treatments-xml/data/BF/5D/B4/BF5DB417DA48FFA0FF38FAD527CCD980.xml
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<document id="3E7A8952977F6293AFD72E1EC73D8FAF" ID-DOI="10.11646/zootaxa.4809.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3934146" ID-ZooBank="7DD95C27-8BEF-467A-B24F-B43C7665B1A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594126143582" checkinUser="plazi" docAuthor="Ng, Peter K. L., Forges, Bertrand Richer De &amp; Martin, Joel W." docDate="2020" docId="BF5DB417DA48FFA0FF38FAD527CCD980" docLanguage="en" docName="zootaxa.4809.2.4.pdf" docOrigin="Zootaxa 4809 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homolidae De Haan 1839" docType="key" docVersion="5" lastPageNumber="326" masterDocId="4364CC6FDA5BFFB4FFAFFFA72250D94C" masterDocTitle="The Homolidae of the Hawaiian Islands, with notes on the taxonomy of Moloha major (Kubo, 1936) (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="328" masterPageNumber="306" pageNumber="325" updateTime="1698778880793" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7FF1D660345FE651F198874ABE0CD7A0">The Homolidae of the Hawaiian Islands, with notes on the taxonomy of Moloha major (Kubo, 1936) (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="90E84F1EA2AE3F32E669C21237B92EEA">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="50CB404033FAD2EC13491485471D2D67">Forges, Bertrand Richer De</mods:namePart>
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<mods:namePart id="5C9B4E4ED5DE2034C5DA9E60076C9637">Martin, Joel W.</mods:namePart>
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<mods:title id="CACE34923FBB050F514852ADA6D75BCE">Zootaxa</mods:title>
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<subSubSection id="7FEE568ADA48FFA7FF38FAD521E1DCC0" box="[151,945,1393,1420]" pageId="19" pageNumber="325" type="nomenclature">
<paragraph id="374B0501DA48FFA7FF38FAD521E1DCC0" blockId="19.[151,945,1393,1420]" box="[151,945,1393,1420]" pageId="19" pageNumber="325">
<heading id="6C03B26DDA48FFA7FF38FAD521E1DCC0" bold="true" box="[151,945,1393,1420]" fontSize="11" level="1" pageId="19" pageNumber="325" reason="1">
<emphasis id="0580D913DA48FFA7FF38FAD521E1DCC0" bold="true" box="[151,945,1393,1420]" pageId="19" pageNumber="325">
Key to the known species of
<taxonomicName id="F0F47E82DA48FFA7FE56FAD620D2DCC7" ID-CoL="7NHV8" authorityName="De Haan" authorityYear="1839" box="[505,642,1393,1419]" class="Malacostraca" family="Homolidae" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="family">Homolidae</taxonomicName>
in the Hawaiian Islands
</emphasis>
</heading>
</paragraph>
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<paragraph id="374B0501DA48FFA7FF38FA1127CCDC87" blockId="19.[151,1437,1461,2041]" box="[151,1436,1461,1484]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38FA1127CCDC87" box="[151,1436,1461,1484]" pageId="19" pageNumber="325">
1. Pereopod 5 merus clearly longer than carapace length (including rostrum)..........................
<taxonomicName id="F0F47E82DA48FFA7FB47FA1127CCDC87" ID-CoL="6P4HQ" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[1256,1436,1462,1483]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="okala">
<emphasis id="0580D913DA48FFA7FB47FA1127CCDC87" box="[1256,1436,1462,1483]" italics="true" pageId="19" pageNumber="325">Latreillopsis okala</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="374B0501DA48FFA7FF38FA7427CCDCA4" blockId="19.[151,1437,1461,2041]" box="[151,1436,1491,1513]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38FA7427CCDCA4" box="[151,1436,1491,1513]" pageId="19" pageNumber="325">- Pereopod 5 merus shorter than carapace length (including rostrum.............................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="8C001DA4DA48FFA7FF38FA5727CCDF11" pageId="19" pageNumber="325">
<paragraph id="374B0501DA48FFA7FF38FA5727CDDF6F" blockId="19.[151,1437,1461,2041]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38FA5727CDDF6F" pageId="19" pageNumber="325">
2. Rostrum with pair of dorsal spines, directed anterolaterally, forming trident; carapace with long lateral spine at base of cervical groove and long upright median gastric spine.............................................
<taxonomicName id="F0F47E82DA48FFA7FB15F9A927CDDF6F" ID-CoL="8WB8V" authorityName="Ng" authorityYear="2016" box="[1210,1437,1550,1571]" class="Malacostraca" family="Homolidae" genus="Homologenus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="namakae">
<emphasis id="0580D913DA48FFA7FB15F9A927CDDF6F" box="[1210,1437,1550,1571]" italics="true" pageId="19" pageNumber="325">Homologenus namakae</emphasis>
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<paragraph id="374B0501DA48FFA7FF38F98C27CCDF11" blockId="19.[151,1437,1461,2041]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F98C27CCDF11" pageId="19" pageNumber="325">- Rostrum simple or at most bifid, never tridendate; carapace margin at base of cervical groove unarmed or with spines of similar length to other carapace spines; without long median gastric spine.............................................. 3</keyLead>
</paragraph>
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<paragraph id="374B0501DA48FFA7FF38F9C127CCDF37" blockId="19.[151,1437,1461,2041]" box="[151,1436,1637,1660]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F9C127CCDF37" box="[151,1436,1637,1660]" pageId="19" pageNumber="325">
3. P5 merus long, reaching anteriorly to level of orbit when folded anteriorly..............................
<taxonomicName id="F0F47E82DA48FFA7FABDF9C127CCDF37" ID-CoL="84GZS" baseAuthorityName="Kubo" baseAuthorityYear="1936" box="[1298,1436,1638,1659]" class="Malacostraca" family="Homolidae" genus="Moloha" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="major">
<emphasis id="0580D913DA48FFA7FABDF9C127CCDF37" box="[1298,1436,1638,1659]" italics="true" pageId="19" pageNumber="325">Moloha major</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="374B0501DA48FFA7FF38F92427CCDFD4" blockId="19.[151,1437,1461,2041]" box="[151,1436,1667,1689]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F92427CCDFD4" box="[151,1436,1667,1689]" pageId="19" pageNumber="325">- P5 merus relatively short, not reaching anteriorly to the level of orbit when folded anteriorly......................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="8C001DA4DA48FFA7FF38F90727CCDF98" pageId="19" pageNumber="325">
<paragraph id="374B0501DA48FFA7FF38F90727CCDFFA" blockId="19.[151,1437,1461,2041]" box="[151,1436,1696,1718]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F90727CCDFFA" box="[151,1436,1696,1718]" pageId="19" pageNumber="325">4. Pseudorostral spines well developed, as long as or longer than rostrum........................................... 5</keyLead>
</paragraph>
<paragraph id="374B0501DA48FFA7FF38F91927CCDF98" blockId="19.[151,1437,1461,2041]" box="[151,1436,1726,1748]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F91927CCDF98" box="[151,1436,1726,1748]" pageId="19" pageNumber="325">- Pseudorostral spines small, conical, much shorter than rostrum................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="8C001DA4DA48FFA7FF38F97C27CCDE42" pageId="19" pageNumber="325">
<paragraph id="374B0501DA48FFA7FF38F97C27CCDFBC" blockId="19.[151,1437,1461,2041]" box="[151,1436,1755,1777]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F97C27CCDFBC" box="[151,1436,1755,1777]" pageId="19" pageNumber="325">
5. P5 merus very slender, length more than 10 times width....................................
<taxonomicName id="F0F47E82DA48FFA7FB00F97C27CCDFBC" ID-CoL="5CQW8" authorityName="Ng &amp; Eldredge" authorityYear="2012" box="[1199,1436,1755,1776]" class="Malacostraca" family="Homolidae" genus="Yaldwynopsis" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="hawaiiana">
<emphasis id="0580D913DA48FFA7FB00F97C27CCDFBC" box="[1199,1436,1755,1776]" italics="true" pageId="19" pageNumber="325">Yaldwynopsis hawaiiana</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="374B0501DA48FFA7FF38F95E27CCDE42" blockId="19.[151,1437,1461,2041]" box="[151,1436,1784,1806]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F95E27CCDE42" box="[151,1436,1784,1806]" pageId="19" pageNumber="325">
- P5 merus relatively stout, length about 7 times width...........................................
<taxonomicName id="F0F47E82DA48FFA7FB4DF95F27CCDE42" box="[1250,1436,1784,1806]" class="Malacostraca" family="Homolidae" genus="Paramola" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="0580D913DA48FFA7FB4DF95F27CCDE42" box="[1250,1436,1784,1806]" italics="true" pageId="19" pageNumber="325">Paramola japonica</emphasis>
</taxonomicName>
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</paragraph>
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<keyStep id="8C001DA4DA48FFA7FF38F8B127CCDECF" pageId="19" pageNumber="325">
<paragraph id="374B0501DA48FFA7FF38F8B127CCDE04" blockId="19.[151,1437,1461,2041]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F8B127CCDE04" pageId="19" pageNumber="325">6. Anterior wall of carapace either side of rostrum with elongated, transverse cavity forming orbit-like structure. Outer surface of cheliped palm with dark-coloured sunken depression at base of pollex in males and some females..................... 7</keyLead>
</paragraph>
<paragraph id="374B0501DA48FFA7FF38F8F627CCDECF" blockId="19.[151,1437,1461,2041]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F8F627CCDECF" pageId="19" pageNumber="325">- Anterior wall of carapace lateral to rostrum rounded, without orbit-like structure. Outer surface of cheliped palm never with dark-coloured sunken depression at base of pollex........................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="8C001DA4DA48FFA7FF38F82C27CCDEB5" pageId="19" pageNumber="325">
<paragraph id="374B0501DA48FFA7FF38F82C27CCDEF1" blockId="19.[151,1437,1461,2041]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F82C27CCDEF1" pageId="19" pageNumber="325">
7. Cheliped pollex straight, alignedt with rest of ventral margin of chela; outer surface of chela with scattered granules on median part, lateral parts more distinctly granulated; P2P4 long.........................................
<taxonomicName id="F0F47E82DA48FFA7FB5FF80F27CCDEF1" ID-CoL="3S2G8" baseAuthorityName="Takeda" baseAuthorityYear="1980" box="[1264,1436,1960,1981]" class="Malacostraca" family="Homolidae" genus="Lamoha" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="0580D913DA48FFA7FB5FF80F27CCDEF1" box="[1264,1436,1960,1981]" italics="true" pageId="19" pageNumber="325">Lamoha williamsi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="374B0501DA48FFA7FF38F86127CCDEB5" blockId="19.[151,1437,1461,2041]" pageId="19" pageNumber="325">
<keyLead id="8C05A634DA48FFA7FF38F86127CCDEB5" pageId="19" pageNumber="325">
- Cheliped pollex bent at an angle from ventral margin of chela; outer surface of chela covered with numerous prominent granules on entire surface; P2P4 proportionately shorter...........................................
<taxonomicName id="F0F47E82DA48FFA7FB47F84427CCDEB5" ID-CoL="3S2G6" baseAuthorityName="Guinot &amp; Richer de Forges" baseAuthorityYear="1981" box="[1256,1436,2019,2041]" class="Malacostraca" family="Homolidae" genus="Lamoha" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="325" phylum="Arthropoda" rank="species" species="personata">
<emphasis id="0580D913DA48FFA7FB47F84427CCDEB5" box="[1256,1436,2019,2041]" italics="true" pageId="19" pageNumber="325">Lamoha personata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="8C001DA4DA4FFFA0FF38FF3D27CCD980" pageId="20" pageNumber="326">
<paragraph id="374B0501DA4FFFA0FF38FF3D27CDD9E2" blockId="20.[151,1437,153,205]" box="[151,1437,153,176]" pageId="20" pageNumber="326">
<keyLead id="8C05A634DA4FFFA0FF38FF3D27CDD9E2" box="[151,1437,153,176]" pageId="20" pageNumber="326">
8. Only extremity of tip of rostrum slightly bifurcated, not easily visible..............................
<taxonomicName id="F0F47E82DA4FFFA0FB49FF3E27CDD9E2" ID-CoL="6MB8M" authorityName="Eldredge" authorityYear="1980" box="[1254,1437,153,174]" class="Malacostraca" family="Homolidae" genus="Homola" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="326" phylum="Arthropoda" rank="species" species="dickinsoni">
<emphasis id="0580D913DA4FFFA0FB49FF3E27CDD9E2" box="[1254,1437,153,174]" italics="true" pageId="20" pageNumber="326">Homola dickinsoni</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="374B0501DA4FFFA0FF38FF1027CCD980" blockId="20.[151,1437,153,205]" box="[151,1436,183,205]" pageId="20" pageNumber="326">
<keyLead id="8C05A634DA4FFFA0FF38FF1027CCD980" box="[151,1436,183,205]" pageId="20" pageNumber="326">
- Tip of rostrum clearly bifid even without magnification.........................................
<taxonomicName id="F0F47E82DA4FFFA0FB42FF1027CCD980" ID-CoL="6MB9R" authorityName="Henderson" authorityYear="1888" box="[1261,1436,183,204]" class="Malacostraca" family="Homolidae" genus="Homola" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="326" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="0580D913DA4FFFA0FB42FF1027CCD980" box="[1261,1436,183,204]" italics="true" pageId="20" pageNumber="326">Homola orientalis</emphasis>
</taxonomicName>
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