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<document id="E97D34A057DF33CEBA2BAD44AE9CEC7E" ID-DOI="10.5281/zenodo.3677213" ID-GBIF-Dataset="e75dd27e-8506-4850-9e05-48101e710bec" ID-ISSN="1942-1354" ID-Zenodo-Dep="3677213" ID-ZooBank="E65684F6-5A77-4970-9BCD-A4CE2971CF8D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582218532698" checkinUser="felipe" docAuthor="Santos-Silva, Antonio, Nascimento, Francisco E. de L. &amp; Wappes, James E." docDate="2019" docId="03D66423CE52BA4A52BD84A025774BF0" docLanguage="en" docName="InsectaMundi.0716.1-35.pdf.imf" docOrigin="Insecta Mundi 716 (716)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Skillmania obrienorum Santos-Silva, Nascimento and Wappes 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="FFEF1C5BCE46BA5F527D8712264A4C78" masterDocTitle="Nomenclatural changes in American Apomecynini including description of new genera and species (Coleoptera: Cerambycidae)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="19" updateTime="1698720538319" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="162BBE5A00FABB3A4EF9B6A6BF101202" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="972DAC60AD220D542B1514EB738EDB7B">
<mods:title id="E048E03D325AB964DFE2EE264ECC3A7F">Nomenclatural changes in American Apomecynini including description of new genera and species (Coleoptera: Cerambycidae)</mods:title>
</mods:titleInfo>
<mods:name id="C63C8124364851982CDD72FDB03A6D39" type="personal">
<mods:role id="F2E5B5E716332EE2949C4F36FEB091B7">
<mods:roleTerm id="E783D0FB81D63F5E6C6824BFC0B0E0D8">Author</mods:roleTerm>
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<mods:namePart id="9FCB9D0E14B53980B9A603CBAF771C39">Santos-Silva, Antonio</mods:namePart>
</mods:name>
<mods:name id="D3B322BC3703E27C8FABE0B86395129D" type="personal">
<mods:role id="E922F240F436B28AA7C29EC2B2D23EB5">
<mods:roleTerm id="E1E6B65B42E406139C6164C6387F6C54">Author</mods:roleTerm>
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<mods:namePart id="F5A597BD6E8736E287EE979DCA93005D">Nascimento, Francisco E. de L.</mods:namePart>
</mods:name>
<mods:name id="12ADC296F1336CD2C68203C6CF24F7FE" type="personal">
<mods:role id="33C9323F106CB1AE418E766A7F24F415">
<mods:roleTerm id="5490C108FDA6D835FB6043F42464ADBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0D05C9F1014127087CA00C94A580160">Wappes, James E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="26FF29A7EAD8A7CFBE88FEDCAA1A55F8">text</mods:typeOfResource>
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<mods:titleInfo id="20BE3E9C2BCD19962D46B3FF36D198E7">
<mods:title id="14FAD924C12E5757942640C9BA2989C2">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="1EB9C988ED0474323DEDFF117A3F1436">
<mods:date id="3BF854EE7FD3ECFDCC926EC9AB54FC12">2019</mods:date>
<mods:detail id="205BEB288A1EA4508FC462E2D39D900F" type="pubDate">
<mods:number id="CB716B6F753278813B572B24C028369D">2019-07-26</mods:number>
</mods:detail>
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<mods:number id="BD9994114DFA54961914E0BE611F3BC6">716</mods:number>
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<mods:number id="110F6AFF03FD2F0361A296302BD56B37">716</mods:number>
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<mods:start id="97A7C10362366B6B87E5FED19828A531">1</mods:start>
<mods:end id="A9EDF913E5078BBEFEFD1A8757B9C7F3">35</mods:end>
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<mods:classification id="4120AA4FE7EDF98FF0C614D859A51595">journal article</mods:classification>
<mods:identifier id="A2A8465DE480FB1F553A12723D5EB1D8" type="DOI">10.5281/zenodo.3677213</mods:identifier>
<mods:identifier id="C6D10D29F7EE88196A66CDAC3A4A1FBD" type="GBIF-Dataset">e75dd27e-8506-4850-9e05-48101e710bec</mods:identifier>
<mods:identifier id="6C14895BBD0CCCC466AEF02DEED2DFD7" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="DD4C717780D23CE91488B9566B9E240C" type="Zenodo-Dep">3677213</mods:identifier>
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<treatment id="03D66423CE52BA4A52BD84A025774BF0" ID-DOI="http://doi.org/10.5281/zenodo.3681557" ID-GBIF-Taxon="162237967" ID-Zenodo-Dep="3681557" LSID="urn:lsid:plazi:treatment:03D66423CE52BA4A52BD84A025774BF0" httpUri="http://treatment.plazi.org/id/03D66423CE52BA4A52BD84A025774BF0" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19">
<subSubSection id="C36586BECE52BA4B52BD84A022C14FB3" box="[192,1163,946,971]" pageId="20" pageNumber="19" type="nomenclature">
<paragraph id="8BC0D535CE52BA4B52BD84A022C14FB3" blockId="20.[192,1163,946,1004]" box="[192,1163,946,971]" pageId="20" pageNumber="19">
<heading id="D0886259CE52BA4B52BD84A022C14FB3" bold="true" box="[192,1163,946,971]" fontSize="10" level="2" pageId="20" pageNumber="19" reason="3">
<emphasis id="B90B0927CE52BA4B52BD84A022C14FB3" bold="true" box="[192,1163,946,971]" pageId="20" pageNumber="19">
<taxonomicName id="4C7FAEB6CE52BA4B52BD84A0225C4FB3" ID-CoL="BJZHT" authority="Santos-Silva, Nascimento and Wappes" authorityName="Santos-Silva, Nascimento and Wappes" authorityYear="2019" box="[192,1046,946,971]" class="Insecta" family="Cerambycidae" genus="Skillmania" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="obrienorum" status="sp. nov.">
<emphasis id="B90B0927CE52BA4B52BD84A024494FB2" bold="true" box="[192,515,946,970]" italics="true" pageId="20" pageNumber="19">Skillmania obrienorum</emphasis>
Santos-Silva, Nascimento and Wappes
</taxonomicName>
,
<taxonomicNameLabel id="A238B45CCE52BA4B565E84A122C14FB3" box="[1059,1163,947,971]" pageId="20" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36586BECE52BA4A52BD84C6221E4AB9" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19" type="description">
<paragraph id="8BC0D535CE52BA4B52BD84C627184F93" blockId="20.[192,1163,946,1004]" box="[192,338,979,1004]" pageId="20" pageNumber="19">
(
<figureCitation id="1344C9B0CE52BA4B52B484C627004F93" box="[201,330,979,1004]" captionStart="Figures 5566" captionStartId="33.[192,287,1790,1812]" captionTargetBox="[270,1362,213,1756]" captionTargetId="figure@33.[270,1362,213,1756]" captionTargetPageId="33" captionText="Figures 5566. Apomecynini spp. 5559) Morrisia squamosa Chemsak and Noguera, comb. nov., male: 55) Dorsal habitus. 56) Ventral habitus. 57) Lateral habitus. 58) Head, frontal view. 59) Eye. 60) Morrisia pulchra, holotype female, eye. 6166) Skillmania obrienorum sp. nov.: 61) Dorsal habitus, holotype male. 62) Ventral habitus, holotype male. 63) Lateral habitus, holotype male. 64) Head, frontal view, holotype male. 65) Elytral apex, holotype male. 66) paratype male, elytral apex." figureDoi="http://doi.org/10.5281/zenodo.3677227" httpUri="https://zenodo.org/record/3677227/files/figure.png" pageId="20" pageNumber="19">Fig. 6166</figureCitation>
)
</paragraph>
<paragraph id="8BC0D535CE52BA4B52BD831623EA4864" blockId="20.[192,1440,1028,1053]" box="[192,1440,1028,1053]" pageId="20" pageNumber="19">
<emphasis id="B90B0927CE52BA4B52BD831627F04864" bold="true" box="[192,442,1028,1052]" pageId="20" pageNumber="19">Description. Male.</emphasis>
Integument mostly black; palpomeres reddish-brown, with yellowish-brown apex.
</paragraph>
<paragraph id="8BC0D535CE52BA4B52BD832624604AAE" blockId="20.[192,1441,1076,1750]" pageId="20" pageNumber="19">
<emphasis id="B90B0927CE52BA4B52BD8326275A4834" bold="true" box="[192,272,1076,1100]" pageId="20" pageNumber="19">Head.</emphasis>
Frons, vertex, and area behind eyes coarsely, abundantly punctate, gradually sparser toward margin of prothorax; frons with dense pale-yellow pubescence with light yellowish-brown pubescence interspersed, especially laterally; area between antennal tubercles and posterior margin of upper eye lobes with dense pale-yellow pubescence with light yellowish-brown pubescence interspersed, (the latter more abundant than in frons); area close to prothoracic margin with dense yellowish-brown pubescence; area behind eyes with pubescence as in area between upper eye lobes. Genae slightly longer than 1.5 times length of lower eye lobe; with dense yellowish-brown pubescence close to eye, gradually less dense, pale-yellow toward glabrous apex. Antennal tubercles mostly with dense light yellowish-brown pubescence. Postclypeus with abundant pale-yellow pubescence on wide central area close to frons, sparser close to anteclypeus, glabrous laterally. Labrum coplanar with anteclypeus at posterior third, inclined at anterior 2/3; nearly smooth on posterior third, finely rugose-punctate on inclined area; with sparse, long, erect brownish setae, especially laterally. Distance between upper eye lobes 1.08 times length of scape; in frontal view, distance between lower eye lobes 1.64 times length of scape. Antennae 0.83 times elytral length; scape with dense yellowish-brown pubescence dorsally, sparser laterally and ventrally; pedicel, basal 2/3 of antennomere III, narrow dorsal area of antennomere III, and base of antennomere IV with dense yellowish-brown pubescence, slightly sparser ventrally; remaining areas of antennomeres III and IV, and entire antennomeres VXI with brownish pubescence partially obscuring integument, except a few short yellowish-brown setae at apex of some antennomeres; antennomeres IIIX with long, erect dark setae ventrally, gradually sparser toward X. Antennal formula (ratio) based on length of antennomere III: scape = 0.70; pedicel = 0.27; IV = 0.61; V = 0.42; VI = 0.38; VII = 0.38; VIII = 0.35; IX = 0.34; X = 0.29; XI = 0.39.
</paragraph>
<paragraph id="8BC0D535CE52BA4A52BD81FC24BE4EFE" blockId="20.[192,1441,1774,1928]" lastBlockId="21.[192,1441,221,646]" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19">
<emphasis id="B90B0927CE52BA4B52BD81FC27664B7E" bold="true" box="[192,300,1774,1798]" pageId="20" pageNumber="19">Thorax.</emphasis>
Prothorax nearly equal in width throughout; sides slightly rounded. Pronotum coarsely, abundantly punctate; sides of anterior third with light yellowish-brown pubescence obscuring integument, with white setae interspersed, except brownish pubescent narrow area close to anterior margin and in central area; central area of anterior third with yellowish-brown pubescence, slightly darker and sparser than laterally, with white setae interspersed; central third with white pubescence, denser laterally, sparser centrally, with yellowish-brown pubescence interspersed, except large, brown pubescent irregular spot on each side of center; sides of posterior third with dense, yellowish-brown and white pubescence interspersed; central area of posterior third with yellowish-brown pubescence partially obscuring integument, except denser, yellower pubescence in center of this area. Sides of prothorax with dense yellowish-brown and white pubescence interspersed, except posterior area close to pronotum mostly white pubescent. Ventral surface of meso- and metathorax with yellowish-brown pubescence nearly obscuring integument (although sparser in central area of prosternum), with white setae interspersed in some areas, except abundant yellowish-white pubescence on mesoventral process, and narrow band along superior area of mesocoxal cavities (sides close to mesocoxal cavities glabrous). Metanepisternum and sides of metaventrite with yellowish-brown pubescence interspersed with white pubescence; wide central area of metaventrite with mostly yellowish-white pubescence with light yellowish-brown pubescence interspersed (slightly darker in some areas); anterior and posterior areas of metaventrite with yellowish-brown pubescence. Scutellum with dense pale-yellow pubescence.
</paragraph>
<paragraph id="8BC0D535CE53BA4A52BD85B225014831" blockId="21.[192,1441,672,1097]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD85B227554EC0" bold="true" box="[192,287,672,696]" pageId="21" pageNumber="20">Elytra.</emphasis>
Coarsely, abundantly punctate. Pubescence as follows: sides of basal seventh with yellowishbrown pubescence partially obscuring integument, more pale-yellow at middle of area between humerus and scutellum, sparsely, completely interspersed with white setae (more abundant on inclined area), except in three irregular spots with sparse brownish pubescence, one dorsally, two in inclined area; area between apex of basal seventh and base of posterior third white pubescent, not obscuring integument, abundantly interspersed with yellow setae (white pubescence irregularly denser in some areas, forming spots), this area widely projected along suture to apex of scutellum, and posteriorly triangularly projected along suture toward posterior fifth; posterior third with large, inverted V-shaped pubescent band, with vertex of distinctly denser, pale yellow, and “arms” with slightly sparser yellowish-brown pubescence; area close to inverted V-shaped band with sparse brownish pubescence on inner and posterior side; posterior fifth with dense yellowish pubescence, irregularly interspersed with white pubescence anteriorly and posteriorly, except yellowish-brown pubescent close to apex, and two large, irregular spots with sparse brownish pubescence, one dorsally, another laterally.
</paragraph>
<paragraph id="8BC0D535CE53BA4A52BD837023EB491C" blockId="21.[192,1441,1122,1380]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD8370274D4802" bold="true" box="[192,263,1122,1146]" pageId="21" pageNumber="20">Legs.</emphasis>
Femora with abundant yellowish-brown pubescence (more yellowish depending on light intensity) with white pubescence interspersed (white pubescence gradually denser toward metafemora). Meso- and metatibiae ventrally transversely rugose; protibiae mostly with yellowish-brown pubescence partially obscuring integument dorsally and laterally, sparser and darkened ventrally; mesotibiae with mostly yellowish-brown pubescence with white pubescence interspersed dorsally and laterally, except darkened pubescence on dorsal sulcus, sparser and darkened ventrally; metatibiae with mostly yellowish-white pubescence on basal 2/3, yellowish-brown on dorsal and lateral sides of posterior third, sparser and darkened on ventral surface of posterior third. Tarsi with mostly sparse yellowish-brown pubescence dorsally.
</paragraph>
<paragraph id="8BC0D535CE53BA4A52BD826F236749CF" blockId="21.[192,1439,1405,1463]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD826F270649ED" bold="true" box="[192,332,1405,1429]" pageId="21" pageNumber="20">Abdomen.</emphasis>
Ventrites coarsely punctate (punctures gradually coarser and denser toward V); with yellowish-brown pubescence partially obscuring integument on IIV, distinctly sparser on V.
</paragraph>
<paragraph id="8BC0D535CE53BA4A52BD82C227224A35" blockId="21.[192,1440,1488,1613]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD82C227014990" bold="true" box="[192,331,1488,1512]" pageId="21" pageNumber="20">Variation.</emphasis>
Pronotal pubescence mostly grayish-yellow centrally and posteriorly; pubescence on sides of prothorax mostly grayish-yellow, slightly more yellowish-brown in some areas; posterior fifth of elytra with large, transverse white pubescent band not reaching apex except large, brownish pubescent dorsal spot (
<figureCitation id="1344C9B0CE53BA4A537E812727114A35" box="[259,347,1589,1613]" captionStart="Figures 5566" captionStartId="33.[192,287,1790,1812]" captionTargetBox="[270,1362,213,1756]" captionTargetId="figure@33.[270,1362,213,1756]" captionTargetPageId="33" captionText="Figures 5566. Apomecynini spp. 5559) Morrisia squamosa Chemsak and Noguera, comb. nov., male: 55) Dorsal habitus. 56) Ventral habitus. 57) Lateral habitus. 58) Head, frontal view. 59) Eye. 60) Morrisia pulchra, holotype female, eye. 6166) Skillmania obrienorum sp. nov.: 61) Dorsal habitus, holotype male. 62) Ventral habitus, holotype male. 63) Lateral habitus, holotype male. 64) Head, frontal view, holotype male. 65) Elytral apex, holotype male. 66) paratype male, elytral apex." figureDoi="http://doi.org/10.5281/zenodo.3677227" httpUri="https://zenodo.org/record/3677227/files/figure.png" pageId="21" pageNumber="20">Fig. 66</figureCitation>
).
</paragraph>
<paragraph id="8BC0D535CE53BA4A52BD8174221E4AB9" blockId="21.[192,1440,1638,1729]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD817425204A06" bold="true" box="[192,874,1638,1662]" pageId="21" pageNumber="20">
Dimensions (mm),
<typeStatus id="54C46B97CE53BA4A53B4817424084A06" box="[457,578,1638,1662]" pageId="21" pageNumber="20" type="holotype">holotype</typeStatus>
male/
<typeStatus id="54C46B97CE53BA4A50E78174255E4A06" box="[666,788,1638,1662]" pageId="21" pageNumber="20" type="paratype">paratype</typeStatus>
male.
</emphasis>
Total length 10.30/11.15; prothoracic length 2.50/2.80; anterior prothoracic width 2.05/2.25; posterior prothoracic width 2.10/2.40; maximum prothoracic width 2.30/2.55; humeral width 2.60/2.95; elytral length 7.20/7.80.
</paragraph>
</subSubSection>
<subSubSection id="C36586BECE53BA4A52BD81C922204B4E" pageId="21" pageNumber="20" type="materials_examined">
<paragraph id="8BC0D535CE53BA4A52BD81C922204B4E" blockId="21.[192,1440,1753,1846]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD81C927C04A8A" bold="true" box="[192,394,1754,1779]" pageId="21" pageNumber="20">Type material.</emphasis>
<materialsCitation id="3B17DF68CE53BA4A53EF81C922EE4B6C" ID-GBIF-Occurrence="2569665502" collectingDate="1982-08-09" collectionCode="FSCA, ACMT" collectorName="C. W. and L. O'Brien &amp; G. Wibmer" country="Mexico" location="Estacion de Biologia Chamela" pageId="21" pageNumber="20" specimenCount="1" stateProvince="Jalisco" typeStatus="holotype">
<typeStatus id="54C46B97CE53BA4A53EF81C9244A4A8B" box="[402,512,1755,1779]" pageId="21" pageNumber="20" type="holotype">Holotype</typeStatus>
male from
<collectingCountry id="F36895A5CE53BA4A50F381C925494A8B" box="[654,771,1755,1779]" name="Mexico" pageId="21" pageNumber="20">MEXICO</collectingCountry>
,
<emphasis id="B90B0927CE53BA4A517281C8252F4A8A" box="[783,869,1754,1778]" italics="true" pageId="21" pageNumber="20">
<collectingRegion id="49BB1BD7CE53BA4A517281C8252F4A8A" box="[783,869,1754,1778]" country="Mexico" name="Jalisco" pageId="21" pageNumber="20">Jalisco</collectingRegion>
</emphasis>
, UNAM (
<location id="8EA083EECE53BA4A51A081C9231B4A8B" LSID="urn:lsid:plazi:treatment:03D66423CE52BA4A52BD84A025774BF0:8EA083EECE53BA4A51A081C9231B4A8B" box="[989,1361,1755,1779]" country="Mexico" name="Estacion de Biologia Chamela" pageId="21" pageNumber="20" stateProvince="Jalisco">Estación de Biologia Chamela</location>
; 200),
<date id="FFC1F3F5CE53BA4A52BD81EE27054B6C" box="[192,335,1788,1812]" pageId="21" pageNumber="20" value="1982-08-09">
<collectingDate id="EF850A1DCE53BA4A52BD81EE27054B6C" box="[192,335,1788,1812]" pageId="21" pageNumber="20" value="1982-08-09">9.VIII.1982</collectingDate>
</date>
,
<collectorName id="268AB0E3CE53BA4A532781EE24054B6C" box="[346,591,1788,1812]" pageId="21" pageNumber="20">C.W. and L. OBrien</collectorName>
and
<collectorName id="268AB0E3CE53BA4A50F481EE25444B6C" box="[649,782,1788,1812]" pageId="21" pageNumber="20">G. Wibmer</collectorName>
col. (
<collectionCode id="ED6E4DF0CE53BA4A513681EE25D24B6C" box="[843,920,1788,1812]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="20" type="Museum">FSCA</collectionCode>
, previously in
<collectionCode id="ED6E4DF0CE53BA4A563B81EE22D64B6C" box="[1094,1180,1788,1812]" pageId="21" pageNumber="20">ACMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B17DF68CE53BA4A56D381EF22204B4E" ID-GBIF-Occurrence="2569665500" pageId="21" pageNumber="20" typeStatus="paratype">
<typeStatus id="54C46B97CE53BA4A56D381EF23544B6D" box="[1198,1310,1789,1813]" pageId="21" pageNumber="20" type="paratype">Paratype</typeStatus>
male from
<collectingCountry id="F36895A5CE53BA4A52BD800C277F4B4E" box="[192,309,1822,1846]" name="Mexico" pageId="21" pageNumber="20">MEXICO</collectingCountry>
,
<emphasis id="B90B0927CE53BA4A533D800C27D14B4D" box="[320,411,1822,1845]" italics="true" pageId="21" pageNumber="20">
<collectingRegion id="49BB1BD7CE53BA4A533D800C27D14B4D" box="[320,411,1822,1845]" country="Mexico" name="Oaxaca" pageId="21" pageNumber="20">Oaxaca</collectingRegion>
</emphasis>
: near Totolapan,
<date id="FFC1F3F5CE53BA4A5009800F25484B4D" box="[628,770,1821,1845]" pageId="21" pageNumber="20" value="1965-08-05">5.VIII.1965</date>
, Flint and Ortiz col. (
<collectionCode id="ED6E4DF0CE53BA4A567A800C22144B4E" box="[1031,1118,1822,1846]" pageId="21" pageNumber="20">ACMT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36586BECE53BA4A52BD805D25774BF0" pageId="21" pageNumber="20" type="etymology">
<paragraph id="8BC0D535CE53BA4A52BD805D25774BF0" blockId="21.[192,1439,1871,1928]" pageId="21" pageNumber="20">
<emphasis id="B90B0927CE53BA4A52BD805D27114B1F" bold="true" box="[192,347,1871,1895]" pageId="21" pageNumber="20">Etymology.</emphasis>
Named for the collectors of the
<typeStatus id="54C46B97CE53BA4A50AF805D25774B1F" box="[722,829,1871,1895]" pageId="21" pageNumber="20" type="holotype">holotype</typeStatus>
, Charlie and Lois OBrien, notable insect collectors and taxonomists and close friends of Fred Skillman.
</paragraph>
</subSubSection>
</treatment>
</document>