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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.171.2306" ID-PMC="PMC3288682" ID-Pensoft-Pub="1313-2970-171-39" ID-Pensoft-UUID="F3103053C45BFFED2A35FFF7FFA5FFA6" ID-PubMed="22423195" ID-Zenodo-Dep="576907" ID-ZooBank="1C0AE742AF194DCE8A6E86B6D72ECCB2" ModsDocID="1313-2970-171-39" checkinTime="1451249327470" checkinUser="pensoft" docAuthor="Kataev, Boris M., Liang, Hongbin &amp; Kavanaugh, David H." docDate="2012" docId="29B9E5C795DA533DAE33C4F3E055543C" docLanguage="en" docName="ZooKeys 171: 39-92" docOrigin="ZooKeys 171" docPubDate="2012-02-24" docSource="http://dx.doi.org/10.3897/zookeys.171.2306" docTitle="Chydaeus andrewesi subsp. kumei Ito 1992, stat. n." docType="treatment" docVersion="3" id="F3103053C45BFFED2A35FFF7FFA5FFA6" lastPageNumber="69" masterDocId="F3103053C45BFFED2A35FFF7FFA5FFA6" masterDocTitle="Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini)" masterLastPageNumber="92" masterPageNumber="39" pageNumber="67" updateTime="1668153268186" updateUser="ExternalLinkService">
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<mods:title>Contribution to knowledge of the genus Chydaeus in Xizang Autonomous Region [Tibet] and Yunnan Province, China (Coleoptera, Carabidae, Harpalini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kataev, Boris M.</mods:namePart>
<mods:affiliation>Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kavanaugh, David H.</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, California 94118 USA</mods:affiliation>
<mods:nameIdentifier type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-02-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>171</mods:number>
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<mods:start>39</mods:start>
<mods:end>92</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.171.2306</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.171.2306</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-171-39</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576907</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182236255" LSID="urn:lsid:plazi:treatment:29B9E5C795DA533DAE33C4F3E055543C" httpUri="http://treatment.plazi.org/id/29B9E5C795DA533DAE33C4F3E055543C" lastPageId="30" lastPageNumber="69" pageId="28" pageNumber="67">
<subSubSection pageId="28" pageNumber="67" type="nomenclature">
<paragraph pageId="28" pageNumber="67">
<taxonomicName LSID="29B9E5C7-95DA-533D-AE33-C4F3E055543C" authority="Ito, 1992" authorityName="Ito" authorityYear="1992" baseAuthorityName="Kataev &amp; Liang &amp; Kavanaugh" baseAuthorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Chydaeus andrewesi subsp. kumei" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="subSpecies" species="andrewesi" status="stat. n." subSpecies="kumei">
<pageBreakToken pageId="28" pageNumber="67" start="start">Chydaeus</pageBreakToken>
andrewesi kumei Ito, 1992
</taxonomicName>
<taxonomicNameLabel pageId="28" pageNumber="67">stat. n.</taxonomicNameLabel>
<figureCitation captionStart="Figures 4750" captionStartId="F8" captionText="Figures 47 - 50. Chydaeus andrewesi kumei Ito (Vietnam, Sa Pa area) 47 Terminal lamella of median lobe, dorsal view 48 Median lobe, left lateral view 49 Median lobe, dorsal view. 50 Median lobe, right lateral view. Scale lines: A = 0.5 mm (Fig. 47), B = 1.0 mm (Figs 48 - 50)." httpUri="https://binary.pensoft.net/fig/14940" pageId="28" pageNumber="67">Figs 47-50</figureCitation>
<figureCitation captionStart="Figures 6061" captionStartId="F12" captionText="Figures 60 - 61. Toporelief map of southeastern Asia, illustrating localities for Chydaeus species 60 Chydaeus andrewsi Schauberger: Chydaeus andrewsi andrewsi Schauberger = red dots and Chydaeus andrewsi kumei Ito = green squares 61 Chydaeus salvazae Schauberger = red dots. Only new records reported here are shown. Scale bar = 300 km (in 100 km increments)." httpUri="https://binary.pensoft.net/fig/14948" pageId="28" pageNumber="67">60</figureCitation>
<figureCitation captionStart="Figure 74" captionStartId="F19" captionText="Figure 74. Digital photograph of landscape at Fenshuiling, Jinping County, Yunnan Province, China, ca. 2060 m; locality for Chydaeus andrewsi kumei Ito." httpUri="https://binary.pensoft.net/fig/14959" pageId="28" pageNumber="67">74</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="67" type="reference_group">
<paragraph pageId="28" pageNumber="67">
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" baseAuthorityName="Ito" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="kumei" status="stat. n.">Chydaeus kumei</taxonomicName>
Ito, 1992: 52
</paragraph>
<paragraph pageId="28" pageNumber="67">
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="guangxiensis" status="stat. n." subGenus="Chydaeus">? Chydaeus (Chydaeus) guangxiensis</taxonomicName>
Ito, 2006: 198
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="67" type="material examined">
<paragraph pageId="28" pageNumber="67">Material examined.</paragraph>
<paragraph pageId="28" pageNumber="67">
A total of 35 specimens (25 males and 10 females, including 1 male in IOZ and 24 males and 10 females in cFED, cSCH, cWR, and ZIN) were examined from the following localities:
<emphasis bold="true" pageId="28" pageNumber="67">China</emphasis>
.
<emphasis bold="true" pageId="28" pageNumber="67">Yunnan Province</emphasis>
.
<emphasis italics="true" pageId="28" pageNumber="67">Jinping County</emphasis>
: 1 male, Fenshuiling, Leidazhan, roadside,
<geoCoordinate degrees="22" direction="north" minutes="51" orientation="latitude" precision="1" seconds="41.0" value="22.86139">22°51'41.0&quot;N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="13" orientation="longitude" precision="1" seconds="40.5" value="103.22792">103°13'40.5&quot;E</geoCoordinate>
, 2060 m, 17.XII.2003, H.B. Liang leg. (IOZ).
<emphasis bold="true" pageId="28" pageNumber="67">Vietnam</emphasis>
.
<emphasis bold="true" pageId="28" pageNumber="67">Lao Cai Province</emphasis>
: 2 males, 2 females, 6 km W of Sa Pa, N slope of Phansipan Mt. Area, 2000-2100 m, near Tram don (base of Hoang Lien Nature Park),
<geoCoordinate degrees="22" direction="north" minutes="21" orientation="latitude" precision="925" value="22.35">22°21'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="46" orientation="longitude" precision="925" value="103.76667">103°46'E</geoCoordinate>
, V.2005, A.V. Abramov leg. (Exp. of Russia-Vietnam Tropical Centre) (ZIN); 1 female, same data, but 1930-2000 m, V.2010, A.V. Abramov leg. (ZIN); 18 males, 6 females, 6 km W of Sa Pa, northern slope of Phansipan Mt. area, May 2008, A.V. Abramov leg. (Exp. of Russia-Vietnam Tropical Centre) (cSCH, cWR, ZIN); 4 males, 1 female, Sa Pa env., ca 1600 m, V.2006, A. Anitchkin leg. (cFED).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="67" type="distribution">
<paragraph pageId="28" pageNumber="67">Distribution.</paragraph>
<paragraph pageId="28" pageNumber="67">
<figureCitation captionStart="Figures 6061" captionStartId="F12" captionText="Figures 60 - 61. Toporelief map of southeastern Asia, illustrating localities for Chydaeus species 60 Chydaeus andrewsi Schauberger: Chydaeus andrewsi andrewsi Schauberger = red dots and Chydaeus andrewsi kumei Ito = green squares 61 Chydaeus salvazae Schauberger = red dots. Only new records reported here are shown. Scale bar = 300 km (in 100 km increments)." httpUri="https://binary.pensoft.net/fig/14948" pageId="28" pageNumber="67">Fig. 60</figureCitation>
. Previously, this taxon was known only from the single male collected in Sa Pa, northern Vietnam. Based on the new records presented here, this species occurs not only in the mountains of northern Vietnam, but also in southern Yunnan Province (China) adjacent to the Vietnamese border.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="67" type="habitat">
<paragraph pageId="28" pageNumber="67">Habitat.</paragraph>
<paragraph pageId="28" pageNumber="67">
Specimens were collected in roadside and road cut open areas and other disturbed areas adjacent to moderately disturbed forest (
<figureCitation captionStart="Figure 74" captionStartId="F19" captionText="Figure 74. Digital photograph of landscape at Fenshuiling, Jinping County, Yunnan Province, China, ca. 2060 m; locality for Chydaeus andrewsi kumei Ito." httpUri="https://binary.pensoft.net/fig/14959" pageId="28" pageNumber="67">Fig. 74</figureCitation>
), hidden under stones and other debris during daylight hours.
</paragraph>
</subSubSection>
<subSubSection lastPageId="30" lastPageNumber="69" pageId="28" pageNumber="67" type="remarks">
<paragraph pageId="28" pageNumber="67">Remarks.</paragraph>
<paragraph pageId="28" pageNumber="67">
Based on
<normalizedToken originalValue="Itos">Ito's</normalizedToken>
original description,
<bibRefCitation author="Kataev, BM" editor="Hartmann, M" journalOrPublisher="II. Verein der Freunde und Foderer des Naturkundemuseums Erfurt e. V., Erfurt" pageId="37" pageNumber="76" pagination="133 - 174" refId="B11" refString="Kataev, BM, Schmidt, J, 2006. To the knowledge of the genus Chydaeus Chaudoir, 1854 (Coleoptera, Carabidae, Harpalini). II. Revision of the C. obscurus, C. semenowi (in new sense), and C. salvazae species-groups of the Himalaya and adjacent areas. In: Hartmann, M, Weipert, J, Eds., Biodiversitat und Naturausstattung im Himalaya - Biodiversity and natural heritage of the Himalaya, Bd. II.Verein der Freunde und Foderer des Naturkundemuseums Erfurt e. V., Erfurt: 133 - 174" title="To the knowledge of the genus Chydaeus Chaudoir, 1854 (Coleoptera, Carabidae, Harpalini). II. Revision of the C. obscurus, C. semenowi (in new sense), and C. salvazae species-groups of the Himalaya and adjacent areas." volumeTitle="Biodiversitat und Naturausstattung im Himalaya - Biodiversity and natural heritage of the Himalaya, Bd." year="2006">Kataev and Schmidt (2006)</bibRefCitation>
treated
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" baseAuthorityName="Ito" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="kumei">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus kumei</emphasis>
</taxonomicName>
as a distinct species of the
<emphasis italics="true" pageId="28" pageNumber="67">obscurus</emphasis>
group, closely related to
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus andrewesi</emphasis>
</taxonomicName>
as characterized by
<bibRefCitation author="Ito, N" journalOrPublisher="Entomological Review of Japan" pageId="36" pageNumber="75" pagination="43 - 57" refId="B4" refString="Ito, N, 1992. Study on Asian Carabidae, IV. (Coleoptera). Entomological Review of Japan 47: 43 - 57" title="Study on Asian Carabidae, IV. (Coleoptera)." volume="47" year="1992">Ito (1992)</bibRefCitation>
. Examination of specimens of
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" baseAuthorityName="Ito" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="kumei">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus kumei</emphasis>
</taxonomicName>
revealed that this taxon possesses all the distinctive features of
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus andrewesi</emphasis>
</taxonomicName>
listed by
<bibRefCitation author="Kataev, BM" editor="Hartmann, M" journalOrPublisher="II. Verein der Freunde und Foderer des Naturkundemuseums Erfurt e. V., Erfurt" pageId="37" pageNumber="76" pagination="133 - 174" refId="B11" refString="Kataev, BM, Schmidt, J, 2006. To the knowledge of the genus Chydaeus Chaudoir, 1854 (Coleoptera, Carabidae, Harpalini). II. Revision of the C. obscurus, C. semenowi (in new sense), and C. salvazae species-groups of the Himalaya and adjacent areas. In: Hartmann, M, Weipert, J, Eds., Biodiversitat und Naturausstattung im Himalaya - Biodiversity and natural heritage of the Himalaya, Bd. II.Verein der Freunde und Foderer des Naturkundemuseums Erfurt e. V., Erfurt: 133 - 174" title="To the knowledge of the genus Chydaeus Chaudoir, 1854 (Coleoptera, Carabidae, Harpalini). II. Revision of the C. obscurus, C. semenowi (in new sense), and C. salvazae species-groups of the Himalaya and adjacent areas." volumeTitle="Biodiversitat und Naturausstattung im Himalaya - Biodiversity and natural heritage of the Himalaya, Bd." year="2006">Kataev and Schmidt (2006)</bibRefCitation>
, including the elongate metepisterna (in both taxa, their length along inner margin is much greater than the width along the anterior margin). Moreover, the structure of the aedeagus in
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" baseAuthorityName="Ito" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="kumei">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus kumei</emphasis>
</taxonomicName>
males (
<figureCitation captionStart="Figures 4750" captionStartId="F8" captionText="Figures 47 - 50. Chydaeus andrewesi kumei Ito (Vietnam, Sa Pa area) 47 Terminal lamella of median lobe, dorsal view 48 Median lobe, left lateral view 49 Median lobe, dorsal view. 50 Median lobe, right lateral view. Scale lines: A = 0.5 mm (Fig. 47), B = 1.0 mm (Figs 48 - 50)." httpUri="https://binary.pensoft.net/fig/14940" pageId="28" pageNumber="67">Figs 47-50</figureCitation>
) is virtually identical to that in
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus andrewesi</emphasis>
</taxonomicName>
males and, in our opinion,
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" baseAuthorityName="Ito" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="kumei">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus kumei</emphasis>
</taxonomicName>
should be treated as a subspecies of
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus andrewesi</emphasis>
</taxonomicName>
. The vicariant distributions of these taxa in Yunnan also supports the subspecific status of
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" baseAuthorityName="Ito" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="kumei">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus kumei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph lastPageId="29" lastPageNumber="68" pageId="28" pageNumber="67">
The main distinctive characters of adults of
<taxonomicName authorityName="Ito" authorityYear="1992" baseAuthorityName="Kataev &amp; Liang &amp; Kavanaugh" baseAuthorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="28" pageNumber="67" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="kumei">
<emphasis italics="true" pageId="28" pageNumber="67">Chydaeus andrewesi kumei</emphasis>
</taxonomicName>
are: pronotum markedly narrowed basad, distinctly depressed and comparatively coarsely punctate laterobasally; pronotal basal angles each with a denticulate apex protruded laterad; elytral disc in males with very fine, more or less markedly effaced microsculpture comprised of thin, transverse meshes; proepisterna distinctly punctate anteriorly; and base of the terminal lamella of the median lobe of aedeagus (
<figureCitation captionStart="Figures 4750" captionStartId="F8" captionText="Figures 47 - 50. Chydaeus andrewesi kumei Ito (Vietnam, Sa Pa area) 47 Terminal lamella of median lobe, dorsal view 48 Median lobe, left lateral view 49 Median lobe, dorsal view. 50 Median lobe, right lateral view. Scale lines: A = 0.5 mm (Fig. 47), B = 1.0 mm (Figs 48 - 50)." httpUri="https://binary.pensoft.net/fig/14940" pageId="28" pageNumber="67">Fig. 47</figureCitation>
) slightly wider than that of other subspecies. We add the following mensural features to
<normalizedToken originalValue="Itos">Ito's</normalizedToken>
description:
<pageBreakToken pageId="29" pageNumber="68" start="start">Size</pageBreakToken>
: Body length 8.6-10.2 mm, width 3.8-4.5 mm. Proportions: HWmax/PWmax = 0.71-0.74; HWmin/PWmax = 0.59-0.63; PWmax/PL = 1.41-1.48; PWmax/ PWmin = 1.26-1.40; EL/EW = 1.38-1.43, EL/PL = 2.39-2.59 (2.39-2.51 in male and 2.42-2.59 in female), W/PWmax = 1.19-1.29.
</paragraph>
<paragraph pageId="29" pageNumber="68">
Specimens of
<taxonomicName authorityName="Ito" authorityYear="1992" baseAuthorityName="Kataev &amp; Liang &amp; Kavanaugh" baseAuthorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="kumei">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus andrewesi kumei</emphasis>
</taxonomicName>
are very similar to those of
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="szetschuanus">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus andrewesi szetschuanus</emphasis>
</taxonomicName>
from Sichuan in morphological features, particularly in the shape of the pronotum (with denticulate basal angles), the punctate proepisterna, and the relatively wide elytra; but they differ from the latter mainly in having finer and sparser punctation on the apical half of the pronotum (pronotum densely and coarsely punctate anteriorly in
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="szetschuanus">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus andrewesi szetschuanus</emphasis>
</taxonomicName>
specimens).
</paragraph>
<paragraph lastPageId="30" lastPageNumber="69" pageId="29" pageNumber="68">
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="species" species="guangxiensis">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus guangxiensis</emphasis>
</taxonomicName>
Ito, 2006 was described from one male from Guangxi (&quot;Below Tienshan Ping, Mt. Miao'er, Xing'an Xian&quot;). We examined two females from the same mountain [one labeled: &quot;Guangxi, Mt.
<normalizedToken originalValue="Miaoer">Miao'er</normalizedToken>
, 2000m, 1985.VIII.1, Song Shimei leg. (IOZ); for the other, see
<bibRefCitation author="Kataev, BM" editor="Hartmann, M" journalOrPublisher="II. Verein der Freunde und Foderer des Naturkundemuseums Erfurt e. V., Erfurt" pageId="37" pageNumber="76" pagination="133 - 174" refId="B11" refString="Kataev, BM, Schmidt, J, 2006. To the knowledge of the genus Chydaeus Chaudoir, 1854 (Coleoptera, Carabidae, Harpalini). II. Revision of the C. obscurus, C. semenowi (in new sense), and C. salvazae species-groups of the Himalaya and adjacent areas. In: Hartmann, M, Weipert, J, Eds., Biodiversitat und Naturausstattung im Himalaya - Biodiversity and natural heritage of the Himalaya, Bd. II.Verein der Freunde und Foderer des Naturkundemuseums Erfurt e. V., Erfurt: 133 - 174" title="To the knowledge of the genus Chydaeus Chaudoir, 1854 (Coleoptera, Carabidae, Harpalini). II. Revision of the C. obscurus, C. semenowi (in new sense), and C. salvazae species-groups of the Himalaya and adjacent areas." volumeTitle="Biodiversitat und Naturausstattung im Himalaya - Biodiversity and natural heritage of the Himalaya, Bd." year="2006">Kataev and Schmidt (2006</bibRefCitation>
: 145)], that matched the original description of
<taxonomicName authorityName="Kataev &amp; Liang &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="species" species="guangxiensis">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus guangxiensis</emphasis>
</taxonomicName>
very well. Judging from Ito's original drawings (2006: Fig. 4), the aedeagus of
<taxonomicName class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="species" species="guangxien">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus guangxien</emphasis>
</taxonomicName>
sis males is identical to that of
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="29" pageNumber="68" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis italics="true" pageId="29" pageNumber="68">Chydaeus andrewesi</emphasis>
</taxonomicName>
<pageBreakToken pageId="30" pageNumber="69" start="start">males</pageBreakToken>
. The pronotum of
<taxonomicName class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="guangxien">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus guangxien</emphasis>
</taxonomicName>
sis adults is smooth or finely punctate anteriorly, as in
<taxonomicName authorityName="Ito" authorityYear="1992" baseAuthorityName="Kataev &amp; Liang &amp; Kavanaugh" baseAuthorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="kumei">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewesi kumei</emphasis>
</taxonomicName>
adults, but the denticle at each basal pronotal angle is only slightly evident or absent, as in
<taxonomicName authority="andrewesi" authorityName="andrewesi Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="andrewesi">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewesi andrewesi</emphasis>
</taxonomicName>
specimens. Further study, based on additional specimens, is necessary to determine the taxonomic status of
<taxonomicName class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="guangxien">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus guangxien</emphasis>
</taxonomicName>
sis; but it is very likely that this taxon is either consubspecific with
<taxonomicName authorityName="Ito" authorityYear="1992" baseAuthorityName="Kataev &amp; Liang &amp; Kavanaugh" baseAuthorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="kumei">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewesi kumei</emphasis>
</taxonomicName>
or, at most, another subspecies of
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewesi</emphasis>
</taxonomicName>
.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/14940" pageId="30" pageNumber="69" start="Figures 4750" startId="F8">
<paragraph pageId="30" pageNumber="69">
<emphasis bold="true" pageId="30" pageNumber="69">Figures 47-50.</emphasis>
<taxonomicName authorityName="Ito" authorityYear="1992" baseAuthorityName="Kataev &amp; Liang &amp; Kavanaugh" baseAuthorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="andrewesi" subSpecies="kumei">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewesi kumei</emphasis>
</taxonomicName>
Ito (Vietnam, Sa Pa area)
<emphasis bold="true" pageId="30" pageNumber="69">47</emphasis>
Terminal lamella of median lobe, dorsal view
<emphasis bold="true" pageId="30" pageNumber="69">48</emphasis>
Median lobe, left lateral view
<emphasis bold="true" pageId="30" pageNumber="69">49</emphasis>
Median lobe, dorsal view.
<emphasis bold="true" pageId="30" pageNumber="69">50</emphasis>
Median lobe, right lateral view. Scale lines:
<emphasis bold="true" pageId="30" pageNumber="69">A</emphasis>
= 0.5 mm (Fig. 47),
<emphasis bold="true" pageId="30" pageNumber="69">B</emphasis>
= 1.0 mm (Figs 48-50).
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/14942" pageId="30" pageNumber="69" start="Figures 5155" startId="F9">
<paragraph pageId="30" pageNumber="69">
<emphasis bold="true" pageId="30" pageNumber="69">Figures 51-55.</emphasis>
Dorsal habitus,
<taxonomicName authorityName="de Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus</emphasis>
</taxonomicName>
species.
<emphasis bold="true" pageId="30" pageNumber="69">51</emphasis>
<taxonomicName authorityName="Kataev &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="fugongensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus fugongensis</emphasis>
</taxonomicName>
sp. n. (holotype).
<emphasis bold="true" pageId="30" pageNumber="69">52</emphasis>
<taxonomicName authorityName="Kataev &amp; Liang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="gutangensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus gutangensis</emphasis>
</taxonomicName>
sp. n. (holotype).
<emphasis bold="true" pageId="30" pageNumber="69">53</emphasis>
<taxonomicName authorityName="Kataev &amp; Liang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="hanmiensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus hanmiensis</emphasis>
</taxonomicName>
sp. n. (holotype).
<emphasis bold="true" pageId="30" pageNumber="69">54</emphasis>
<taxonomicName authorityName="Kataev &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="asetosus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus asetosus</emphasis>
</taxonomicName>
sp. n. (holotype)
<emphasis bold="true" pageId="30" pageNumber="69">55</emphasis>
<taxonomicName authorityName="Kataev &amp; Liang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="baoshanensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus baoshanensis</emphasis>
</taxonomicName>
sp. n. (holotype). Scale lines = 1.0 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/14943" pageId="30" pageNumber="69" start="Figures 5657" startId="F10">
<paragraph pageId="30" pageNumber="69">
<emphasis bold="true" pageId="30" pageNumber="69">Figures 56-57.</emphasis>
Toporelief map of southeastern Asia, illustrating localities for
<taxonomicName authorityName="de Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus</emphasis>
</taxonomicName>
species
<emphasis bold="true" pageId="30" pageNumber="69">56</emphasis>
<taxonomicName authorityName="Ito" authorityYear="2006" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="shunichii">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus shunichii</emphasis>
</taxonomicName>
Ito = red dots,
<taxonomicName authorityName="Kataev &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="fugongensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus fugongensis</emphasis>
</taxonomicName>
sp. n. = pink diamond,
<taxonomicName authorityName="Kataev &amp; Liang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="gutangensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus gutangensis</emphasis>
</taxonomicName>
sp. n. = yellow square, and
<taxonomicName authorityName="Kataev &amp; Liang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="hanmiensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus hanmiensis</emphasis>
</taxonomicName>
sp. n. = light blue triangle
<emphasis bold="true" pageId="30" pageNumber="69">57</emphasis>
<taxonomicName authorityName="Kataev &amp; Liang" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="baoshanensis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus baoshanensis</emphasis>
</taxonomicName>
sp. n. = dark blue square,
<taxonomicName authorityName="Kataev &amp; Kavanaugh" authorityYear="2012" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="asetosus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus asetosus</emphasis>
</taxonomicName>
sp. n. = yellow triangles, and
<taxonomicName authorityName="Schauberger" authorityYear="1932" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="obtusicollis">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus obtusicollis</emphasis>
</taxonomicName>
Schauberger = red dots. Only new records reported here are shown. Scale bar = 300 km (in 100 km increments).
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/14946" pageId="30" pageNumber="69" start="Figures 5859" startId="F11">
<paragraph pageId="30" pageNumber="69">
<emphasis bold="true" pageId="30" pageNumber="69">Figures 58-59.</emphasis>
Toporelief map of southeastern Asia, illustrating localities for
<taxonomicName authorityName="de Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus</emphasis>
</taxonomicName>
species
<emphasis bold="true" pageId="30" pageNumber="69">58</emphasis>
<taxonomicName authorityName="Ito" authorityYear="2003" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="satoi">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus satoi</emphasis>
</taxonomicName>
Ito = yellow square and
<taxonomicName authorityName="Ito" authorityYear="2002" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="convexus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus convexus</emphasis>
</taxonomicName>
Ito = red dots
<emphasis bold="true" pageId="30" pageNumber="69">59</emphasis>
<taxonomicName baseAuthorityName="Tschitscherine" baseAuthorityYear="1899" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="semenowi">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus semenowi</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Tschitschérine">Tschitscherine</normalizedToken>
) = red dots and
<taxonomicName authorityName="Kataev &amp; Schmidt" authorityYear="2006" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus malaisei</emphasis>
</taxonomicName>
Kataev &amp; Schmidt = yellow squares. Only new records reported here are shown. Scale bar = 300 km (in 100 km increments).
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/14948" pageId="30" pageNumber="69" start="Figures 6061" startId="F12">
<paragraph pageId="30" pageNumber="69">
<emphasis bold="true" pageId="30" pageNumber="69">Figures 60-61.</emphasis>
Toporelief map of southeastern Asia, illustrating localities for
<taxonomicName authorityName="de Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus</emphasis>
</taxonomicName>
species
<emphasis bold="true" pageId="30" pageNumber="69">60</emphasis>
<taxonomicName class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="andrewsi">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewsi</emphasis>
</taxonomicName>
Schauberger:
<taxonomicName authority="andrewsi" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="andrewsi" subSpecies="andrewsi">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewsi andrewsi</emphasis>
</taxonomicName>
Schauberger = red dots and
<taxonomicName class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="andrewsi" subSpecies="kumei">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus andrewsi kumei</emphasis>
</taxonomicName>
Ito = green squares
<emphasis bold="true" pageId="30" pageNumber="69">61</emphasis>
<taxonomicName authority="Schauberger" authorityName="Schauberger" authorityYear="1934" class="Insecta" family="Carabidae" genus="Chydaeus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="30" pageNumber="69" phylum="Arthropoda" rank="species" species="salvazae">
<emphasis italics="true" pageId="30" pageNumber="69">Chydaeus salvazae Schauberger</emphasis>
</taxonomicName>
= red dots. Only new records reported here are shown. Scale bar = 300 km (in 100 km increments).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>