterior view; (B) Medial view; (C) Posterior view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="BB3636ADFF8FB77544BFFF6765FCA059" blockId="12.[146,770,148,256]" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB77544BFFF6765E6A1FE" bold="true" box="[146,243,148,167]" italics="true" pageId="12" pageNumber="13">Variation:</emphasis>
Individuals of
<taxonomicName id="7C894D2EFF8FB77545A2FF676758A1FE" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[399,589,148,168]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camptera" status="sp. nov.">
<emphasis id="89FDEABFFF8FB77545A2FF676758A1FE" box="[399,589,148,168]" italics="true" pageId="12" pageNumber="13">Brachoria camptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8FB7754675FF66679AA1F1" box="[600,655,149,168]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
are known to exhibit two color morphs, 1) two-spotted red (
<figureCitation id="23B22A28FF8FB77546ADFF4267DBA19C" box="[640,718,177,197]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="12" pageNumber="13">Fig. 5M</figureCitation>
) and 2) two-spotted yellow-orange with a faint light blue caudal stripe (
<figureCitation id="23B22A28FF8FB77544B4FF1F65C9A059" box="[153,220,236,256]" captionStart="Fig" captionStartId="9.[146,173,1782,1799]" captionTargetBox="[284,1303,147,1747]" captionTargetId="figure-2@9.[284,1303,147,1747]" captionTargetPageId="9" captionText="Fig. 5. Color photographs of new species of xystodesmid millipedes and their color morphs. Appalachioria bondi n. sp. (AG): AThree-spotted yellow morph with yellow legs, dorsal view, B Two-spotted pink morph with pink legs, dorsal view, C Three-spotted white morph with white legs, DThree-spotted orange morph with orange legs, ETwo-spotted yellow morph with pink legs, F.Two-spotted white morph with white legs, GThree-spotted yellow morph with pink legs, dorsal view. Appalachioria sierwaldae, n. sp. (HI): H. Three-spotted morph with yellow paranotal spots and orange medial spots and orange legs. I. Threespotted yellow morph with yellow legs. Appalachioria brownae n. sp. (JK): J. Three-spotted yellow morph with yellow legs. K. Two-spotted yellow morph with yellow legs. Brachoria camptera n. sp. (LM): L. Two-spotted orange morph with white legs. M. Two-spotted red morph with red legs. Brachoria cryocybe n. sp. (N): N. Two-spotted morph with red paranotal spots and white stripe and red legs. Brachoria platana n. sp. (OP): O. Striped red morph with red legs. P. Threespotted yellow morph with yellow legs. Rudiloria charityae n. sp. (Q): Q. Three-spotted morph with yellow paranotal spots and orange medial spots and yellow legs. Sigmoria beameri n. sp. (R): R. Two-spotted red morph with light red legs. Nannaria aenigma n. sp. (ST): S. Two-spotted red morph with light red legs. T. Two-spotted white morph with white legs. Nannaria hokie n. sp. (U):Two-spotted red morph with gray legs." figureDoi="http://doi.org/10.5281/zenodo.5384543" httpUri="https://zenodo.org/record/5384543/files/figure.png" pageId="12" pageNumber="13">Fig. 5L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8FB77544BFFEDD6458A025" pageId="12" pageNumber="13" type="biology_ecology">
<paragraph id="BB3636ADFF8FB77544BFFEDD6458A025" blockId="12.[146,770,302,380]" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB77544BFFEDD65F7A018" bold="true" box="[146,226,302,321]" italics="true" pageId="12" pageNumber="13">Ecology:</emphasis>
Individuals of
<taxonomicName id="7C894D2EFF8FB775455DFEDD6730A018" authority="Means &amp; Hennen &amp; Tanabe &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Tanabe &amp; Marek" authorityYear="2021" box="[368,549,302,322]" class="Diplopoda" family="Xystodesmidae" genus="Brachoria" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="camptera" status="sp. nov.">
<emphasis id="89FDEABFFF8FB775455DFEDD6730A018" box="[368,549,302,322]" italics="true" pageId="12" pageNumber="13">Brachoria camptera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92CE57C4FF8FB7754601FEDC674AA01B" box="[556,607,303,322]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
were collected in a mesic deciduous forest with dominant tree species of oak, maple, beech, and tuliptree.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8FB77544BFFE596741A0E7" box="[146,596,426,446]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="BB3636ADFF8FB77544BFFE596741A0E7" blockId="12.[146,596,426,446]" box="[146,596,426,446]" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB77544BFFE59641EA0E4" bold="true" box="[146,267,426,445]" italics="true" pageId="12" pageNumber="13">Distribution:</emphasis>
Known only from the
<typeStatus id="6432880FFF8FB77545F3FE586710A0E7" box="[478,517,427,446]" pageId="12" pageNumber="13">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="F3936526FF8FB77544BFFE1F65DAA301" pageId="12" pageNumber="13" type="etymology">
<paragraph id="BB3636ADFF8FB77544BFFE1F65DAA301" blockId="12.[146,770,492,600]" pageId="12" pageNumber="13">
<emphasis id="89FDEABFFF8FB77544BFFE1F65E8A0A6" bold="true" box="[146,253,492,511]" italics="true" pageId="12" pageNumber="13">Etymology:</emphasis>
This species is named for the shape of its acropodite, specifically the angular nature of its anterior bend. The specific name is a noun in apposition derived from the Greek
<emphasis id="89FDEABFFF8FB7754642FDD567A9A363" box="[623,700,550,570]" italics="true" pageId="12" pageNumber="13">kampter</emphasis>
, bend, angle.
</paragraph>
</subSubSection>
</treatment>
</document>