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<mods:title id="3174B45C37DAC2F559D0E08528FB29DE">Taxonomic revision of Schausiana Viette with two new species from Guatemala and notes on biogeography and correlated tectonics (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="428ED08892180B1B2D00C13E464FACB4">Grehan, John R.</mods:namePart>
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<mods:namePart id="BEB2D699688F1E7077DDDB386A93E47E">Monzón-Sierra, José</mods:namePart>
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<mods:affiliation id="92BA176039BAA21B06A9BE16CDA246DD">Centro de Estudios Ambientales y Biodiversidad, Universidad del Valle de Guatemala, 18 Avenida 11 - 95 Zona 15, Vista Hermosa III, Guatemala, Guatemala, C. A. jmonzon @ uvg. edu. gt; https: // orcid. org / 0000 - 0002 - 0733 - 1895</mods:affiliation>
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<paragraph id="8B95993A7E21FFE34390FE5342D11F02" blockId="2.[151,462,510,537]" box="[151,462,510,537]" pageId="2" pageNumber="69">
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<taxonomicName id="4C2AE2B97E21FFE34390FE5342D11F02" ID-CoL="92MG9" authority="Viette, 1950 a" authorityName="Viette" authorityYear="1949" box="[151,462,510,537]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
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<emphasis id="B95E45287E21FFE34390FE53423A1F03" bold="true" box="[151,293,510,537]" italics="true" pageId="2" pageNumber="69">Schausiana</emphasis>
<bibRefCitation id="EFBBE4CB7E21FFE3422BFE5342D11F02" author="Viette, P. L." box="[300,462,510,536]" pageId="2" pageNumber="69" pagination="73 - 81" refId="ref15143" refString="Viette, P. L. (1950 a) Contribution a l'etude des Hepialidae (11 e note). Sur quelques especes sud-americaines. Annales de la Societe entomologique de France, 116, 73 - 81." type="journal article" year="1950">Viette, 1950a</bibRefCitation>
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Type-species: 
<taxonomicName id="4C2AE2B97E21FFE3422FFDEE41481F41" ID-CoL="97Z4T" authority="Schaus, 1901" authorityName="Schaus" authorityYear="1901" box="[296,599,579,603]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="trojesa">
<emphasis id="B95E45287E21FFE3422FFDEE42D81F41" box="[296,455,579,603]" italics="true" pageId="2" pageNumber="69">Phassus trojesa</emphasis>
<bibRefCitation id="EFBBE4CB7E21FFE342C9FDEE41481F41" author="Schaus, W." box="[462,599,579,603]" pageId="2" pageNumber="69" pagination="73 - 77" refId="ref14953" refString="Schaus, W. (1901) New species of Heterocera from tropical America-II. Journal of the New York Entomological Society, 9 (2), 73 - 77. https: // doi. org / 10.1080 / 03745480109442893" type="journal article" year="1901">Schaus, 1901</bibRefCitation>
</taxonomicName>
, by original designation; monotypic.
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<paragraph id="8B95993A7E21FFE34390FD2D40831FEE" blockId="2.[151,1437,640,757]" pageId="2" pageNumber="69">
<bibRefCitation id="EFBBE4CB7E21FFE34390FD2D42531F82" author="Viette, P. L." box="[151,332,640,664]" pageId="2" pageNumber="69" pagination="73 - 81" refId="ref15143" refString="Viette, P. L. (1950 a) Contribution a l'etude des Hepialidae (11 e note). Sur quelques especes sud-americaines. Annales de la Societe entomologique de France, 116, 73 - 81." type="journal article" year="1950">Viette (1950a: 80)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE3426EFD2D413D1F82" author="Viette, P. L." box="[361,546,640,664]" pageId="2" pageNumber="69" pagination="73 - 80" refId="ref15229" refString="Viette, P. L. (1951 a) Contribution a l'etude des Hepialidae (17 e note). Les genres et leur espece type. Lambillionea, 50 (7 - 10), 73 - 80." type="journal article" year="1951">Viette (1951a: 79)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE34147FD2D41F11F82" author="Paclt, J." box="[576,750,640,664]" pageId="2" pageNumber="69" pagination="141 - 148" refId="ref14622" refString="Paclt, J. (1953) Genera of the Hepialidae (Insecta, Lepidoptera). Journal of the Asiatic Society, 19 (2), 141 - 148." type="journal article" year="1953">Paclt (1953: 143)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE3400AFD2D47781F82" author="Edwards, M. A. &amp; Hopwood, A. T." box="[781,1127,640,665]" pageId="2" pageNumber="69" refId="ref12983" refString="Edwards, M. A. &amp; Hopwood, A. T. (1966) Nomenclator Zoologicus. Vol. 6. Zoological Society of London, London, x + 329 pp." type="book" year="1966">Edwards &amp; Hopwood (1966: 266)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE34782FD2D43FA1FAD" author="Nielsen, E. S. &amp; Robinson, G. S." pageId="2" pageNumber="69" pagination="1 - 192" refId="ref14419" refString="Nielsen, E. S. &amp; Robinson, G. S. (1983) Ghost moths of southern South America (Lepidoptera: Hepialidae) [With a summary in Spanish prepared by P. Gentili: &quot; Mariposas Fantasmas del sur de Sudamerica &quot;]. Entomograph, 4, 1 - 192, 431 figs." type="journal article" year="1983">Nielsen &amp; Robinson (1983: 18)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE34204FD32415D1FAD" author="Robinson, G. S. &amp; Nielsen, E. S." box="[259,578,671,695]" pageId="2" pageNumber="69" pagination="1 - 112" refId="ref14895" refString="Robinson, G. S. &amp; Nielsen, E. S. (1984) Hepialidae. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Checklist: Part 1. Micropterigoidea-Immoidea. W. Junk, Den Haag, pp. i-xxvii + 1 - 112." type="book chapter" year="1984">Robinson &amp; Nielsen (1984: 17)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE34166FD3240641FAD" author="Nye, I. W. B. &amp; Fletcher, D. S." box="[609,891,671,695]" pageId="2" pageNumber="69" pagination="1 - 368" refId="ref14545" refString="Nye, I. W. B. &amp; Fletcher, D. S. (1991) Microlepidoptera. In: Nye, I. W. B. (Ed.), The generic names of moths of the world. Vol. 6. Trustees of the NHMUK, London, pp. i-xxix + 1 - 368. https: // doi. org / 10.5962 / bhl. title. 119516" type="book chapter" year="1991">Nye &amp; Fletcher (1991: 273)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE3409EFD3247861FAD" author="Nielsen, E. S. &amp; Robinson, G. S. &amp; Wagner, D. L." box="[921,1177,671,695]" pageId="2" pageNumber="69" pagination="823 - 878" refId="ref14479" refString="Nielsen, E. S., Robinson, G. S. &amp; Wagner, D. L. (2000) Ghost-moths of the world: a global inventory and bibliography of the Exoporia (Mnesarchaeoidea and Hepialoidea) (Lepidoptera). Journal of Natural History, 34, 823 - 878. https: // doi. org / 10.1080 / 002229300299282" type="journal article" year="2000">
Nielsen 
<emphasis id="B95E45287E21FFE340E8FD0D47041FAD" box="[1007,1051,671,695]" italics="true" pageId="2" pageNumber="69">et al</emphasis>
. (2000: 841)
</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE347BFFD32466F1FAD" author="Grehan, J. R." box="[1208,1392,671,695]" pageId="2" pageNumber="69" pagination="43 - 63" refId="ref13156" refString="Grehan, J. R. (2010) Structural variants in the morphology of the first abdominal tergite supporting the monophyly of the Latin American genera Cibyra Walker, Druceiella Viette, Pfitzneriana Viette and Trichophassus Le Cerf (Lepidoptera: Hepialidae). Bulletin of the Buffalo Museum of Science, 39, 43 - 63." type="journal article" year="2010">Grehan (2010: 47</bibRefCitation>
; fig. V, appendix).— 
<bibRefCitation id="EFBBE4CB7E21FFE34261FD13418B1FCC" author="Mielke, C. G. C. &amp; Grehan, J. R." box="[358,660,702,726]" pageId="2" pageNumber="69" pagination="131 - 158" refId="ref14177" refString="Mielke, C. G. C. &amp; Grehan, J. R. (2012) Catalogue of the Latin American Hepialidae with taxonomic remarks (Lepidoptera). Nachrichten entomologische Vereins Apollo, Neue Folge, 32 (3 / 4), 131 - 158." type="journal article" year="2012">Mielke &amp; Grehan (2012: 148)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE341B5FD1340711FCC" author="Grehan, J. R." box="[690,878,702,726]" pageId="2" pageNumber="69" pagination="1 - 30" refId="ref13215" refString="Grehan, J. R. (2012) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Society of Natural Sciences, 42, 1 - 30." type="journal article" year="2012">Grehan (2012: 14)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE3408BFD1347A31FCC" author="Mielke, C. G. C. &amp; Grehan, J. R." box="[908,1212,702,726]" pageId="2" pageNumber="69" pagination="113 - 134" refId="ref14227" refString="Mielke, C. G. C. &amp; Grehan, J. R. (2015) Description of a new species of Phassus Walker, 1856 from Costa Rica, Pallas, gen. n., with a new species from Guatemala, and taxonomic notes on Sthenopis Packard, [1856] (Lepidoptera, Hepialidae). The European Entomologist, 7, 113 - 134." type="journal article" year="2015">Mielke &amp; Grehan (2015: 113)</bibRefCitation>
.— 
<bibRefCitation id="EFBBE4CB7E21FFE347DCFD13422F1FEE" author="Grehan, J. R. &amp; Rawlins, J. E." pageId="2" pageNumber="69" pagination="47 - 57" refId="ref13563" refString="Grehan, J. R. &amp; Rawlins, J. E. (2016) A remarkable new genus and species of ghost moth from Peru (Lepidoptera: Exoporia: Hepialidae). Annals of Carnegie Museum, 84, 47 - 57. https: // doi. org / 10.2992 / 007.084.0106" type="journal article" year="2016">Grehan &amp; Rawlins (2016: 47)</bibRefCitation>
.—Mielke &amp; 
<bibRefCitation id="EFBBE4CB7E21FFE342BEFD7141601FEE" author="Grehan, J. R. &amp; Mielke, C. G. C." box="[441,639,732,757]" pageId="2" pageNumber="69" pagination="434 - 440" refId="ref13253" refString="Grehan, J. R. &amp; Mielke, C. G. C. (2017) Re-characterization of Gymelloxes with a re-description of Gymelloxes terea from Central America (Lepidoptera: Hepialidae). Zootaxa, 4363 (3), 434 - 440. https: // doi. org / 10.11646 / zootaxa. 4363.3.8" type="journal article" year="2017">Grehan (2017: 136)</bibRefCitation>
.—Grehan 
<emphasis id="B95E45287E21FFE341F5FD7040011FEE" box="[754,798,732,756]" italics="true" pageId="2" pageNumber="69">et al</emphasis>
. (2018: 63).
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E21FFE34390FC8F41261EB3" pageId="2" pageNumber="69" type="diagnosis">
<paragraph id="8B95993A7E21FFE34390FC8F41261EB3" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE34390FC8F42151E26" bold="true" box="[151,266,802,828]" pageId="2" pageNumber="69">Diagnosis</emphasis>
. Distinguished from all other 
<taxonomicName id="4C2AE2B97E21FFE34165FC8E41C41E27" box="[610,731,803,829]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
by unique scales on the FW veins, and by the combination of the following characters: (i) labial palpus unisegmented, sometimes with remnants of segment fusion, (ii) antenna filiform, (iii) 
<specimenCount id="9D2C52B37E21FFE3422CFCC6425F1E9F" box="[299,320,875,901]" pageId="2" pageNumber="69" type="male">♂</specimenCount>
metatibia hypertrophied and androconia present, (iv) ‘hepialine’ venation, (v) 
<specimenCount id="9D2C52B37E21FFE347A3FCC647A61E9F" box="[1188,1209,875,901]" pageId="2" pageNumber="69" type="male">♂</specimenCount>
HW 1A incomplete, and (vi) 
<specimenCount id="9D2C52B37E21FFE343F1FC2242141EB3" box="[246,267,911,937]" pageId="2" pageNumber="69" type="female">♀</specimenCount>
HW CuP barely complete.
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E21FFE343C0FC1F461B1A6F" pageId="2" pageNumber="69" type="description">
<paragraph id="8B95993A7E21FFE343C0FC1F472F1ED4" blockId="2.[151,1437,802,2017]" box="[199,1072,946,974]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0FC1F42721ED6" bold="true" box="[199,365,946,972]" pageId="2" pageNumber="69">Redescription</emphasis>
. 
<emphasis id="B95E45287E21FFE3427BFC1E42DB1ED7" bold="true" box="[380,452,947,973]" pageId="2" pageNumber="69">Males</emphasis>
(
<figureCitation id="131185BF7E21FFE342D5FC1E41281ED7" box="[466,567,947,973]" captionStart="FIGURES 6–8" captionStartId="5.[151,264,1140,1164]" captionTargetBox="[154,1433,181,1116]" captionTargetId="figure-20@5.[154,1433,181,1116]" captionTargetPageId="5" captionText="FIGURES 6–8. Schausiana trojesa. 6, ♂ LT: dorsal view (USNM). 7, ♂ (CGCM 39.417): 7a, dorsal view; 7b, ventral view (CGCM). 8, ♀ (CGCM 33.184): dorsal view (CGCM). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413512" httpUri="https://zenodo.org/record/4413512/files/figure.png" pageId="2" pageNumber="69">Figs 6–7</figureCitation>
, 
<figureCitation id="131185BF7E21FFE34144FC1E414C1ED7" box="[579,595,947,973]" captionStart="FIGURES 9–11" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[232,1410,193,1899]" captionTargetId="figure-20@7.[153,1434,181,1921]" captionTargetPageId="7" captionText="FIGURES 9–11. Schausiana maishei, sp. nov. 9, ♂ PT (CGCM 32.071): 9a, dorsal view; 9b, ventral view (CGCM). 10, ♀ PT (CGCM 36.646): 10a, dorsal view; 10b, ventral view (CGCM). 11, ♀ HT: dorsal view (UVG). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413514" httpUri="https://zenodo.org/record/4413514/files/figure.png" pageId="2" pageNumber="69">9</figureCitation>
, 
<figureCitation id="131185BF7E21FFE34158FC1E41621ED7" box="[607,637,947,973]" captionStart="FIGURES 12–13" captionStartId="9.[151,264,1782,1806]" captionTargetBox="[162,1425,211,1724]" captionTargetId="figure-20@9.[151,1435,181,1757]" captionTargetPageId="9" captionText="FIGURES 12–13. Schausiana phalerus. 12, ♂ HT: 12a, dorsal view; 12b, ventral view (NHMUK). 13, ♀ (CGCM 24.122): 13a, dorsal view; 13b, ventral view (CGCM). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413516" httpUri="https://zenodo.org/record/4413516/files/figure.png" pageId="2" pageNumber="69">12</figureCitation>
, 
<figureCitation id="131185BF7E21FFE3418EFC1E41B81ED7" box="[649,679,947,973]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[161,1423,195,1893]" captionTargetId="figure-20@11.[151,1435,181,1905]" captionTargetPageId="11" captionText="FIGURES 14–16. Schausiana chalciope, sp. nov. 14, ♂ PT (CGCM 6.216): 14a, dorsal view; 14b, ventral view (CGCM). 15," figureDoi="http://doi.org/10.5281/zenodo.4413518" httpUri="https://zenodo.org/record/4413518/files/figure.png" pageId="2" pageNumber="69">14</figureCitation>
, 
<figureCitation id="131185BF7E21FFE341B4FC1E41E41ED7" box="[691,763,947,973]" captionStart="FIGURES 17–21" captionStartId="13.[151,264,1946,1970]" captionTargetBox="[152,1434,813,1922]" captionTargetId="figure-311@13.[152,1435,813,1922]" captionTargetPageId="13" captionText="FIGURES 17–21. Schausiana pharus. 17, ♂ LT: dorsal view (NHMUK). 18, ♂ (CGCM 26.702): 18a, dorsal view; 18b, ventral view (CGCM). 19, ♀ (CGCM 25.341): dorsal view (CGCM). 20, ♀ (CGCM 36.194): dorsal view (CGCM). 21, ♀ (CGCM 33.341): dorsal view (CGCM). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413520" httpUri="https://zenodo.org/record/4413520/files/figure.png" pageId="2" pageNumber="69">17–18</figureCitation>
, 
<figureCitation id="131185BF7E21FFE34000FC1E40501ED7" box="[775,847,947,973]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">23–26</figureCitation>
, 
<figureCitation id="131185BF7E21FFE3405CFC1E40D21ED7" box="[859,973,947,973]" captionStart="FIGURES 28–33" captionStartId="15.[151,264,1922,1946]" captionTargetBox="[179,1396,633,1892]" captionTargetId="figure-214@15.[167,1420,633,1898]" captionTargetPageId="15" captionText="FIGURES 28–33. Schausiana spp. abdomen (upper side, tergites; lower side, sternites). 28–29, S. trojesa: 28, ♂ (CGCM 39417); 29, ♀ (CGCM 33.184). 30, S. maishei, sp. nov., ♂ PT (CGCM 23.236). 31, S. chalciope, sp. nov., ♀ PT (CGCM 41.258). 32–33, S. pharus: 32, ♂ (CGCM 7.110); 33, ♀ (CGCM 5.797). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413524" httpUri="https://zenodo.org/record/4413524/files/figure.png" pageId="2" pageNumber="69">28, 30, 32</figureCitation>
, 
<figureCitation id="131185BF7E21FFE340DEFC1E473C1ED4" box="[985,1059,947,974]" captionStart="FIGURES 34–36" captionStartId="16.[151,264,1795,1819]" captionTargetBox="[264,1317,196,1761]" captionTargetId="figure-26@16.[242,1345,181,1771]" captionTargetPageId="16" captionText="FIGURES 34–36. Schausiana spp. ♂ genitalia. 34, S. trojesa (CGCM 39.417): 34a, ventral view; 34b, ventral view with base of valva exposed (arrow indicates the hook at the base of the valva); 34c, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 34d, lateral view. 35, S. maishei, sp. nov., ♂ PT (CGCM 23.236): 35a, ventral view (arrow indicates the hook at the base of the valva); 35b, ventral view with pseudotegumen moved upwards (arrow indicating the mesal ridge of fultura inferior); 35c, dorsal view (internal); 35d, lateral view. 36, S. pharus (CGCM 7.110): 36a, ventral view (arrow indicates the hook at the base of the valva); 36b, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 36c, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4413526" httpUri="https://zenodo.org/record/4413526/files/figure.png" pageId="2" pageNumber="69">34–36</figureCitation>
).
</paragraph>
<paragraph id="8B95993A7E21FFE343C0FC7A47461947" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0FC7A421B1EEB" box="[199,260,983,1009]" italics="true" pageId="2" pageNumber="69">Head</emphasis>
. Clypeus glabrous anteriorly, mesally projected and differentiated from the frons. Frons with piliform and porrect scales. Vertex scales as for the frons, but shorter. Eyes large, occupying 4/5 of the head in anterior view. Labial palps unisegmented (
<figureCitation id="131185BF7E21FFE342DAFBB2412D1923" box="[477,562,1055,1081]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">Fig. 23</figureCitation>
) with indentations sometimes indicating fusion of at least two palpomeres. Antenna filiform, segments cylindrical, sensilla caetica and sensilla trichodea present.
</paragraph>
<paragraph id="8B95993A7E21FFE343C0FBCA42E01843" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0FBCA423F199B" box="[199,288,1127,1153]" italics="true" pageId="2" pageNumber="69">Thorax.</emphasis>
Legs (
<figureCitation id="131185BF7E21FFE3426AFBCA42ED199B" box="[365,498,1126,1153]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">Figs 24–25</figureCitation>
): metatibia hypertrophied (twice as long as wide) with androconia; arolium absent. Venation (
<figureCitation id="131185BF7E21FFE3420DFB26424019BC" box="[266,351,1163,1190]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">Fig. 26</figureCitation>
): FW without Sc1; HW without Sc1, CuP complete, 1A incomplete, and 2A+3A complete. DFW with specialized scales along the veins, long (
<quantity id="4CD234DF7E21FFE3419EFB02401919D3" box="[665,774,1199,1225]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="2" pageNumber="69" unit="mm" value="5.5" valueMax="10.0" valueMin="1.0">1–10 mm</quantity>
) and piliform in 
<taxonomicName id="4C2AE2B97E21FFE340C2FB02473019D2" authorityName="Schaus" authorityYear="1901" box="[965,1071,1199,1225]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="trojesa">
<emphasis id="B95E45287E21FFE340C2FB0240CC19D3" box="[965,979,1199,1225]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE340E5FB02473019D2" box="[994,1071,1199,1224]" italics="true" pageId="2" pageNumber="69">trojesa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2AE2B97E21FFE34761FB0247C619D3" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[1126,1241,1199,1225]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="maishei" status="sp. nov.">
<emphasis id="B95E45287E21FFE34761FB02476B19D3" box="[1126,1140,1199,1225]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE34785FB0247C619D3" box="[1154,1241,1199,1225]" italics="true" pageId="2" pageNumber="69">maishei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E21FFE347E7FB02460019D3" box="[1248,1311,1199,1225]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B95E45287E21FFE347E7FB0247E519D3" bold="true" box="[1248,1274,1199,1225]" pageId="2" pageNumber="69">sp</emphasis>
. 
<emphasis id="B95E45287E21FFE3460EFB02460619D3" bold="true" box="[1289,1305,1199,1225]" pageId="2" pageNumber="69">n</emphasis>
.
</taxonomicNameLabel>
, white and shortened spines in 
<taxonomicName id="4C2AE2B97E21FFE3427DFB7E41BE19F7" authority="(Druce, 1887)" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" baseAuthorityName="Druce" baseAuthorityYear="1887" box="[378,673,1235,1261]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="phalerus" status="comb. nov.">
<emphasis id="B95E45287E21FFE3427DFB7E429719F7" box="[378,392,1235,1261]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE34290FB7E42E619F7" box="[407,505,1235,1261]" italics="true" pageId="2" pageNumber="69">phalerus</emphasis>
(Druce, 1887)
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E21FFE341AEFB7E400F19F7" box="[681,784,1235,1261]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B95E45287E21FFE341AEFB7E41F519F7" bold="true" box="[681,746,1235,1261]" pageId="2" pageNumber="69">comb</emphasis>
. 
<emphasis id="B95E45287E21FFE341FDFB7E401519F7" bold="true" box="[762,778,1235,1261]" pageId="2" pageNumber="69">n</emphasis>
.
</taxonomicNameLabel>
, and 
<taxonomicName id="4C2AE2B97E21FFE34055FB7E40C319F7" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[850,988,1235,1261]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="chalciope" status="sp. nov.">
<emphasis id="B95E45287E21FFE34055FB7E407F19F7" box="[850,864,1235,1261]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE34068FB7E40C319F7" box="[879,988,1235,1261]" italics="true" pageId="2" pageNumber="69">chalciope</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E21FFE340E2FB7E473A19F7" box="[997,1061,1235,1261]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B95E45287E21FFE340E2FB7E40E019F7" bold="true" box="[997,1023,1235,1261]" pageId="2" pageNumber="69">sp</emphasis>
. 
<emphasis id="B95E45287E21FFE34708FB7E470019F7" bold="true" box="[1039,1055,1235,1261]" pageId="2" pageNumber="69">n</emphasis>
.
</taxonomicNameLabel>
, and thinly piliform, longer than regular porrect scales in 
<taxonomicName id="4C2AE2B97E21FFE342AFFB5A410E180B" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" baseAuthorityName="Druce" baseAuthorityYear="1887" box="[424,529,1271,1297]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="pharus" status="comb. nov.">
<emphasis id="B95E45287E21FFE342AFFB5A42A9180B" box="[424,438,1271,1297]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE342C4FB5A410E180B" box="[451,529,1271,1297]" italics="true" pageId="2" pageNumber="69">pharus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E21FFE34110FB5A4162180B" box="[535,637,1271,1297]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B95E45287E21FFE34110FB5A4147180B" bold="true" box="[535,600,1271,1297]" pageId="2" pageNumber="69">comb</emphasis>
. 
<emphasis id="B95E45287E21FFE34161FB5A4169180B" bold="true" box="[614,630,1271,1297]" pageId="2" pageNumber="69">n</emphasis>
.
</taxonomicNameLabel>
(figured for 
<specimenCount id="9D2C52B37E21FFE3400DFB5A4000180B" box="[778,799,1271,1297]" pageId="2" pageNumber="69" type="female">♀</specimenCount>
, 
<figureCitation id="131185BF7E21FFE3402AFB5A408E180B" box="[813,913,1270,1297]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[156,1431,194,1830]" captionTargetId="figure-26@4.[156,1432,181,1841]" captionTargetPageId="4" captionText="FIGURES 1–5. Schausiana spp. FW areas and specialized scales. Areas: 1, S. trojesa ♀. Specialized scales: 2, S. trojesa ♀; 3, S. maishei, sp. nov. ♀; 4, S. chalciope, sp. nov. ♀; 5. S. pharus ♀. Arrows indicate the position of such scales." figureDoi="http://doi.org/10.5281/zenodo.4413510" httpUri="https://zenodo.org/record/4413510/files/figure.png" pageId="2" pageNumber="69">Figs 2–5</figureCitation>
). Although absent in 
<taxonomicName id="4C2AE2B97E21FFE3477AFB5A47F9180B" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" baseAuthorityName="Druce" baseAuthorityYear="1887" box="[1149,1254,1271,1297]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="pharus" status="comb. nov.">
<emphasis id="B95E45287E21FFE3477AFB5A4794180B" box="[1149,1163,1271,1297]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE3479FFB5A47F9180B" box="[1176,1254,1271,1297]" italics="true" pageId="2" pageNumber="69">pharus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E21FFE347EBFB5A4654180B" box="[1260,1355,1271,1297]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B95E45287E21FFE347EBFB5A4632180B" bold="true" box="[1260,1325,1271,1297]" pageId="2" pageNumber="69">comb</emphasis>
. 
<emphasis id="B95E45287E21FFE3463CFB5A4654180B" bold="true" box="[1339,1355,1271,1297]" pageId="2" pageNumber="69">n</emphasis>
</taxonomicNameLabel>
, all the other species have black ovoid markings in the proximal anal area of the DFW. The FW U-band generally extends posteriorly almost to the A vein.
</paragraph>
<paragraph id="8B95993A7E21FFE343C0FACE476A18BB" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0FACE422E1867" box="[199,305,1379,1405]" italics="true" pageId="2" pageNumber="69">Abdomen</emphasis>
(
<figureCitation id="131185BF7E21FFE34239FACE42FA1867" box="[318,485,1379,1405]" captionStart="FIGURES 28–33" captionStartId="15.[151,264,1922,1946]" captionTargetBox="[179,1396,633,1892]" captionTargetId="figure-214@15.[167,1420,633,1898]" captionTargetPageId="15" captionText="FIGURES 28–33. Schausiana spp. abdomen (upper side, tergites; lower side, sternites). 28–29, S. trojesa: 28, ♂ (CGCM 39417); 29, ♀ (CGCM 33.184). 30, S. maishei, sp. nov., ♂ PT (CGCM 23.236). 31, S. chalciope, sp. nov., ♀ PT (CGCM 41.258). 32–33, S. pharus: 32, ♂ (CGCM 7.110); 33, ♀ (CGCM 5.797). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413524" httpUri="https://zenodo.org/record/4413524/files/figure.png" pageId="2" pageNumber="69">Figs 28, 30, 32</figureCitation>
). Tergosternal sclerite simple, intermediate zone without any protruding knob. Sternite II compressed laterally with shallow concave edges, tergite and sternite VIII sclerotized.
</paragraph>
<paragraph id="8B95993A7E21FFE343C0FA06466F1B4F" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0FA06427518DF" box="[199,362,1451,1477]" italics="true" pageId="2" pageNumber="69">Male genitalia</emphasis>
(
<figureCitation id="131185BF7E21FFE34271FA0642E818DC" box="[374,503,1451,1478]" captionStart="FIGURES 34–36" captionStartId="16.[151,264,1795,1819]" captionTargetBox="[264,1317,196,1761]" captionTargetId="figure-26@16.[242,1345,181,1771]" captionTargetPageId="16" captionText="FIGURES 34–36. Schausiana spp. ♂ genitalia. 34, S. trojesa (CGCM 39.417): 34a, ventral view; 34b, ventral view with base of valva exposed (arrow indicates the hook at the base of the valva); 34c, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 34d, lateral view. 35, S. maishei, sp. nov., ♂ PT (CGCM 23.236): 35a, ventral view (arrow indicates the hook at the base of the valva); 35b, ventral view with pseudotegumen moved upwards (arrow indicating the mesal ridge of fultura inferior); 35c, dorsal view (internal); 35d, lateral view. 36, S. pharus (CGCM 7.110): 36a, ventral view (arrow indicates the hook at the base of the valva); 36b, dorsal view (internal; arrow indicating the mesal ridge of fultura inferior); 36c, lateral view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4413526" httpUri="https://zenodo.org/record/4413526/files/figure.png" pageId="2" pageNumber="69">Figs 34–36</figureCitation>
). Tegumen fused to the pseudotegumen, but distinguished by a less sclerotized fusion line, ventrally articulated with saccus. Saccus U-shaped on posterior and anterior margins. Tergal lobes reduced and fused to the pseudotegumen, but distinctly sclerotized; each dorso-mesally connected by a thick membrane. Pseudotegumen dorso-mesally unfused and ventro-mesally fused. Fultura inferior and fultura superior present. Valva curved with a distinct elbow, and with a basal and densely sclerotized process. Phallus without cornutus.
</paragraph>
<paragraph id="8B95993A7E21FFE343C0F9F2471B1B63" blockId="2.[151,1437,802,2017]" box="[199,1028,1630,1657]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F9F242361B63" bold="true" box="[199,297,1631,1657]" pageId="2" pageNumber="69">Females</emphasis>
(
<figureCitation id="131185BF7E21FFE3423FF9F242A61B63" box="[312,441,1630,1657]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[156,1431,194,1830]" captionTargetId="figure-26@4.[156,1432,181,1841]" captionTargetPageId="4" captionText="FIGURES 1–5. Schausiana spp. FW areas and specialized scales. Areas: 1, S. trojesa ♀. Specialized scales: 2, S. trojesa ♀; 3, S. maishei, sp. nov. ♀; 4, S. chalciope, sp. nov. ♀; 5. S. pharus ♀. Arrows indicate the position of such scales." figureDoi="http://doi.org/10.5281/zenodo.4413510" httpUri="https://zenodo.org/record/4413510/files/figure.png" pageId="2" pageNumber="69">Figs 1, 2–5</figureCitation>
, 
<figureCitation id="131185BF7E21FFE342C2F9F242CB1B63" box="[453,468,1631,1657]" captionStart="FIGURES 6–8" captionStartId="5.[151,264,1140,1164]" captionTargetBox="[154,1433,181,1116]" captionTargetId="figure-20@5.[154,1433,181,1116]" captionTargetPageId="5" captionText="FIGURES 6–8. Schausiana trojesa. 6, ♂ LT: dorsal view (USNM). 7, ♂ (CGCM 39.417): 7a, dorsal view; 7b, ventral view (CGCM). 8, ♀ (CGCM 33.184): dorsal view (CGCM). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413512" httpUri="https://zenodo.org/record/4413512/files/figure.png" pageId="2" pageNumber="69">8</figureCitation>
, 
<figureCitation id="131185BF7E21FFE342E6F9F241371B63" box="[481,552,1631,1657]" captionStart="FIGURES 9–11" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[232,1410,193,1899]" captionTargetId="figure-20@7.[153,1434,181,1921]" captionTargetPageId="7" captionText="FIGURES 9–11. Schausiana maishei, sp. nov. 9, ♂ PT (CGCM 32.071): 9a, dorsal view; 9b, ventral view (CGCM). 10, ♀ PT (CGCM 36.646): 10a, dorsal view; 10b, ventral view (CGCM). 11, ♀ HT: dorsal view (UVG). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413514" httpUri="https://zenodo.org/record/4413514/files/figure.png" pageId="2" pageNumber="69">10–11</figureCitation>
, 
<figureCitation id="131185BF7E21FFE34133F9F2414D1B63" box="[564,594,1631,1657]" captionStart="FIGURES 12–13" captionStartId="9.[151,264,1782,1806]" captionTargetBox="[162,1425,211,1724]" captionTargetId="figure-20@9.[151,1435,181,1757]" captionTargetPageId="9" captionText="FIGURES 12–13. Schausiana phalerus. 12, ♂ HT: 12a, dorsal view; 12b, ventral view (NHMUK). 13, ♀ (CGCM 24.122): 13a, dorsal view; 13b, ventral view (CGCM). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4413516" httpUri="https://zenodo.org/record/4413516/files/figure.png" pageId="2" pageNumber="69">13</figureCitation>
, 
<figureCitation id="131185BF7E21FFE34159F9F341B91B63" box="[606,678,1630,1657]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,1929,1953]" captionTargetBox="[161,1423,195,1893]" captionTargetId="figure-20@11.[151,1435,181,1905]" captionTargetPageId="11" captionText="FIGURES 14–16. Schausiana chalciope, sp. nov. 14, ♂ PT (CGCM 6.216): 14a, dorsal view; 14b, ventral view (CGCM). 15," figureDoi="http://doi.org/10.5281/zenodo.4413518" httpUri="https://zenodo.org/record/4413518/files/figure.png" pageId="2" pageNumber="69">15–16</figureCitation>
, 
<figureCitation id="131185BF7E21FFE341B5F9F2403B1B63" box="[690,804,1631,1657]" captionStart-0="FIGURES 17–21" captionStart-1="FIGURES 22–27" captionStartId-0="13.[151,264,1946,1970]" captionStartId-1="14.[151,264,1900,1924]" captionTargetBox-0="[152,1434,813,1922]" captionTargetBox-1="[159,1418,181,1870]" captionTargetId-0="figure-311@13.[152,1435,813,1922]" captionTargetId-1="figure-26@14.[158,1429,181,1877]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 17–21. Schausiana pharus. 17, ♂ LT: dorsal view (NHMUK). 18, ♂ (CGCM 26.702): 18a, dorsal view; 18b, ventral view (CGCM). 19, ♀ (CGCM 25.341): dorsal view (CGCM). 20, ♀ (CGCM 36.194): dorsal view (CGCM). 21, ♀ (CGCM 33.341): dorsal view (CGCM). Scale bar = 1 cm." captionText-1="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi-0="http://doi.org/10.5281/zenodo.4413520" figureDoi-1="http://doi.org/10.5281/zenodo.4413522" httpUri-0="https://zenodo.org/record/4413520/files/figure.png" httpUri-1="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">19–22, 27</figureCitation>
, 
<figureCitation id="131185BF7E21FFE34037F9F240BD1B63" box="[816,930,1631,1657]" captionStart="FIGURES 28–33" captionStartId="15.[151,264,1922,1946]" captionTargetBox="[179,1396,633,1892]" captionTargetId="figure-214@15.[167,1420,633,1898]" captionTargetPageId="15" captionText="FIGURES 28–33. Schausiana spp. abdomen (upper side, tergites; lower side, sternites). 28–29, S. trojesa: 28, ♂ (CGCM 39417); 29, ♀ (CGCM 33.184). 30, S. maishei, sp. nov., ♂ PT (CGCM 23.236). 31, S. chalciope, sp. nov., ♀ PT (CGCM 41.258). 32–33, S. pharus: 32, ♂ (CGCM 7.110); 33, ♀ (CGCM 5.797). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413524" httpUri="https://zenodo.org/record/4413524/files/figure.png" pageId="2" pageNumber="69">29, 31, 33</figureCitation>
, 
<figureCitation id="131185BF7E21FFE340A9F9F240E71B63" box="[942,1016,1631,1657]" captionStart="FIGURES 37–47. 37–41" captionStartId="17.[151,264,1856,1880]" captionTargetBox="[186,1396,313,1825]" captionTargetId="figure-83@17.[170,1417,309,1833]" captionTargetPageId="17" captionText="FIGURES 37–47. 37–41. Schausiana spp. ♀ genitalia. 37, S. trojesa (CGCM 33.184): 37a, posterior view; 37b, lateral view. 38, S. chalciope, sp. nov., PT (CGCM 41.258): posterior view. 39, S. pharus (CGCM 5.797): posterior view. Ductus and corpus bursae: 40, S. trojesa (CGCM 33.184); 41, S. pharus (CGCM 5.797). 42–47. Dorsal forewing cubital patch (yellow dotted line). 42, Pallas reynaudi C. Mielke &amp; Grehan, 2015. 43, Schausiana trojesa. 44, S. maishei, sp. nov. 45, S. phalerus. 46, S. chalciope, sp. nov. 47. S. pharus." figureDoi="http://doi.org/10.5281/zenodo.4413528" httpUri="https://zenodo.org/record/4413528/files/figure.png" pageId="2" pageNumber="69">37–41</figureCitation>
).
</paragraph>
<paragraph id="8B95993A7E21FFE343C0F92E40581B87" blockId="2.[151,1437,802,2017]" box="[199,839,1667,1693]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F92E421B1B87" box="[199,260,1667,1693]" italics="true" pageId="2" pageNumber="69">Head</emphasis>
. As for the 
<specimenCount id="9D2C52B37E21FFE34280F92E42801B87" box="[391,415,1667,1693]" pageId="2" pageNumber="69" type="male">♂</specimenCount>
. Labial palps unisegmented (
<figureCitation id="131185BF7E21FFE341E1F92E40251B87" box="[742,826,1667,1693]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8B95993A7E21FFE343C0F90A42F81BFF" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F90A42071BDB" box="[199,280,1703,1729]" italics="true" pageId="2" pageNumber="69">Thorax</emphasis>
. Venation (
<figureCitation id="131185BF7E21FFE3429FF90A42F21BDB" box="[408,493,1703,1729]" captionStart="FIGURES 22–27" captionStartId="14.[151,264,1900,1924]" captionTargetBox="[159,1418,181,1870]" captionTargetId="figure-26@14.[158,1429,181,1877]" captionTargetPageId="14" captionText="FIGURES 22–27. Schausiana spp. labial palpi, legs and wing venation. 22–23, labial palpi of S. trojesa: 22, ♀ (Morelos, Cuernavaca (UNAM)); 23, ♂ (Morelos, Tepoztlán (UNAM)). Unscaled. 24–25, legs: 24, S. maishei, sp. nov. (CGCM 23.236); 25, S. pharus (CGCM 7.110). Scale bar = 5 mm. 26–27, wing venation of S. trojesa: 26, ♂ (arrows indicating 1A incomplete and 2A+3A); 27, ♀ (arrow indicating CuP mesally incomplete)." figureDoi="http://doi.org/10.5281/zenodo.4413522" httpUri="https://zenodo.org/record/4413522/files/figure.png" pageId="2" pageNumber="69">Fig. 27</figureCitation>
): HW with CuP mesally barely complete, 1A and 2A+3A complete. FW, specialized scales as for the 
<specimenCount id="9D2C52B37E21FFE34257F966427A1BFF" box="[336,357,1739,1765]" pageId="2" pageNumber="69" type="male">♂</specimenCount>
(
<figureCitation id="131185BF7E21FFE34273F96642C41BFF" box="[372,475,1738,1765]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1864,1888]" captionTargetBox="[156,1431,194,1830]" captionTargetId="figure-26@4.[156,1432,181,1841]" captionTargetPageId="4" captionText="FIGURES 1–5. Schausiana spp. FW areas and specialized scales. Areas: 1, S. trojesa ♀. Specialized scales: 2, S. trojesa ♀; 3, S. maishei, sp. nov. ♀; 4, S. chalciope, sp. nov. ♀; 5. S. pharus ♀. Arrows indicate the position of such scales." figureDoi="http://doi.org/10.5281/zenodo.4413510" httpUri="https://zenodo.org/record/4413510/files/figure.png" pageId="2" pageNumber="69">Figs 2–5</figureCitation>
).
</paragraph>
<paragraph id="8B95993A7E21FFE343C0F942418E1A13" blockId="2.[151,1437,802,2017]" box="[199,657,1775,1801]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F942422E1A13" box="[199,305,1775,1801]" italics="true" pageId="2" pageNumber="69">Abdomen</emphasis>
. As for the 
<specimenCount id="9D2C52B37E21FFE342B3F94242D61A13" box="[436,457,1775,1801]" pageId="2" pageNumber="69" type="male">♂</specimenCount>
(
<figureCitation id="131185BF7E21FFE342DFF942419B1A13" box="[472,644,1775,1801]" captionStart="FIGURES 28–33" captionStartId="15.[151,264,1922,1946]" captionTargetBox="[179,1396,633,1892]" captionTargetId="figure-214@15.[167,1420,633,1898]" captionTargetPageId="15" captionText="FIGURES 28–33. Schausiana spp. abdomen (upper side, tergites; lower side, sternites). 28–29, S. trojesa: 28, ♂ (CGCM 39417); 29, ♀ (CGCM 33.184). 30, S. maishei, sp. nov., ♂ PT (CGCM 23.236). 31, S. chalciope, sp. nov., ♀ PT (CGCM 41.258). 32–33, S. pharus: 32, ♂ (CGCM 7.110); 33, ♀ (CGCM 5.797). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4413524" httpUri="https://zenodo.org/record/4413524/files/figure.png" pageId="2" pageNumber="69">Figs 29, 31, 33</figureCitation>
).
</paragraph>
<paragraph id="8B95993A7E21FFE343C0F8BE41981A4B" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F8BE42991A37" box="[199,390,1811,1837]" italics="true" pageId="2" pageNumber="69">Female genitalia</emphasis>
(
<figureCitation id="131185BF7E21FFE34292F8BE41071A37" box="[405,536,1811,1837]" captionStart="FIGURES 37–47. 37–41" captionStartId="17.[151,264,1856,1880]" captionTargetBox="[186,1396,313,1825]" captionTargetId="figure-83@17.[170,1417,309,1833]" captionTargetPageId="17" captionText="FIGURES 37–47. 37–41. Schausiana spp. ♀ genitalia. 37, S. trojesa (CGCM 33.184): 37a, posterior view; 37b, lateral view. 38, S. chalciope, sp. nov., PT (CGCM 41.258): posterior view. 39, S. pharus (CGCM 5.797): posterior view. Ductus and corpus bursae: 40, S. trojesa (CGCM 33.184); 41, S. pharus (CGCM 5.797). 42–47. Dorsal forewing cubital patch (yellow dotted line). 42, Pallas reynaudi C. Mielke &amp; Grehan, 2015. 43, Schausiana trojesa. 44, S. maishei, sp. nov. 45, S. phalerus. 46, S. chalciope, sp. nov. 47. S. pharus." figureDoi="http://doi.org/10.5281/zenodo.4413528" httpUri="https://zenodo.org/record/4413528/files/figure.png" pageId="2" pageNumber="69">Figs 37–41</figureCitation>
). Corpus bursae with diverticulum for at least 
<taxonomicName id="4C2AE2B97E21FFE34718F8BE47971A36" authorityName="Schaus" authorityYear="1901" box="[1055,1160,1811,1837]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="trojesa">
<emphasis id="B95E45287E21FFE34718F8BE47321A37" box="[1055,1069,1811,1837]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE3473CF8BE47971A36" box="[1083,1160,1811,1836]" italics="true" pageId="2" pageNumber="69">trojesa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2AE2B97E21FFE347B9F8BE46371A37" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" baseAuthorityName="Druce" baseAuthorityYear="1887" box="[1214,1320,1811,1837]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="pharus" status="comb. nov.">
<emphasis id="B95E45287E21FFE347B9F8BE47D31A37" box="[1214,1228,1811,1837]" italics="true" pageId="2" pageNumber="69">S</emphasis>
. 
<emphasis id="B95E45287E21FFE347DDF8BE46371A37" box="[1242,1320,1811,1837]" italics="true" pageId="2" pageNumber="69">pharus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E21FFE34628F8BE468B1A37" box="[1327,1428,1811,1837]" pageId="2" pageNumber="69" rank="species">
<emphasis id="B95E45287E21FFE34628F8BE466F1A37" bold="true" box="[1327,1392,1811,1837]" pageId="2" pageNumber="69">comb</emphasis>
. 
<emphasis id="B95E45287E21FFE34678F8BE46901A37" bold="true" box="[1407,1423,1811,1837]" pageId="2" pageNumber="69">n</emphasis>
.
</taxonomicNameLabel>
, where is the internal genitalia are preserved.
</paragraph>
<paragraph id="8B95993A7E21FFE343C0F8F7461B1A6F" blockId="2.[151,1437,802,2017]" box="[199,1284,1882,1909]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F8F742E11A6E" bold="true" box="[199,510,1882,1908]" pageId="2" pageNumber="69">Geographical distribution</emphasis>
. 
<taxonomicName id="4C2AE2B97E21FFE3410BF8F641921A6F" authorityName="Viette" authorityYear="1949" box="[524,653,1883,1909]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B95E45287E21FFE3410BF8F641921A6F" box="[524,653,1883,1909]" italics="true" pageId="2" pageNumber="69">Schausiana</emphasis>
</taxonomicName>
is known to occur from 
<collectingCountry id="F33DD9AA7E21FFE340A7F8F640E91A6F" box="[928,1014,1883,1909]" name="Mexico" pageId="2" pageNumber="69">Mexico</collectingCountry>
to 
<collectingCountry id="F33DD9AA7E21FFE3471EF8F6478C1A6F" box="[1049,1171,1883,1909]" name="Costa Rica" pageId="2" pageNumber="69">Costa Rica</collectingCountry>
(
<figureCitation id="131185BF7E21FFE347A4F8F647E81A6F" box="[1187,1271,1883,1909]" captionStart="FIGURE 48" captionStartId="18.[151,250,1727,1751]" captionTargetBox="[227,1436,921,1704]" captionTargetId="figure-316@18.[151,1436,921,1704]" captionTargetPageId="18" captionText="FIGURE 48. Distribution of Schausiana spp. within southern North and Central America Eastern boundary of Guerrero terrane shown as blue dashed line." figureDoi="http://doi.org/10.5281/zenodo.4413530" httpUri="https://zenodo.org/record/4413530/files/figure.png" pageId="2" pageNumber="69">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E21FFE343C0F8D240401AA7" pageId="2" pageNumber="69" type="etymology">
<paragraph id="8B95993A7E21FFE343C0F8D240401AA7" blockId="2.[151,1437,802,2017]" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F8D242571A83" bold="true" box="[199,328,1919,1945]" pageId="2" pageNumber="69">Etymology</emphasis>
. Viette (1950) did not explicitly state the etymology for this genus, but he likely proposed it in honour of William Schaus, a prominent American lepidopterist.
</paragraph>
</subSubSection>
<subSubSection id="C330CAB17E21FFE243C0F86A41481C07" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69" type="discussion">
<paragraph id="8B95993A7E21FFE243C0F86A41481C07" blockId="2.[151,1437,802,2017]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
<emphasis id="B95E45287E21FFE343C0F86A422B1AFB" bold="true" box="[199,308,1991,2017]" pageId="2" pageNumber="69">Remarks</emphasis>
. Some other potential diagnostic characters have been found only where male and female genitalia have been dissected: (i) pseudotegumen dorso-mesally unfused while ventro-mesally fused, (ii) basal portion of the valve with spiny or hook process (both detected in 
<taxonomicName id="4C2AE2B97E20FFE241D5FF1640241DCE" authorityName="Schaus" authorityYear="1901" box="[722,827,187,213]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trojesa">
<emphasis id="B95E45287E20FFE241D5FF1641FF1DCF" box="[722,736,187,213]" italics="true" pageId="3" pageNumber="70">S</emphasis>
. 
<emphasis id="B95E45287E20FFE241E9FF1640241DCE" box="[750,827,187,212]" italics="true" pageId="3" pageNumber="70">trojesa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2AE2B97E20FFE2404EFF1640A31DCF" authority="Mielke &amp; Grehan &amp; Monzón-Sierra, 2020" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" box="[841,956,187,213]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="maishei" status="sp. nov.">
<emphasis id="B95E45287E20FFE2404EFF1640481DCF" box="[841,855,187,213]" italics="true" pageId="3" pageNumber="70">S</emphasis>
. 
<emphasis id="B95E45287E20FFE24062FF1640A31DCF" box="[869,956,187,213]" italics="true" pageId="3" pageNumber="70">maishei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E20FFE240C4FF16471E1DCF" box="[963,1025,187,213]" pageId="3" pageNumber="70" rank="species">
<emphasis id="B95E45287E20FFE240C4FF1640C21DCF" bold="true" box="[963,989,187,213]" pageId="3" pageNumber="70">sp</emphasis>
. 
<emphasis id="B95E45287E20FFE240ECFF1640E41DCF" bold="true" box="[1003,1019,187,213]" pageId="3" pageNumber="70">n</emphasis>
.
</taxonomicNameLabel>
, and 
<taxonomicName id="4C2AE2B97E20FFE24738FF1646521DCF" authority="(Druce, 1887)" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" baseAuthorityName="Druce" baseAuthorityYear="1887" box="[1087,1357,187,213]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="pharus" status="comb. nov.">
<emphasis id="B95E45287E20FFE24738FF1647521DCF" box="[1087,1101,187,213]" italics="true" pageId="3" pageNumber="70">S</emphasis>
. 
<emphasis id="B95E45287E20FFE2475CFF1647B61DCF" box="[1115,1193,187,213]" italics="true" pageId="3" pageNumber="70">pharus</emphasis>
(Druce, 1887)
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E20FFE24653FF1643B81DE3" pageId="3" pageNumber="70" rank="species">
<emphasis id="B95E45287E20FFE24653FF16468A1DCF" bold="true" box="[1364,1429,187,213]" pageId="3" pageNumber="70">comb</emphasis>
. 
<emphasis id="B95E45287E20FFE24390FF7243B81DE3" bold="true" box="[151,167,223,249]" pageId="3" pageNumber="70">n</emphasis>
</taxonomicNameLabel>
), and (iii) corpus bursae with diverticulum (detected in 
<taxonomicName id="4C2AE2B97E20FFE24027FF7240961DE2" authorityName="Schaus" authorityYear="1901" box="[800,905,223,249]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="trojesa">
<emphasis id="B95E45287E20FFE24027FF7240311DE3" box="[800,814,223,249]" italics="true" pageId="3" pageNumber="70">S</emphasis>
. 
<emphasis id="B95E45287E20FFE2403BFF7240961DE2" box="[828,905,223,248]" italics="true" pageId="3" pageNumber="70">trojesa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2AE2B97E20FFE240C6FF7247341DE3" authorityName="Mielke &amp; Grehan &amp; Monzón-Sierra" authorityYear="2020" baseAuthorityName="Druce" baseAuthorityYear="1887" box="[961,1067,223,249]" class="Insecta" family="Hepialidae" genus="Schausiana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="pharus" status="comb. nov.">
<emphasis id="B95E45287E20FFE240C6FF7240D01DE3" box="[961,975,223,249]" italics="true" pageId="3" pageNumber="70">S</emphasis>
. 
<emphasis id="B95E45287E20FFE240DAFF7247341DE3" box="[989,1067,223,249]" italics="true" pageId="3" pageNumber="70">pharus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26DF8537E20FFE24734FF7247871DE3" box="[1075,1176,223,249]" pageId="3" pageNumber="70" rank="species">
<emphasis id="B95E45287E20FFE24734FF72476B1DE3" bold="true" box="[1075,1140,223,249]" pageId="3" pageNumber="70">comb</emphasis>
. 
<emphasis id="B95E45287E20FFE24784FF72478C1DE3" bold="true" box="[1155,1171,223,249]" pageId="3" pageNumber="70">n</emphasis>
.
</taxonomicNameLabel>
). For comparison with other genera, see the discussion section.
</paragraph>
</subSubSection>
</treatment>
</document>