1027 lines
196 KiB
XML
1027 lines
196 KiB
XML
<document id="C89446616AE3A2A84E2FA58DE8A2BA76" ID-CLB-Dataset="8760" ID-DOI="10.11646/zootaxa.4822.2.5" ID-GBIF-Dataset="c45ae0a1-a4c5-4cd5-93c5-abdbb5a2a727" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401461" ID-ZooBank="6FC281F4-CC48-41E4-9C73-80B57A2A2D17" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609336114936" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Kawai, Kei & Ng, Peter K. L." docDate="2020" docId="131587D99505427A73A2FF50FAA3C008" docLanguage="en" docName="zootaxa.4822.2.5.pdf" docOrigin="Zootaxa 4822 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcotheres ocularius Komai & Kawai & Ng 2020, n. sp." docType="treatment" docUuid="85D9E4FC-CF7C-43B7-B21C-7070705CEE69" docUuidSource="ZooBank" docVersion="8" lastPageNumber="244" masterDocId="EF2CFFA1950E426D7335FFC7FFD4C269" masterDocTitle="On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the Southwest Pacific (Decapoda: Brachyura: Pinnotheridae)" masterLastPageNumber="247" masterPageNumber="221" pageNumber="232" updateTime="1701998130601" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="CD9FC7802D721587D02F30EEAA8AC21E">On the identity of the poorly known pea crab, Pinnothera obesa Dana, 1852, and description of a new species of Arcotheres Manning, 1993 from the Southwest Pacific (Decapoda: Brachyura: Pinnotheridae)</mods:title>
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<mods:name id="1096DF6426DADEB275431EA346A5C4B5" type="personal">
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<mods:namePart id="E2238A3E82E5DC64ED07C5FE20EF4C2F">Komai, Tomoyuki</mods:namePart>
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<mods:affiliation id="EC6AE418CFB0E545B9E67B4763AE1344">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, 260 - 8682 Japan.</mods:affiliation>
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<mods:namePart id="575D630FE780FAEDE5963254953B7CA5">Kawai, Kei</mods:namePart>
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<mods:affiliation id="016379DAC9B8F30AC5C0B8BE4AEEEB7B">International Center for Island Studies, Kagoshima University, Korimoto 1 - 21 - 24, Kagoshima, 890 - 8580.</mods:affiliation>
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<mods:namePart id="C42F020BFED855863FEBB5D91D0F61C7">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="B7E66E30E6D6C1225BA663324DEFED6D">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:nameIdentifier id="57FC8B8DBF066D5BF5E85BDF159693B8" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:title id="C6D4F08BED4AD4B2F29635422A7C47D1">Zootaxa</mods:title>
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<mods:date id="426C088D3027B2A0096807E3B13AE20B">2020</mods:date>
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<mods:number id="D69D23211634AD44691EF57E63E4BCCC">2020-08-05</mods:number>
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<mods:number id="85F118055DA979E21987479EDA569206">4822</mods:number>
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<treatment id="131587D99505427A73A2FF50FAA3C008" ID-DOI="http://doi.org/10.5281/zenodo.4401459" ID-GBIF-Taxon="177010450" ID-Zenodo-Dep="4401459" LSID="urn:lsid:zoobank.org:act:85D9E4FC-CF7C-43B7-B21C-7070705CEE69" httpUri="http://treatment.plazi.org/id/131587D99505427A73A2FF50FAA3C008" lastPageId="23" lastPageNumber="244" pageId="11" pageNumber="232">
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<subSubSection id="D3A665449505426673A2FF50FE0AC2D8" box="[151,478,151,178]" pageId="11" pageNumber="232" type="nomenclature">
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<paragraph id="9B0336CF9505426673A2FF50FE0AC2D8" blockId="11.[151,478,151,213]" box="[151,478,151,178]" pageId="11" pageNumber="232">
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<heading id="C04B81A39505426673A2FF50FE0AC2D8" bold="true" box="[151,478,151,178]" fontSize="11" level="1" pageId="11" pageNumber="232" reason="1">
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<emphasis id="A9C8EADD9505426673A2FF50FE0AC2D8" bold="true" box="[151,478,151,178]" pageId="11" pageNumber="232">
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<taxonomicName id="5CBC4D4C9505426673A2FF50FE41C2DB" ID-CoL="84GPY" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[151,405,151,178]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="232" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
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<emphasis id="A9C8EADD9505426673A2FF50FE41C2DB" bold="true" box="[151,405,151,178]" italics="true" pageId="11" pageNumber="232">Arcotheres ocularius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B2FB57A69505426672A9FF50FE0AC2D8" box="[412,478,151,177]" pageId="11" pageNumber="232" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A665449505426673A2FF7CFEF4C2BC" box="[151,288,187,213]" pageId="11" pageNumber="232" type="description">
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<paragraph id="9B0336CF9505426673A2FF7CFEF4C2BC" blockId="11.[151,478,151,213]" box="[151,288,187,213]" pageId="11" pageNumber="232">
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(
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<figureCitation id="03872A4A9505426673AAFF7CFECCC2BC" box="[159,280,187,213]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-10="FIGURE 17" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStart-5="FIGURE 12" captionStart-6="FIGURE 13" captionStart-7="FIGURE 14" captionStart-8="FIGURE 15" captionStart-9="FIGURE 16" captionStartId-0="13.[151,250,1943,1967]" captionStartId-1="14.[151,250,1828,1852]" captionStartId-10="23.[151,250,1620,1644]" captionStartId-2="15.[151,250,1866,1890]" captionStartId-3="16.[151,250,1553,1577]" captionStartId-4="17.[151,250,1157,1181]" captionStartId-5="18.[151,250,2007,2031]" captionStartId-6="19.[151,250,1850,1874]" captionStartId-7="20.[151,250,1151,1175]" captionStartId-8="21.[151,250,1870,1894]" captionStartId-9="22.[151,250,1901,1925]" captionTargetBox-0="[163,1425,181,1919]" captionTargetBox-1="[161,1426,190,1794]" captionTargetBox-10="[151,1435,633,1597]" captionTargetBox-2="[206,1379,190,1831]" captionTargetBox-3="[160,1425,190,1519]" captionTargetBox-4="[160,1425,189,1127]" captionTargetBox-5="[159,1427,1065,1983]" captionTargetBox-6="[161,1425,191,1815]" captionTargetBox-7="[157,1428,188,1117]" captionTargetBox-8="[175,1414,188,1838]" captionTargetBox-9="[180,1407,855,1867]" captionTargetId-0="figure-17@13.[163,1425,181,1919]" captionTargetId-1="figure-20@14.[151,1436,181,1805]" captionTargetId-10="figure-260@23.[151,1435,633,1597]" captionTargetId-2="figure-17@15.[197,1390,181,1842]" captionTargetId-3="figure-20@16.[151,1436,181,1529]" captionTargetId-4="figure-17@17.[151,1436,181,1133]" captionTargetId-5="figure-437@18.[159,1427,1065,1983]" captionTargetId-6="figure-17@19.[151,1436,181,1826]" captionTargetId-7="figure-20@20.[151,1436,181,1127]" captionTargetId-8="figure-17@21.[166,1422,181,1847]" captionTargetId-9="figure-370@22.[173,1414,849,1878]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-10="23" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionTargetPageId-5="18" captionTargetPageId-6="19" captionTargetPageId-7="20" captionTargetPageId-8="21" captionTargetPageId-9="22" captionText-0="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." captionText-1="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." captionText-10="FIGURE 17. Arcotheres ocularius n. sp., female of post-hard stage V, not specified, living animal associated with Anadara sp." captionText-2="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." captionText-3="FIGURE 10. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, habitus, dorsal view; B, thoracic sternite 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, pleon and telson, ventral view." captionText-4="FIGURE 11. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus." captionText-5="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." captionText-6="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." captionText-7="FIGURE 14. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view." captionText-8="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." captionText-9="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.4401477" figureDoi-1="http://doi.org/10.5281/zenodo.4401479" figureDoi-10="http://doi.org/10.5281/zenodo.4401499" figureDoi-2="http://doi.org/10.5281/zenodo.4401481" figureDoi-3="http://doi.org/10.5281/zenodo.4401483" figureDoi-4="http://doi.org/10.5281/zenodo.4401485" figureDoi-5="http://doi.org/10.5281/zenodo.4401487" figureDoi-6="http://doi.org/10.5281/zenodo.4401489" figureDoi-7="http://doi.org/10.5281/zenodo.4401491" figureDoi-8="http://doi.org/10.5281/zenodo.4401493" figureDoi-9="http://doi.org/10.5281/zenodo.4401497" httpUri-0="https://zenodo.org/record/4401477/files/figure.png" httpUri-1="https://zenodo.org/record/4401479/files/figure.png" httpUri-10="https://zenodo.org/record/4401499/files/figure.png" httpUri-2="https://zenodo.org/record/4401481/files/figure.png" httpUri-3="https://zenodo.org/record/4401483/files/figure.png" httpUri-4="https://zenodo.org/record/4401485/files/figure.png" httpUri-5="https://zenodo.org/record/4401487/files/figure.png" httpUri-6="https://zenodo.org/record/4401489/files/figure.png" httpUri-7="https://zenodo.org/record/4401491/files/figure.png" httpUri-8="https://zenodo.org/record/4401493/files/figure.png" httpUri-9="https://zenodo.org/record/4401497/files/figure.png" pageId="11" pageNumber="232">Figs. 7–17</figureCitation>
|
||
)
|
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A665449505426673A2FEC4FCA0C134" pageId="11" pageNumber="232" type="materials_examined">
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<paragraph id="9B0336CF9505426673A2FEC4FF3CC30C" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
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<emphasis id="A9C8EADD9505426673A2FEC4FE9EC374" bold="true" box="[151,330,259,285]" pageId="11" pageNumber="232">Type material.</emphasis>
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<typeStatus id="4407886D950542667260FEC4FE16C374" box="[341,450,259,285]" pageId="11" pageNumber="232" type="holotype">Holotype</typeStatus>
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||
:
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<collectionCode id="FDADAE0A9505426672E4FEC4FDE8C374" box="[465,572,259,285]" pageId="11" pageNumber="232">CBM-ZC</collectionCode>
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15952, ovigerous female (10.2×
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<quantity id="5C449B2A950542667089FEC4FBFDC377" box="[956,1065,259,286]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="11" pageNumber="232" unit="mm" value="11.8">11.8 mm</quantity>
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), Kumi Village,
|
||
<collectingCountry id="E3AB765F9505426677DBFEC4FACCC374" box="[1262,1304,259,285]" name="Fiji" pageId="11" pageNumber="232">Viti</collectingCountry>
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Leve,
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<collectingCountry id="E3AB765F95054266765BFEC4FA4CC374" box="[1390,1432,259,285]" name="Fiji" pageId="11" pageNumber="232">Fiji</collectingCountry>
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,
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,
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, intertidal, associated with
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<taxonomicName id="5CBC4D4C9505426671CDFEE0FC8EC328" box="[760,858,295,321]" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcoida" pageId="11" pageNumber="232" phylum="Mollusca" rank="genus">
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<emphasis id="A9C8EADD9505426671CDFEE0FC8EC328" box="[760,858,295,321]" italics="true" pageId="11" pageNumber="232">Anadara</emphasis>
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</taxonomicName>
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sp. (
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<taxonomicName id="5CBC4D4C9505426670A2FEE0FC28C328" authorityName="Linnaeus" authorityYear="1758" box="[919,1020,295,321]" class="Bivalvia" kingdom="Animalia" pageId="11" pageNumber="232" phylum="Mollusca" rank="class">Bivalvia</taxonomicName>
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:
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<taxonomicName id="5CBC4D4C95054266773DFEE0FBB1C32B" authorityName="Lamarck" authorityYear="1809" box="[1032,1125,295,322]" class="Bivalvia" family="Arcidae" kingdom="Animalia" order="Arcoida" pageId="11" pageNumber="232" phylum="Mollusca" rank="family">Arcidae</taxonomicName>
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),
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||
<date id="EF02100F950542667749FEE0FAE3C328" box="[1148,1335,295,321]" pageId="11" pageNumber="232" value="2010-08-16">16 August 2010</date>
|
||
, coll. K. Kawai.
|
||
</paragraph>
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<paragraph id="9B0336CF9505426673F2FEB7FCD8C0A4" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
<materialsCitation id="2BD43C929505426673F2FEB7FF6CC3C7" ID-GBIF-Occurrence="3019935311" collectionCode="V" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15962" specimenCount="2" typeStatus="paratype">
|
||
<typeStatus id="4407886D9505426673F2FEB7FEE0C3E3" box="[199,308,368,394]" pageId="11" pageNumber="232" type="paratype">Paratypes</typeStatus>
|
||
(all but
|
||
<specimenCode id="CB1A9EB49505426672A1FEA8FD99C3E3" box="[404,589,367,394]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15962</specimenCode>
|
||
from same locality and host as
|
||
<typeStatus id="4407886D950542667083FEA8FBC8C3E0" box="[950,1052,367,393]" pageId="11" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
; females are all in post-hard stage
|
||
<collectionCode id="FDADAE0A9505426673A2FE53FF7BC3C7" box="[151,175,404,430]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="232" type="Museum">V</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="2BD43C9295054266738CFE53FF07C3B8" ID-GBIF-Occurrence="3019935301" collectingDate="2010-08-16" collectionCode="ZRC" pageId="11" pageNumber="232" specimenCode="ZRC 2019.1877" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
:
|
||
<specimenCode id="CB1A9EB49505426673F0FE54FEA9C3C4" box="[197,381,403,429]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">ZRC 2019.1877</specimenCode>
|
||
,
|
||
<specimenCount id="8DBAFD469505426672BCFE54FE3FC3C4" box="[393,491,403,429]" pageId="11" pageNumber="232" type="female">1 female</specimenCount>
|
||
(8.2×
|
||
<quantity id="5C449B2A95054266711AFE54FD5FC3C7" box="[559,651,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="11" pageNumber="232" unit="mm" value="8.9">8.9 mm</quantity>
|
||
), 3 ovigerous females (8.4×9.6, 9.7×10.9, 11.9×
|
||
<quantity id="5C449B2A950542667799FE54FAC1C3C7" box="[1196,1301,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="11" pageNumber="232" unit="mm" value="12.3">12.3 mm</quantity>
|
||
),
|
||
<date id="EF02100F95054266761CFE54FF07C3B8" pageId="11" pageNumber="232" value="2010-08-16">
|
||
<collectingDate id="FF46E9E795054266761CFE54FF07C3B8" pageId="11" pageNumber="232" value="2010-08-16">16 August 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C929505426673EBFE70FAF2C3BB" ID-GBIF-Occurrence="3019935310" box="[222,1318,439,466]" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15953" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB49505426673EBFE70FE4CC3BB" box="[222,408,439,466]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15953</specimenCode>
|
||
,
|
||
<specimenCount id="8DBAFD46950542667296FE70FDDBC3B8" box="[419,527,439,465]" pageId="11" pageNumber="232" type="female">2 females</specimenCount>
|
||
(5.0×5.5, 6.4×
|
||
<quantity id="5C449B2A950542667181FE70FCDAC3BB" box="[692,782,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="11" pageNumber="232" unit="mm" value="6.9">6.9 mm</quantity>
|
||
), 17 ovigerous females (7.4×8.5–9.6×
|
||
<quantity id="5C449B2A950542667783FE70FACAC3BB" box="[1206,1310,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="11" pageNumber="232" unit="mm" value="13.4">13.4 mm</quantity>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C92950542667604FE70FC93C39C" ID-GBIF-Occurrence="3019935322" collectingDate="2010-08-16" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15954, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB4950542667604FE70FF34C39F" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15954</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB49505426673DFFE1CFF2CC39C" box="[234,248,475,501]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
post-hard stage male (6.8×
|
||
<quantity id="5C449B2A950542667113FE1CFD55C39F" box="[550,641,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="11" pageNumber="232" unit="mm" value="6.2">6.2 mm</quantity>
|
||
),
|
||
<date id="EF02100F9505426671A6FE1CFC93C39C" box="[659,839,475,501]" pageId="11" pageNumber="232" value="2010-08-16">
|
||
<collectingDate id="FF46E9E79505426671A6FE1CFC93C39C" box="[659,839,475,501]" pageId="11" pageNumber="232" value="2010-08-16">16 August 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C92950542667064FE1CFE9AC070" ID-GBIF-Occurrence="3019935320" collectingDate="2010-08-16" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15956, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB4950542667064FE1CFBDFC39F" box="[849,1035,475,502]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15956</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667720FE1CFBF7C39C" box="[1045,1059,475,501]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
hard stage female, (5.2×
|
||
<quantity id="5C449B2A950542667603FE1CFA44C39F" box="[1334,1424,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="11" pageNumber="232" unit="mm" value="5.2">5.2 mm</quantity>
|
||
),
|
||
<date id="EF02100F9505426673A2FE38FE9AC070" box="[151,334,511,537]" pageId="11" pageNumber="232" value="2010-08-16">
|
||
<collectingDate id="FF46E9E79505426673A2FE38FE9AC070" box="[151,334,511,537]" pageId="11" pageNumber="232" value="2010-08-16">16 August 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C9295054266726FFE38FB80C070" ID-GBIF-Occurrence="3019935308" box="[346,1108,511,538]" collectingDate="2010-08-16" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15957, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB495054266726FFE38FDC2C073" box="[346,534,511,538]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15957</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667117FE38FDE4C070" box="[546,560,511,537]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
hard stage male (5.0×
|
||
<quantity id="5C449B2A950542667019FE38FC5DC073" box="[812,905,511,538]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="232" unit="mm" value="4.6">4.6 mm</quantity>
|
||
),
|
||
<date id="EF02100F9505426670A9FE38FB80C070" box="[924,1108,511,537]" pageId="11" pageNumber="232" value="2010-08-16">
|
||
<collectingDate id="FF46E9E79505426670A9FE38FB80C070" box="[924,1108,511,537]" pageId="11" pageNumber="232" value="2010-08-16">16 August 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C9295054266776AFE38FD01C054" ID-GBIF-Occurrence="3019935316" collectingDate="2020-08-16" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15958, 2" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB495054266776AFE38FAC8C073" box="[1119,1308,511,538]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15958</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667612FE38FAE1C070" box="[1319,1333,511,537]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">2</specimenCode>
|
||
pre-hard stage male (3.2×3.2, 3.4×
|
||
<quantity id="5C449B2A950542667281FDE4FDDAC057" box="[436,526,547,574]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="232" unit="mm" value="3.3">3.3 mm</quantity>
|
||
),
|
||
<date id="EF02100F950542667115FDE4FD01C054" box="[544,725,547,573]" pageId="11" pageNumber="232" value="2020-08-16">
|
||
<collectingDate id="FF46E9E7950542667115FDE4FD01C054" box="[544,725,547,573]" pageId="11" pageNumber="232" value="2020-08-16">16 August 2020</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C929505426671EAFDE4FEDEC008" ID-GBIF-Occurrence="3019935309" collectingDate="2010-08-16" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15959, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB49505426671EAFDE4FC4EC057" box="[735,922,547,574]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15959</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667090FDE4FC67C054" box="[933,947,547,573]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
post-hard stage male (5.8×
|
||
<quantity id="5C449B2A9505426677D7FDE4FAE9C057" box="[1250,1341,547,574]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="11" pageNumber="232" unit="mm" value="5.8">5.8 mm</quantity>
|
||
),
|
||
<date id="EF02100F950542667665FDE4FEDEC008" pageId="11" pageNumber="232" value="2010-08-16">
|
||
<collectingDate id="FF46E9E7950542667665FDE4FEDEC008" pageId="11" pageNumber="232" value="2010-08-16">16 August 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C92950542667223FD80FB90C008" ID-GBIF-Occurrence="3019935315" box="[278,1092,583,610]" collectingDate="2020-08-16" collectionCode="ZRC" pageId="11" pageNumber="232" specimenCode="ZRC 2020.0007, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB4950542667223FD80FE1AC008" box="[278,462,583,609]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">ZRC 2020.0007</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB49505426672EFFD80FE3CC008" box="[474,488,583,609]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">1</specimenCode>
|
||
post-hard stage male (5.4×
|
||
<quantity id="5C449B2A950542667028FD80FCADC00B" box="[797,889,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="232" unit="mm" value="4.8">4.8 mm</quantity>
|
||
),
|
||
<date id="EF02100F9505426670B9FD80FB90C008" box="[908,1092,583,609]" pageId="11" pageNumber="232" value="2020-08-16">
|
||
<collectingDate id="FF46E9E79505426670B9FD80FB90C008" box="[908,1092,583,609]" pageId="11" pageNumber="232" value="2020-08-16">16 August 2020</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C9295054266777AFD80FDA6C0EC" ID-GBIF-Occurrence="3019935307" collectingDate="2011-09-06" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15960, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB495054266777AFD80FAD8C00B" box="[1103,1292,583,610]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15960</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB495054266762DFD80FAF2C008" box="[1304,1318,583,609]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
ovigerous female (9.1×
|
||
<quantity id="5C449B2A950542667213FDACFE44C0EF" box="[294,400,619,646]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="11" pageNumber="232" unit="mm" value="10.7">10.7 mm</quantity>
|
||
),
|
||
<date id="EF02100F950542667296FDACFDA6C0EC" box="[419,626,619,645]" pageId="11" pageNumber="232" value="2011-09-06">
|
||
<collectingDate id="FF46E9E7950542667296FDACFDA6C0EC" box="[419,626,619,645]" pageId="11" pageNumber="232" value="2011-09-06">6 September 2011</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C9295054266714AFDACFF06C0C0" ID-GBIF-Occurrence="3019935304" collectingDate="2018-08" collectionCode="CBM-ZC" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15961, 7" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB495054266714AFDACFCEFC0EF" box="[639,827,619,646]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15961</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667073FDABFC80C0EF" box="[838,852,620,646]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">7</specimenCode>
|
||
ovigerous females (7.2×8.6–9.5×
|
||
<quantity id="5C449B2A9505426677E4FDACFAEDC0EF" box="[1233,1337,619,646]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="11" pageNumber="232" unit="mm" value="10.7">10.7 mm</quantity>
|
||
),
|
||
<date id="EF02100F95054266767EFDACFF06C0C0" pageId="11" pageNumber="232" value="2018-08">
|
||
<collectingDate id="FF46E9E795054266767EFDACFF06C0C0" pageId="11" pageNumber="232" value="2018-08">August 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C929505426673EAFD48FB95C0C0" ID-GBIF-Occurrence="3019935323" box="[223,1089,655,682]" collectingDate="2018-08" collectionCode="ZRC" pageId="11" pageNumber="232" specimenCode="ZRC 2020.0008, 5" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB49505426673EAFD48FE4CC0C0" box="[223,408,655,681]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">ZRC 2020.0008</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667291FD48FE66C0C3" box="[420,434,655,682]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">5</specimenCode>
|
||
ovigerous females (8.0×9.4–9.3×
|
||
<quantity id="5C449B2A950542667004FD48FC4EC0C3" box="[817,922,655,682]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="11" pageNumber="232" unit="mm" value="10.7">10.7 mm</quantity>
|
||
),
|
||
<date id="EF02100F950542667098FD48FB95C0C0" box="[941,1089,655,681]" pageId="11" pageNumber="232" value="2018-08">
|
||
<collectingDate id="FF46E9E7950542667098FD48FB95C0C0" box="[941,1089,655,681]" pageId="11" pageNumber="232" value="2018-08">August 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C9295054266777BFD48FCDDC0A4" ID-GBIF-Occurrence="3019935302" collectingDate="2019-02-22" collectionCode="CBM-ZC" collectorName="Ucikumi" location="Kumi Village" pageId="11" pageNumber="232" specimenCode="CBM-ZC 19562, 1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="CB1A9EB495054266777BFD48FADFC0C3" box="[1102,1291,655,682]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 19562</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB4950542667622FD48FAF1C0C0" box="[1303,1317,655,681]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
ovigerous female,
|
||
<collectorName id="364953199505426673C4FD74FE82C0A4" box="[241,342,691,717]" pageId="11" pageNumber="232">Ucikumi</collectorName>
|
||
, near
|
||
<location id="9E6360149505426672ACFD74FDE7C0A4" LSID="urn:lsid:plazi:treatment:131587D99505427A73A2FF50FAA3C008:9E6360149505426672ACFD74FDE7C0A4" box="[409,563,691,717]" name="Kumi Village" pageId="11" pageNumber="232">Kumi Village</location>
|
||
,
|
||
<date id="EF02100F95054266710BFD74FCDDC0A4" box="[574,777,691,717]" pageId="11" pageNumber="232" value="2019-02-22">
|
||
<collectingDate id="FF46E9E795054266710BFD74FCDDC0A4" box="[574,777,691,717]" pageId="11" pageNumber="232" value="2019-02-22">22 February 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426673F2FD10FAD8C17C" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
<materialsCitation id="2BD43C929505426673F2FD10FF29C17C" ID-GBIF-Occurrence="3019935305" collectionCode="CBM-ZC" country="Fiji" pageId="11" pageNumber="232" specimenCode="CBM-ZC 15955, 1" specimenCount="1">
|
||
<emphasis id="A9C8EADD9505426673F2FD10FE2FC098" bold="true" box="[199,507,727,753]" pageId="11" pageNumber="232">Other material examined.</emphasis>
|
||
<collectingCountry id="E3AB765F950542667134FD10FDF8C098" box="[513,556,727,753]" name="Fiji" pageId="11" pageNumber="232">Fiji</collectingCountry>
|
||
.
|
||
<specimenCode id="CB1A9EB4950542667103FD10FD25C09B" box="[566,753,727,754]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">CBM-ZC 15955</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB49505426671CEFD10FCDDC098" box="[763,777,727,753]" collectionCode="CBM-ZC" pageId="11" pageNumber="232">1</specimenCode>
|
||
young female (post-hard stage; 4.0×
|
||
<quantity id="5C449B2A950542667791FD10FB29C09B" box="[1188,1277,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="232" unit="mm" value="3.9">3.9 mm</quantity>
|
||
), same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2BD43C9295054266723DFD3CFADDC17C" ID-GBIF-Occurrence="3019935314" box="[264,1289,763,790]" collectionCode="ZRC" country="Fiji" pageId="11" pageNumber="232" specimenCode="ZRC 2019.1878, 4" specimenCount="1">
|
||
<specimenCode id="CB1A9EB495054266723DFD3CFE6BC17C" box="[264,447,763,789]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">ZRC 2019.1878</specimenCode>
|
||
,
|
||
<specimenCode id="CB1A9EB49505426672FFFD3CFE0CC17C" box="[458,472,763,789]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">4</specimenCode>
|
||
poorly preserved females (3.3×4.2–4.6×6.0 mm), same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426673F2FCD8FCA0C134" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
<collectingCountry id="E3AB765F9505426673F2FCD8FEECC150" box="[199,312,799,825]" name="Indonesia" pageId="11" pageNumber="232">Indonesia</collectingCountry>
|
||
.
|
||
<collectionCode id="FDADAE0A950542667271FCD8FEAEC150" box="[324,378,799,825]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="232">ZRC</collectionCode>
|
||
2019.1025,
|
||
<specimenCount id="8DBAFD4695054266713FFCD8FDB9C150" box="[522,621,799,825]" pageId="11" pageNumber="232" type="female">1 female</specimenCount>
|
||
(8.4×
|
||
<quantity id="5C449B2A950542667185FCD8FCD9C153" box="[688,781,799,826]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="11" pageNumber="232" unit="mm" value="9.4">9.4 mm</quantity>
|
||
), Teluk Kodek, Lombok, intertidal, in
|
||
<taxonomicName id="5CBC4D4C9505426677F3FCD8FE18C137" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcoida" pageId="11" pageNumber="232" phylum="Mollusca" rank="species" species="antiquata">
|
||
<emphasis id="A9C8EADD9505426677F3FCD8FA48C150" box="[1222,1436,799,825]" italics="true" pageId="11" pageNumber="232">Anadara antiquata</emphasis>
|
||
(
|
||
<bibRefCitation id="FF2D4B3E9505426673AAFC84FE9AC137" author="Linnaeus, C." box="[159,334,835,862]" pageId="11" pageNumber="232" refId="ref14047" refString="Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Vol. 1. 10 th Revised Edition. Laurentius Salvius, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
) (Arcidae)
|
||
</taxonomicName>
|
||
,
|
||
<date id="EF02100F9505426672E2FC84FD66C134" box="[471,690,835,861]" pageId="11" pageNumber="232" value="2013-11-12">12 November 2013</date>
|
||
, coll. S. Dwiono.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A665449505427973F2FCA0FF17C4B1" lastPageId="20" lastPageNumber="241" pageId="11" pageNumber="232" type="description">
|
||
<paragraph id="9B0336CF9505426673F2FCA0FCA4C184" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
<emphasis id="A9C8EADD9505426673F2FCA0FE8DC1E8" bold="true" box="[199,345,871,897]" pageId="11" pageNumber="232">Description.</emphasis>
|
||
<emphasis id="A9C8EADD950542667255FCA0FD1AC1E8" box="[352,718,871,897]" italics="true" pageId="11" pageNumber="232">Adult females (post-hard stages)</emphasis>
|
||
. Carapace (
|
||
<figureCitation id="03872A4A950542667063FCA0FC77C1E8" box="[854,931,871,897]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7</figureCitation>
|
||
A–D, 8A) subcircular, 1.1–1.3 times as wide as long; dorsal surface gently convex, smooth, glabrous; lateral surfaces also smooth, glabrous; front projecting anteriorly beyond orbits, entire, margin slightly convex; anterior quarter of anterolateral margin gently sloping posteriorly, forming slight angle with rest of slightly convex margin.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426673F2FC3FFB65C678" blockId="11.[151,1437,259,2014]" box="[199,1201,1015,1042]" pageId="11" pageNumber="232">
|
||
Eyes (
|
||
<figureCitation id="03872A4A950542667238FC30FE46C67B" box="[269,402,1015,1042]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7C, D</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9505426672A8FC30FE34C67B" box="[413,480,1015,1042]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="11" pageNumber="232">8A, B</figureCitation>
|
||
) small, visible in dorsal view, movable, completely filling orbit.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426673F2FBDCFD4DC630" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
Antennular peduncles (
|
||
<figureCitation id="03872A4A9505426672E4FBDCFDFDC65C" box="[465,553,1051,1077]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="11" pageNumber="232">Fig. 8B</figureCitation>
|
||
) folded obliquely into antennular fossae. Antenna (
|
||
<figureCitation id="03872A4A95054266774EFBDCFB00C65C" box="[1147,1236,1051,1077]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="11" pageNumber="232">Fig. 8B</figureCitation>
|
||
) extending along lateral margin of front; article 2 subtriangular.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426673F2FBA4FD6AC683" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
Maxilliped 3 (
|
||
<figureCitation id="03872A4A95054266725DFBA4FE2AC614" box="[360,510,1123,1150]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="11" pageNumber="232">Fig. 8C, F, G</figureCitation>
|
||
) outer surface glabrous; propodus moderately stout, distinctly longer than carpus; dactylus short, digitiform, inserted submedially on flexor margin of propodus, tip just reaching propodal apex; ischiomerus completely fused, distinctly longer than wide, mesial margin angular at widest point. Exopod slender, about half length of ischiomerus (not illustrated).
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426673F2FB34FC10C43B" blockId="11.[151,1437,259,2014]" pageId="11" pageNumber="232">
|
||
Chelipeds (
|
||
<figureCitation id="03872A4A950542667273FB34FE3CC767" box="[326,488,1267,1294]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7A, B, E</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9505426672C1FB34FDD5C764" box="[500,513,1267,1293]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="11" pageNumber="232">9</figureCitation>
|
||
A–E) moderately long, symmetrical. Dactylus about 0.6 times as long as palm, terminating in hooked, acute tip; upper margin rounded, gently convex; occlusal margin with 1 prominent, proximally directed, triangular tooth proximally and row of sparse short setae distal to occlusal tooth. Palm slightly widened distally, 1.9 times as long as high; outer surface glabrous; inner surface almost glabrous, but with longitudinal row of short setae adjacent to lower margin, extending onto fixed finger, becoming multiple rows. Fixed finger terminating in hooked, acute tip, bearing short longitudinal row of stiff setae on inner surface distally; occlusal margin with 2 small teeth flanking occlusal tooth on dactylus, margin between 2 teeth denticulate, margin distal to occlusal tooth slightly convex, laminar, bearing row of minute stiff setae. Carpus short, cup-shaped, glabrous on dorsal and outer surfaces; inner surface with sparse thin setae. Merus nearly straight slightly compressed laterally, slightly widened distally; outer and inner surfaces almost glabrous, former gently convex.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9505426173F2F99CFBA2C3C4" blockId="11.[151,1437,259,2014]" lastBlockId="12.[151,1437,151,1977]" lastPageId="12" lastPageNumber="233" pageId="11" pageNumber="232">
|
||
P2, P3 and P5 symmetrical, P4 longest, asymmetrical; relative length of meri P4>P3>P2>P5. P2 (
|
||
<figureCitation id="03872A4A9505426677C8F99CFA98C41F" box="[1277,1356,1627,1654]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950542667663F99CFA40C41C" box="[1366,1428,1627,1653]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="11" pageNumber="232">9F, G</figureCitation>
|
||
) moderately long, slender, entirely glabrous; dactylus short, noticeably curved, terminating in acute tip, 0.5–0.6 times as long as propodus; propodus slightly narrowing distally, longer than carpus; merus somewhat compressed, almost straight. P3 (
|
||
<figureCitation id="03872A4A950542667210F900FEA7C48B" box="[293,371,1735,1762]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A95054266724AF900FE6CC48B" box="[383,440,1735,1762]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="11" pageNumber="232">9H, I</figureCitation>
|
||
) generally similar to but slightly longer than P2; dactylus short, curved, about half-length of propodus. Longer P4 (
|
||
<figureCitation id="03872A4A950542667281F92CFDD0C56F" box="[436,516,1771,1798]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950542667125F92CFD8CC56F" box="[528,600,1771,1798]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="11" pageNumber="232">9L, M</figureCitation>
|
||
)) about 1.2 times as long as shorter P4; dactylus slender, 0.9–1.0 times as long as propodus, approximately 10 times as long as wide, gently curved proximally, terminating in acute tip; surfaces almost glabrous except for subapical long seta on flexor margin; propodus slightly curved, more slender than those of P2 and P3; carpus curved at base; merus more slender than those of P2 and P3, glabrous; ischium-merus articulation oblique, ischium glabrous. Shorter P4 (
|
||
<figureCitation id="03872A4A9505426671F9F8BCFCCEC5FF" box="[716,794,1915,1942]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="11" pageNumber="232">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950542667010F8BCFCB4C5FF" box="[805,864,1915,1942]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="11" pageNumber="232">9J, K</figureCitation>
|
||
) dactylus slightly arcuate, approximately 6 times as long as wide, 0.6–0.7 times as long as propodus, still longer than dactyli of P2 and P3, terminating in curved corneous claw; extensor surface with scattered minute setae; flexor margin with irregular rows of short, feather-like setae; propodus, carpus and merus generally similar to those of longer P4, but respective article shorter than corresponding article of longer side; ischium with sparse short setae on posterior surface. P5 (
|
||
<figureCitation id="03872A4A950242617723FF7CFBB2C2BC" box="[1046,1126,187,213]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="12" pageNumber="233">
|
||
Figs.
|
||
<geoCoordinate id="FE885008950242617763FF7CFBB2C2BC" box="[1110,1126,187,213]" degrees="7, 9" direction="north" orientation="latitude" pageId="12" pageNumber="233" precision="5555" value="7.9">7</geoCoordinate>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950242617746FF7CFB6DC2BC" box="[1139,1209,187,213]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="12" pageNumber="233">9N, O</figureCitation>
|
||
) propodus to merus shorter and slender than corresponding articles of P2 and P3; dactylus long (but shorter than that of longer P4), subequal in length to propodus, approximately 9 times as long as wide, slightly arcuate, gradually tapering to acute tip, extensor margin and upper surface with scattered short setae in distal half, flexor margin with numerous short setae becoming dense and longer distally, no comb-like rows of minute spiniform setae; propodus faintly curved, almost glabrous except for subterminal tuft of long setae on flexor margin; carpus glabrous, substantially curved; merus faintly sinuous, not narrowing proximally, glabrous; merus-ischium articulation straight.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FE70FC33C070" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
Thoracic sternite very wide. Sternites 1–4 (
|
||
<figureCitation id="03872A4A95024261718EFE70FCC0C3B8" box="[699,788,439,465]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="12" pageNumber="233">Fig. 8D</figureCitation>
|
||
) fused, slightly narrowing anteriorly; narrow incision at base of maxilliped 3, suggesting boundary between sternites 3 and 4, still discernible; anteromedian margin faintly concave; anterolateral angles produced in rounded lobe; surface almost flat.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FDE4FCC9C008" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
Pleon (
|
||
<figureCitation id="03872A4A950242617223FDE4FEB1C054" box="[278,357,547,573]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="12" pageNumber="233">
|
||
Figs.
|
||
<geoCoordinate id="FE885008950242617260FDE4FEB1C054" box="[341,357,547,573]" degrees="7, 8" direction="east" orientation="longitude" pageId="12" pageNumber="233" precision="5555" value="7.8">7</geoCoordinate>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950242617244FDE4FE45C054" box="[369,401,547,573]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="12" pageNumber="233">8E</figureCitation>
|
||
) extremely wide, extending to buccal region, covering bases of ambulatory legs; telson very broad, partially recessed into distal margin of pleomere 6.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FDACFC54C17C" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
<emphasis id="A9C8EADD9502426173F2FDACFE43C0ED" box="[199,407,619,645]" italics="true" pageId="12" pageNumber="233">Female hard stage</emphasis>
|
||
. Generally similar to male hard stage (see below) (
|
||
<figureCitation id="03872A4A9502426170E5FDACFBECC0EC" box="[976,1080,619,645]" captionStart="FIGURE 10" captionStartId="16.[151,250,1553,1577]" captionTargetBox="[160,1425,190,1519]" captionTargetId="figure-20@16.[151,1436,181,1529]" captionTargetPageId="16" captionText="FIGURE 10. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, habitus, dorsal view; B, thoracic sternite 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, pleon and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4401483" httpUri="https://zenodo.org/record/4401483/files/figure.png" pageId="12" pageNumber="233">Fig. 10A</figureCitation>
|
||
). P3 and P4 provided with natatory setae (
|
||
<figureCitation id="03872A4A950242617225FD48FEB9C0C0" box="[272,365,655,681]" captionStart="FIGURE 10" captionStartId="16.[151,250,1553,1577]" captionTargetBox="[160,1425,190,1519]" captionTargetId="figure-20@16.[151,1436,181,1529]" captionTargetPageId="16" captionText="FIGURE 10. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, habitus, dorsal view; B, thoracic sternite 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, pleon and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4401483" httpUri="https://zenodo.org/record/4401483/files/figure.png" pageId="12" pageNumber="233">Figs. 10</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A95024261724DFD48FE45C0C0" box="[376,401,655,681]" captionStart="FIGURE 11" captionStartId="17.[151,250,1157,1181]" captionTargetBox="[160,1425,189,1127]" captionTargetId="figure-17@17.[151,1436,181,1133]" captionTargetPageId="17" captionText="FIGURE 11. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus." figureDoi="http://doi.org/10.5281/zenodo.4401485" httpUri="https://zenodo.org/record/4401485/files/figure.png" pageId="12" pageNumber="233">11</figureCitation>
|
||
C–E). P2 (
|
||
<figureCitation id="03872A4A950242617133FD48FDBCC0C0" box="[518,616,655,681]" captionStart="FIGURE 11" captionStartId="17.[151,250,1157,1181]" captionTargetBox="[160,1425,189,1127]" captionTargetId="figure-17@17.[151,1436,181,1133]" captionTargetPageId="17" captionText="FIGURE 11. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus." figureDoi="http://doi.org/10.5281/zenodo.4401485" httpUri="https://zenodo.org/record/4401485/files/figure.png" pageId="12" pageNumber="233">Fig. 11B</figureCitation>
|
||
) dactylus with row of short setae on flexor margin. P3 (
|
||
<figureCitation id="03872A4A9502426177FAFD48FAE4C0C0" box="[1231,1328,655,681]" captionStart="FIGURE 11" captionStartId="17.[151,250,1157,1181]" captionTargetBox="[160,1425,189,1127]" captionTargetId="figure-17@17.[151,1436,181,1133]" captionTargetPageId="17" captionText="FIGURE 11. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus." figureDoi="http://doi.org/10.5281/zenodo.4401485" httpUri="https://zenodo.org/record/4401485/files/figure.png" pageId="12" pageNumber="233">Fig. 11C</figureCitation>
|
||
) dactylus more stout than in hard stage male. Longer P4 (
|
||
<figureCitation id="03872A4A95024261719FFD74FCDFC0A4" box="[682,779,691,717]" captionStart="FIGURE 11" captionStartId="17.[151,250,1157,1181]" captionTargetBox="[160,1425,189,1127]" captionTargetId="figure-17@17.[151,1436,181,1133]" captionTargetPageId="17" captionText="FIGURE 11. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus." figureDoi="http://doi.org/10.5281/zenodo.4401485" httpUri="https://zenodo.org/record/4401485/files/figure.png" pageId="12" pageNumber="233">Fig. 11E</figureCitation>
|
||
) dactylus only with subapical setae on flexor margin, as in post-hard stage. Pleon (
|
||
<figureCitation id="03872A4A950242617295FD10FDD7C098" box="[416,515,727,753]" captionStart="FIGURE 10" captionStartId="16.[151,250,1553,1577]" captionTargetBox="[160,1425,190,1519]" captionTargetId="figure-20@16.[151,1436,181,1529]" captionTargetPageId="16" captionText="FIGURE 10. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, habitus, dorsal view; B, thoracic sternite 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, pleon and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4401483" httpUri="https://zenodo.org/record/4401483/files/figure.png" pageId="12" pageNumber="233">Fig. 10E</figureCitation>
|
||
) slightly wider than that of hard males; pleomeres 2–4 each with paired biramous pleopods; telson 1.3 times wider than long, roundly subhexagonal.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FCD8FBF2C1CC" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
<emphasis id="A9C8EADD9502426173F2FCD8FE56C151" box="[199,386,799,825]" italics="true" pageId="12" pageNumber="233">Male hard stage.</emphasis>
|
||
Carapace (
|
||
<figureCitation id="03872A4A9502426172CBFCD8FD8EC150" box="[510,602,799,825]" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950242617150FCD8FD41C150" box="[613,661,799,825]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="12" pageNumber="233">13A</figureCitation>
|
||
) well calcified, rounded, subcircular, 1.1 times as long as wide; dorsal surface convex, without demarcation of regions, with scattered minute pits laterally; front weakly projecting anteriorly beyond orbits, margin nearly straight with faint median notch in dorsal view; lateral angle of front produced into small triangular process directed downward, but not closing orbit (
|
||
<figureCitation id="03872A4A950242617086FC4CFBCDC1CC" box="[947,1049,907,933]" captionStart="FIGURE 11" captionStartId="17.[151,250,1157,1181]" captionTargetBox="[160,1425,189,1127]" captionTargetId="figure-17@17.[151,1436,181,1133]" captionTargetPageId="17" captionText="FIGURE 11. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus." figureDoi="http://doi.org/10.5281/zenodo.4401485" httpUri="https://zenodo.org/record/4401485/files/figure.png" pageId="12" pageNumber="233">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FC68FB1CC1A0" blockId="12.[151,1437,151,1977]" box="[199,1224,943,969]" pageId="12" pageNumber="233">
|
||
Eyes (
|
||
<figureCitation id="03872A4A950242617238FC68FE47C1A0" box="[269,403,943,969]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="12" pageNumber="233">Fig. 13A, B</figureCitation>
|
||
) moderately small, clearly visible in dorsal view completely filling orbit.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FC14FC57C678" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
Antennular peduncles (
|
||
<figureCitation id="03872A4A9502426172F9FC14FDE5C184" box="[460,561,979,1005]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="12" pageNumber="233">Fig. 13B</figureCitation>
|
||
) folded obliquely into antennular fossae. Antenna (
|
||
<figureCitation id="03872A4A950242617741FC14FB0CC184" box="[1140,1240,979,1005]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="12" pageNumber="233">Fig. 13B</figureCitation>
|
||
) extending along lateral margin of front; article 2 subtrapezoidal, narrowing distally.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FBDCFF2FC630" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
Maxilliped 3 (
|
||
<figureCitation id="03872A4A95024261725EFBDCFE05C65C" box="[363,465,1051,1077]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="12" pageNumber="233">Fig. 13C</figureCitation>
|
||
) generally similar to that of females, but inner margin of ischiomerus more strongly concave.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FBA4FBAEC7F4" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
Chelipeds (
|
||
<figureCitation id="03872A4A950242617273FBA4FE76C614" box="[326,418,1123,1149]" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950242617298FBA4FE2AC614" box="[429,510,1123,1149]" captionStart="FIGURE 14" captionStartId="20.[151,250,1151,1175]" captionTargetBox="[157,1428,188,1117]" captionTargetId="figure-20@20.[151,1436,181,1127]" captionTargetPageId="20" captionText="FIGURE 14. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401491" httpUri="https://zenodo.org/record/4401491/files/figure.png" pageId="12" pageNumber="233">14A, B</figureCitation>
|
||
) moderately large, symmetrical. Dactylus about 0.8 times as long as palm, strongly curved, terminating in hooked, acute tip; upper margin rounded; occlusal margin with 1 conspicuous triangular tooth proximal to midlength and with row of sparse short setae. Palm slightly widened distally, 1.3 times as long as high; outer surface almost glabrous, but with sparse minute pits; inner surface almost glabrous, but with longitudinal row of short to long setae adjacent to lower margin, extending onto fixed finger, becoming multiple rows. Fixed finger terminating in hooked, acute tip; occlusal margin with 2 or 3 small teeth proximal to midlength, flanking occlusal tooth on dactylus, and with row of minute setae distal to midlength. Carpus distinctly shorter than palm, somewhat inflated, with sparse minute pits. Merus slightly curved inwards, slightly compressed laterally; outer surface convex, glabrous, lower distal angle somewhat produced; inner surface with few short setae.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9502426173F2FA60FBCFC5D0" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="233">
|
||
P2, P3 and P5 symmetrical, P4 longest, asymmetrical; natatory setae present on propodi and carpi of P3 and P4 (
|
||
<figureCitation id="03872A4A9502426173F0FA0CFEF7C78C" box="[197,291,1483,1509]" captionStart="FIGURE 10" captionStartId="16.[151,250,1553,1577]" captionTargetBox="[160,1425,190,1519]" captionTargetId="figure-20@16.[151,1436,181,1529]" captionTargetPageId="16" captionText="FIGURE 10. Arcotheres ocularius n. sp., paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, habitus, dorsal view; B, thoracic sternite 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, pleon and telson, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4401483" httpUri="https://zenodo.org/record/4401483/files/figure.png" pageId="12" pageNumber="233">Figs. 10</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A950242617205FA0CFE9FC78C" box="[304,331,1483,1509]" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">12</figureCitation>
|
||
D–F). P2 (
|
||
<figureCitation id="03872A4A9502426172F6FA0CFDF5C78C" box="[451,545,1483,1509]" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A95024261711BFA0CFD8AC78C" box="[558,606,1483,1509]" captionStart="FIGURE 14" captionStartId="20.[151,250,1151,1175]" captionTargetBox="[157,1428,188,1117]" captionTargetId="figure-20@20.[151,1436,181,1127]" captionTargetPageId="20" captionText="FIGURE 14. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401491" httpUri="https://zenodo.org/record/4401491/files/figure.png" pageId="12" pageNumber="233">14C</figureCitation>
|
||
) generally glabrous except for row of sparse short setae on flexor margin of propodus; dactylus curved, half-length of propodus; carpus curved proximally; merus almost straight, slightly widened distally. P3 (
|
||
<figureCitation id="03872A4A9502426172B9F9D4FE3FC444" box="[396,491,1555,1581]" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9502426172CDF9D4FDFEC444" box="[504,554,1555,1581]" captionStart="FIGURE 14" captionStartId="20.[151,250,1151,1175]" captionTargetBox="[157,1428,188,1117]" captionTargetId="figure-20@20.[151,1436,181,1127]" captionTargetPageId="20" captionText="FIGURE 14. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401491" httpUri="https://zenodo.org/record/4401491/files/figure.png" pageId="12" pageNumber="233">14D</figureCitation>
|
||
) dactylus slender than that of P2, about 0.6 times as long as propodus, extensor margin with row of short setae; propodus with low of long natatory setae on upper surface adjacent to extensor margin and flexor margin, respectively; carpus upper surface with oblique low of long natatory setae. P4 (
|
||
<figureCitation id="03872A4A950242617651F99CFF1EC4F0" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">Figs. 12A</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9502426173E2F9B8FEF0C4F0" box="[215,292,1663,1689]" captionStart="FIGURE 14" captionStartId="20.[151,250,1151,1175]" captionTargetBox="[157,1428,188,1117]" captionTargetId="figure-20@20.[151,1436,181,1127]" captionTargetPageId="20" captionText="FIGURE 14. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401491" httpUri="https://zenodo.org/record/4401491/files/figure.png" pageId="12" pageNumber="233">14E, F</figureCitation>
|
||
) longer dactylus 1.3 times as long as shorter dactylus, slightly arcuate, gradually tapering distally to acute tip, extensor margin glabrous, flexor margin with row of sparse setae (distalmost subterminal seta elongate); shorter dactylus with low of long setae on extensor margin and sparse short setae on flexor margin; propodus with long natatory setae on upper surface adjacent to extensor margin and on flexor margin; carpus with oblique row of natatory setae on upper surface; merus nearly straight, glabrous. P5 (
|
||
<figureCitation id="03872A4A9502426170A0F8C8FC26C540" box="[917,1010,1807,1833]" captionStart="FIGURE 12" captionStartId="18.[151,250,2007,2031]" captionTargetBox="[159,1427,1065,1983]" captionTargetId="figure-437@18.[159,1427,1065,1983]" captionTargetPageId="18" captionText="FIGURE 12. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401487" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="12" pageNumber="233">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9502426170CBF8C8FBE4C540" box="[1022,1072,1807,1833]" captionStart="FIGURE 14" captionStartId="20.[151,250,1151,1175]" captionTargetBox="[157,1428,188,1117]" captionTargetId="figure-20@20.[151,1436,181,1127]" captionTargetPageId="20" captionText="FIGURE 14. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401491" httpUri="https://zenodo.org/record/4401491/files/figure.png" pageId="12" pageNumber="233">14G</figureCitation>
|
||
) shortest; dactylus slightly arcuate, 0.8 times as long as propodus, approximately 8 times as long as wide; extensor margin with rows of short setae at least on distal half, flexor margin with sparse longer setae; propodus almost glabrous except for sparse long setae on lower margin; carpus glabrous, gently curved; merus subcylindrical, straight, with row of moderately long setae on upper margin; ischium slightly wider than merus, about half-length of merus.
|
||
</paragraph>
|
||
<caption id="CFC366479503426073A2F850FD59C591" ID-DOI="http://doi.org/10.5281/zenodo.4401477" ID-Zenodo-Dep="4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="13" pageNumber="234" startId="13.[151,250,1943,1967]" targetBox="[163,1425,181,1919]" targetPageId="13">
|
||
<paragraph id="9B0336CF9503426073A2F850FD59C591" blockId="13.[151,1437,1943,2040]" pageId="13" pageNumber="234">
|
||
<emphasis id="A9C8EADD9503426073A2F850FEC0C5D9" bold="true" box="[151,276,1943,1968]" pageId="13" pageNumber="234">FIGURE 7.</emphasis>
|
||
<taxonomicName id="5CBC4D4C95034260722EF85FFE24C5D9" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[283,496,1944,1968]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="234" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD95034260722EF85FFE24C5D9" box="[283,496,1944,1968]" italics="true" pageId="13" pageNumber="234">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9503426072C2F85FFDE5C5D9" bold="true" box="[503,561,1944,1968]" pageId="13" pageNumber="234">
|
||
<taxonomicNameLabel id="B2FB57A69503426072C2F85FFDE5C5D9" box="[503,561,1944,1968]" pageId="13" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji. A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC366479500426373A2F8E3FE8FC5A4" ID-DOI="http://doi.org/10.5281/zenodo.4401479" ID-Zenodo-Dep="4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="14" pageNumber="235" startId="14.[151,250,1828,1852]" targetBox="[161,1426,190,1794]" targetPageId="14">
|
||
<paragraph id="9B0336CF9500426373A2F8E3FE8FC5A4" blockId="14.[151,1437,1828,1997]" pageId="14" pageNumber="235">
|
||
<emphasis id="A9C8EADD9500426373A2F8E3FEC7C554" bold="true" box="[151,275,1828,1853]" pageId="14" pageNumber="235">FIGURE 8.</emphasis>
|
||
<taxonomicName id="5CBC4D4C95004263722EF8E2FE3BC554" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[283,495,1829,1853]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="235" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD95004263722EF8E2FE3BC554" box="[283,495,1829,1853]" italics="true" pageId="14" pageNumber="235">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9500426372C0F8E2FDFBC554" bold="true" box="[501,559,1829,1853]" pageId="14" pageNumber="235">
|
||
<taxonomicNameLabel id="B2FB57A69500426372C0F8E2FDFBC554" box="[501,559,1829,1853]" pageId="14" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC366479501426273A2F88DFDE7C59B" ID-DOI="http://doi.org/10.5281/zenodo.4401481" ID-Zenodo-Dep="4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="15" pageNumber="236" startId="15.[151,250,1866,1890]" targetBox="[206,1379,190,1831]" targetPageId="15">
|
||
<paragraph id="9B0336CF9501426273A2F88DFDE7C59B" blockId="15.[151,1436,1866,2034]" pageId="15" pageNumber="236">
|
||
<emphasis id="A9C8EADD9501426273A2F88DFEC1C50B" bold="true" box="[151,277,1866,1890]" pageId="15" pageNumber="236">FIGURE 9.</emphasis>
|
||
<taxonomicName id="5CBC4D4C95014262722BF88DFE21C50B" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[286,501,1866,1890]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="236" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD95014262722BF88DFE21C50B" box="[286,501,1866,1890]" italics="true" pageId="15" pageNumber="236">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9501426272C8F88DFDEDC50B" bold="true" box="[509,569,1866,1890]" pageId="15" pageNumber="236">
|
||
<taxonomicNameLabel id="B2FB57A69501426272C8F88DFDEDC50B" box="[509,569,1866,1890]" pageId="15" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC36647951E427D73A2F9D6FBDDC418" ID-DOI="http://doi.org/10.5281/zenodo.4401483" ID-Zenodo-Dep="4401483" httpUri="https://zenodo.org/record/4401483/files/figure.png" pageId="16" pageNumber="237" startId="16.[151,250,1553,1577]" targetBox="[160,1425,190,1519]" targetPageId="16">
|
||
<paragraph id="9B0336CF951E427D73A2F9D6FBDDC418" blockId="16.[151,1436,1553,1649]" pageId="16" pageNumber="237">
|
||
<emphasis id="A9C8EADD951E427D73A2F9D6FEF6C440" bold="true" box="[151,290,1553,1577]" pageId="16" pageNumber="237">FIGURE 10.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951E427D721FF9D6FE2BC440" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[298,511,1553,1577]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="237" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951E427D721FF9D6FE2BC440" box="[298,511,1553,1577]" italics="true" pageId="16" pageNumber="237">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951E427D7132F9D6FD97C440" bold="true" box="[519,579,1553,1577]" pageId="16" pageNumber="237">
|
||
<taxonomicNameLabel id="B2FB57A6951E427D7132F9D6FD97C440" box="[519,579,1553,1577]" pageId="16" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, habitus, dorsal view; B, thoracic sternite 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, pleon and telson, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B0336CF951E427D73F2F958FE55C5DC" blockId="16.[151,1437,1695,1973]" pageId="16" pageNumber="237">
|
||
Thoracic sternum (
|
||
<figureCitation id="03872A4A951E427D7295F958FE3BC4D0" box="[416,495,1695,1721]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="16" pageNumber="237">Fig. 13</figureCitation>
|
||
D–G) broad. Sternites 1–4 fused, slightly narrowing anteriorly; narrow incision at base of maxilliped 3, suggesting boundary between sternites 3 and 4, still discernible; broad anteromedian concavity well delimited; anterolateral angles produced in rounded lobe; surface almost flat. Exposed parts of sternites 5–7 nearly smooth, each posteriorly with small but conspicuous, rounded episternites. Sternite 8 clearly visible in lateral part; lateral margin almost entire. Sterno-pleonal cavity (
|
||
<figureCitation id="03872A4A951E427D706DF8E8FC6DC520" box="[856,953,1839,1865]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="16" pageNumber="237">Fig. 13F</figureCitation>
|
||
) deep, defined by clearly delimited margin, widened anteriorly (corresponding to telson), reaching anteriorly to level of anterior ends of cheliped coxae; press button on sternite 5 relatively large, rounded; sternite 6 with pair of large, rounded protuberance accommodating lateral margin of G1.
|
||
</paragraph>
|
||
<caption id="CFC36647951F427C73A2FB42FACFC68C" ID-DOI="http://doi.org/10.5281/zenodo.4401485" ID-Zenodo-Dep="4401485" httpUri="https://zenodo.org/record/4401485/files/figure.png" pageId="17" pageNumber="238" startId="17.[151,250,1157,1181]" targetBox="[160,1425,189,1127]" targetPageId="17">
|
||
<paragraph id="9B0336CF951F427C73A2FB42FACFC68C" blockId="17.[151,1436,1157,1253]" pageId="17" pageNumber="238">
|
||
<emphasis id="A9C8EADD951F427C73A2FB42FECBC6F4" bold="true" box="[151,287,1157,1181]" pageId="17" pageNumber="238">FIGURE 11.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951F427C7213FB42FE2EC6F4" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[294,506,1157,1181]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="238" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951F427C7213FB42FE2EC6F4" box="[294,506,1157,1181]" italics="true" pageId="17" pageNumber="238">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951F427C7135FB42FDEEC6F4" bold="true" box="[512,570,1157,1181]" pageId="17" pageNumber="238">
|
||
<taxonomicNameLabel id="B2FB57A6951F427C7135FB42FDEEC6F4" box="[512,570,1157,1181]" pageId="17" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, female of hard stage (5.2×5.2 mm), CBM-ZC 15956. A, right chela, outer view; B, right P2, dactylus to carpus, dorsal view; C, right P3, dactylus to carpus, dorsal view; D, right P4, dactylus to carpus (shorter side), dorsal view; E, left P4, dactylus to carpus (longer side), dorsal view; F, right P5, dactylus to carpus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B0336CF951F427C73F2FAD4FA99C7F0" blockId="17.[151,1437,1299,2010]" pageId="17" pageNumber="238">
|
||
Pleon (
|
||
<figureCitation id="03872A4A951F427C7222FAD4FEA9C747" box="[279,381,1299,1326]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="17" pageNumber="238">Fig. 13H</figureCitation>
|
||
) moderately narrow, tapering distally to pleomere 6; lateral margins of pleomeres 3 generally rounded, those of sternite 4 slightly concave. Pleomeres 1 and 2 subequal in width, longitudinally very short. Pleomere 3 widely subtrapezoidal. Pleomeres 4–6 all subtrapezoidal. Telson subhexagonal, 3.6 times wider than long, maximal width greater than distal width of pleomere 6; distal margin nearly straight, lateral margin angular.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951F427C73F2FA63FAA7C440" blockId="17.[151,1437,1299,2010]" pageId="17" pageNumber="238">
|
||
G1 (
|
||
<figureCitation id="03872A4A951F427C73CFFA64FEA5C7D7" box="[250,369,1443,1470]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="17" pageNumber="238">Fig. 13I, J</figureCitation>
|
||
) noticeably arcuate outward and slightly twisted, dorsoventrally compressed, gradually tapering to subacutely pointed apex, reaching to anterolateral end of sterno-pleonal cavity; ventral surface with irregular rows of setae extending to lateral margin. G2 (
|
||
<figureCitation id="03872A4A951F427C71ABFA2CFCD6C46F" box="[670,770,1515,1542]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="17" pageNumber="238">Fig. 13K</figureCitation>
|
||
) biramous; endopod narrowing distal to midlength; exopod well developed, flattened, evenly tapering to rounded apex, slightly longer than endopod. Pleopods 3–5 absent.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951F427C73F2F9F4FCE1C4D0" blockId="17.[151,1437,1299,2010]" pageId="17" pageNumber="238">
|
||
<emphasis id="A9C8EADD951F427C73F2F9F4FE60C424" box="[199,436,1587,1613]" italics="true" pageId="17" pageNumber="238">Male post-hard stage</emphasis>
|
||
. Carapace (
|
||
<figureCitation id="03872A4A951F427C710DF9F4FD48C427" box="[568,668,1587,1614]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="17" pageNumber="238">Fig. 15A</figureCitation>
|
||
) well chitinised but less calcified than in hard stage, rounded, subcircular, 1.1 times as long as wide; dorsal surface convex, without demarcation of regions, glabrous (no minute pits); front weakly projecting anteriorly beyond orbits, margin broadly rounded; lateral angle of front produced into small triangular process directed downward, but not closing orbit.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951F427C73F2F903FBFCC4B4" blockId="17.[151,1437,1299,2010]" box="[199,1064,1731,1758]" pageId="17" pageNumber="238">
|
||
Eyes (
|
||
<figureCitation id="03872A4A951F427C7238F904FEA6C4B7" box="[269,370,1731,1758]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="17" pageNumber="238">Fig. 15A</figureCitation>
|
||
) small, clearly visible in dorsal view, completely filling orbit.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951F427C73F2F920FDA9C54C" blockId="17.[151,1437,1299,2010]" pageId="17" pageNumber="238">Antennular peduncles folded obliquely into antennular fossae. Antenna extending along lateral margin of front; article 2 subtrapezoidal, narrowing distally.</paragraph>
|
||
<paragraph id="9B0336CF951F427C73F2F8E8FCEEC520" blockId="17.[151,1437,1299,2010]" box="[199,826,1839,1866]" pageId="17" pageNumber="238">
|
||
Maxilliped 3 (
|
||
<figureCitation id="03872A4A951F427C7253F8E8FE18C523" box="[358,460,1839,1866]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="17" pageNumber="238">Fig. 16A</figureCitation>
|
||
) similar to that of adult females.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951F427F73F2F894FB7DC374" blockId="17.[151,1437,1299,2010]" lastBlockId="18.[151,1437,151,1041]" lastPageId="18" lastPageNumber="239" pageId="17" pageNumber="238">
|
||
Chelipeds (
|
||
<figureCitation id="03872A4A951F427C727FF894FE60C507" box="[330,436,1875,1902]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="17" pageNumber="238">Fig. 15A</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A951F427C72F6F894FE26C507" box="[451,498,1875,1902]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="17" pageNumber="238">16B</figureCitation>
|
||
) rather small, symmetrical. Dactylus 0.6 times as long as palm, strongly curved, terminating in hooked, acute tip; upper margin rounded; occlusal margin with 1 conspicuous tooth proximal to midlength and with row of sparse short setae. Palm slightly widened distally, 1.6 times as long as high; outer surface glabrous; inner surface almost glabrous, but with longitudinal row of short setae adjacent to lower margin, extending onto fixed finger, becoming multiple rows. Fixed finger terminating in hooked, acute tip; occlusal margin with 2 small teeth proximal to midlength, flanking occlusal tooth on dactylus, and with row of short setae distal to proximal teeth. Carpus distinctly shorter than palm, widened distally, glabrous. Merus slightly curved inwards, slightly compressed laterally; outer surface convex, glabrous, lower distal angle somewhat produced.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951C427F73F2FEE0FAD2C098" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="239">
|
||
Ambulatory legs (
|
||
<figureCitation id="03872A4A951C427F72A5FEE0FE21C328" box="[400,501,294,321]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Fig. 15A</figureCitation>
|
||
) relatively shorter and slender compared with those of hard stage males, P3 longest; no natatory setae on every pereopod; P2, P3 and P5 symmetrical, P4 with asymmetrical dactyli. P2 (
|
||
<figureCitation id="03872A4A951C427F7616FE8CFA43C30C" box="[1315,1431,330,357]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Figs. 15A</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A951C427F73A2FEA8FF1DC3E3" box="[151,201,367,394]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="18" pageNumber="239">16D</figureCitation>
|
||
) entirely glabrous; dactylus curved, half-length of propodus; carpus curved proximally; merus almost straight, slightly widened distally. P3 (
|
||
<figureCitation id="03872A4A951C427F72D6FE54FD81C3C4" box="[483,597,402,429]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Figs. 15A</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A951C427F7155FE54FD5BC3C7" box="[608,655,403,430]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="18" pageNumber="239">16E</figureCitation>
|
||
) generally similar to P2 except for length; dactylus about 0.6 times as long as propodus; carpus lower surface with scattered short setae, otherwise glabrous. P4 (
|
||
<figureCitation id="03872A4A951C427F77A5FE70FAD6C3B8" box="[1168,1282,438,465]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Figs. 15A</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A951C427F763BFE70FA8FC3B8" box="[1294,1371,439,465]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="18" pageNumber="239">16F, G</figureCitation>
|
||
) with left dactylus slightly longer than right dactylus (left 1.1 times as long as right), 0.9 times as long as propodus, extensor margin glabrous, flexor margin with 2 long subterminal setae followed by row of sparse short setae; shorter right dactylus 0.7 times as long as propodus, flexor margin with 1 long subterminal seta followed by fewer short setae; propodus to merus glabrous; carpus longest among ambulatory legs; merus nearly straight. P5 (
|
||
<figureCitation id="03872A4A951C427F77D4FD80FA81C008" box="[1249,1365,582,609]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Figs. 15A</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A951C427F7656FD80FA40C00B" box="[1379,1428,583,610]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="18" pageNumber="239">16H</figureCitation>
|
||
) shortest; dactylus slightly arcuate, subequal in length to propodus, approximately 9 times as long as wide; extensor margin with irregular rows of short setae on distal half, lower margin with row of setae becoming longer distally; propodus almost glabrous except for sparse short setae on lower margin; carpus glabrous, gently curved; merus subcylindrical, straight, also glabrous; ischium slightly wider than merus, about half-length of merus.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951C427F73F2FD3CFDAAC678" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="239">
|
||
Thoracic sternum broad. Sternites 1–4 (Fig. 153B) fused, narrowing anteriorly; narrow incision at base of maxilliped 3, suggesting boundary between sternites 3 and 4, still discernible; broad anteromedian concavity well delimited; anterolateral angles produced in rounded lobe; surface almost flat. Exposed parts of sternites 5–7 (
|
||
<figureCitation id="03872A4A951C427F765AFC84FF13C1E8" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Fig. 15C</figureCitation>
|
||
) nearly smooth, each posteriorly with small but conspicuous, rounded episternites. Sternite 8 (
|
||
<figureCitation id="03872A4A951C427F77E8FCA0FA96C1E8" box="[1245,1346,870,897]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Fig. 15D</figureCitation>
|
||
) clearly visible in lateral part, lateral margin deeply notched. Sterno-pleonal cavity (
|
||
<figureCitation id="03872A4A951C427F70DFFC4CFB99C1CC" box="[1002,1101,906,933]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="18" pageNumber="239">Fig. 15E</figureCitation>
|
||
) deep, defined by sharply delimited margin, widened anteriorly (corresponding to telson), reaching anteriorly to level of anterior ends of cheliped coxae; pleon locking apparatus consisting of 2 press buttons on either side on sternite 5 inside of cavity; sternite 6 with trace of low elevation on either side.
|
||
</paragraph>
|
||
<caption id="CFC36647951C427F73A2F810FA53C586" ID-DOI="http://doi.org/10.5281/zenodo.4401487" ID-Zenodo-Dep="4401487" box="[151,1415,2007,2031]" httpUri="https://zenodo.org/record/4401487/files/figure.png" pageId="18" pageNumber="239" startId="18.[151,250,2007,2031]" targetBox="[159,1427,1065,1983]" targetPageId="18">
|
||
<paragraph id="9B0336CF951C427F73A2F810FA53C586" blockId="18.[151,1415,2007,2031]" box="[151,1415,2007,2031]" pageId="18" pageNumber="239">
|
||
<emphasis id="A9C8EADD951C427F73A2F810FEF4C586" bold="true" box="[151,288,2007,2031]" pageId="18" pageNumber="239">FIGURE 12.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951C427F7213F810FE2EC586" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[294,506,2007,2031]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="239" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951C427F7213F810FE2EC586" box="[294,506,2007,2031]" italics="true" pageId="18" pageNumber="239">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951C427F7135F810FDEDC586" bold="true" box="[512,569,2007,2031]" pageId="18" pageNumber="239">
|
||
<taxonomicNameLabel id="B2FB57A6951C427F7135F810FDEDC586" box="[512,569,2007,2031]" pageId="18" pageNumber="239" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957, habitus, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC36647951D427E73A2F8FDFBFEC58B" ID-DOI="http://doi.org/10.5281/zenodo.4401489" ID-Zenodo-Dep="4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="19" pageNumber="240" startId="19.[151,250,1850,1874]" targetBox="[161,1425,191,1815]" targetPageId="19">
|
||
<paragraph id="9B0336CF951D427E73A2F8FDFBFEC58B" blockId="19.[151,1436,1850,2018]" pageId="19" pageNumber="240">
|
||
<emphasis id="A9C8EADD951D427E73A2F8FDFEF4C53B" bold="true" box="[151,288,1850,1874]" pageId="19" pageNumber="240">FIGURE 13.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951D427E7212F8FDFE2FC53B" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[295,507,1850,1874]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="240" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951D427E7212F8FDFE2FC53B" box="[295,507,1850,1874]" italics="true" pageId="19" pageNumber="240">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951D427E7137F8FDFDEFC53B" bold="true" box="[514,571,1850,1874]" pageId="19" pageNumber="240">
|
||
<taxonomicNameLabel id="B2FB57A6951D427E7137F8FDFDEFC53B" box="[514,571,1850,1874]" pageId="19" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC36647951A427973A2FBB8FB72C76A" ID-DOI="http://doi.org/10.5281/zenodo.4401491" ID-Zenodo-Dep="4401491" httpUri="https://zenodo.org/record/4401491/files/figure.png" pageId="20" pageNumber="241" startId="20.[151,250,1151,1175]" targetBox="[157,1428,188,1117]" targetPageId="20">
|
||
<paragraph id="9B0336CF951A427973A2FBB8FB72C76A" blockId="20.[151,1437,1151,1283]" pageId="20" pageNumber="241">
|
||
<emphasis id="A9C8EADD951A427973A2FBB8FEF4C6FE" bold="true" box="[151,288,1151,1175]" pageId="20" pageNumber="241">FIGURE 14.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951A42797213FBB8FE2EC6FE" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[294,506,1151,1175]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951A42797213FBB8FE2EC6FE" box="[294,506,1151,1175]" italics="true" pageId="20" pageNumber="241">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951A42797134FBB8FDEEC6FE" bold="true" box="[513,570,1151,1175]" pageId="20" pageNumber="241">
|
||
<taxonomicNameLabel id="B2FB57A6951A42797134FBB8FDEEC6FE" box="[513,570,1151,1175]" pageId="20" pageNumber="241" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, chela and carpus of left cheliped, outer view; B, same, dorsal view; C, right P2, dactylus to distal part of carpus, dorsal view; D, right P3, dactylus to distal part of carpus, dorsal view; E, right P4 (longer side), dactylus to distal part of carpus, dorsal view; F, left P4 (shorter side), dactylus to distal part of carpus, dorsal view; G, right P5, dactylus to distal part of carpus, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B0336CF951A427973F2FAF6FBF7C7DE" blockId="20.[151,1437,1329,2040]" pageId="20" pageNumber="241">
|
||
Pleon (
|
||
<figureCitation id="03872A4A951A4279722CFAF6FEA8C725" box="[281,380,1329,1356]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="20" pageNumber="241">Fig. 15F</figureCitation>
|
||
) moderately narrow, tapering distally to pleomere 6. Pleomeres 1 and 2 subequal in width, longitudinally short. Pleomere 3 widely subtrapezoidal; lateral margins convex. Pleomeres 4–6 all subtrapezoidal; lateral margins of pleomere 4 concave. Telson (
|
||
<figureCitation id="03872A4A951A4279718FFABEFCC9C7FD" box="[698,797,1401,1428]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="20" pageNumber="241">Fig. 15F</figureCitation>
|
||
) pentagonal, 1.8 times wider than long, maximal width greater than distal width of pleomere 6, distal margin slightly produced medially.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951A427973F2FA05FB4DC42E" blockId="20.[151,1437,1329,2040]" pageId="20" pageNumber="241">
|
||
G1 (
|
||
<figureCitation id="03872A4A951A427973CCFA06FE54C7B5" box="[249,384,1473,1500]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="20" pageNumber="241">Fig. 15G, H</figureCitation>
|
||
) noticeably arcuate outward and slightly twisted, dorsoventrally compressed, gradually tapering to bluntly pointed apex; ventral surface with irregular rows of short setae adjacent to lateral margin. G2 (
|
||
<figureCitation id="03872A4A951A4279765AFA22FF69C44D" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="20" pageNumber="241">Fig. 15I</figureCitation>
|
||
) endopod abruptly tapering at midlength to blunt apex, about 0.2 length of G1; exopod well developed, slender, flattened, evenly tapering to blunt apex, slightly longer than endopod. Pleopods 3–5 absent.
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951A427973F2F996FADFC4F9" blockId="20.[151,1437,1329,2040]" pageId="20" pageNumber="241">
|
||
<emphasis id="A9C8EADD951A427973F2F996FEEBC402" bold="true" box="[199,319,1617,1643]" pageId="20" pageNumber="241">Variation.</emphasis>
|
||
Carapace shape is not a stable character; even in the series of specimens of
|
||
<taxonomicName id="5CBC4D4C951A427977ADF995FAF5C402" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[1176,1313,1617,1644]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="ocularius">
|
||
<emphasis id="A9C8EADD951A427977ADF995FAF5C402" box="[1176,1313,1617,1644]" italics="true" pageId="20" pageNumber="241">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
examined, the carapace of ovigerous females varies from broadly ovate (
|
||
<figureCitation id="03872A4A951A4279707EF9B2FC77C4F9" box="[843,931,1653,1680]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="20" pageNumber="241">Fig. 7A</figureCitation>
|
||
) to almost circular (
|
||
<figureCitation id="03872A4A951A427977B0F9B2FB2AC4F9" box="[1157,1278,1653,1680]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="20" pageNumber="241">Fig. 7B, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9B0336CF951A427973F2F95EFF17C4B1" blockId="20.[151,1437,1329,2040]" pageId="20" pageNumber="241">
|
||
<emphasis id="A9C8EADD951A427973F2F95EFEA7C4DA" bold="true" box="[199,371,1689,1715]" pageId="20" pageNumber="241">Colour in life.</emphasis>
|
||
Entirely whitish; female ovary, visible through integument, orange; cornea of eye red (
|
||
<figureCitation id="03872A4A951A4279765AF95EFF62C4B1" captionStart="FIGURE 17" captionStartId="23.[151,250,1620,1644]" captionTargetBox="[151,1435,633,1597]" captionTargetId="figure-260@23.[151,1435,633,1597]" captionTargetPageId="23" captionText="FIGURE 17. Arcotheres ocularius n. sp., female of post-hard stage V, not specified, living animal associated with Anadara sp." figureDoi="http://doi.org/10.5281/zenodo.4401499" httpUri="https://zenodo.org/record/4401499/files/figure.png" pageId="20" pageNumber="241">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A66544951A427973F2F926FF00C549" pageId="20" pageNumber="241" type="distribution">
|
||
<paragraph id="9B0336CF951A427973F2F926FF00C549" blockId="20.[151,1437,1329,2040]" pageId="20" pageNumber="241">
|
||
<emphasis id="A9C8EADD951A427973F2F926FEB5C492" bold="true" box="[199,353,1761,1787]" pageId="20" pageNumber="241">Distribution.</emphasis>
|
||
<collectingCountry id="E3AB765F951A4279725DF926FE5BC492" box="[360,399,1761,1787]" name="Fiji" pageId="20" pageNumber="241">Fiji</collectingCountry>
|
||
and
|
||
<collectingCountry id="E3AB765F951A427972FDF926FDE1C492" box="[456,565,1761,1787]" name="Indonesia" pageId="20" pageNumber="241">Indonesia</collectingCountry>
|
||
(Lombok); intertidal; exclusively associated with of
|
||
<taxonomicName id="5CBC4D4C951A427977BBF926FB24C492" box="[1166,1264,1761,1787]" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcoida" pageId="20" pageNumber="241" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A9C8EADD951A427977BBF926FB24C492" box="[1166,1264,1761,1787]" italics="true" pageId="20" pageNumber="241">Anadara</emphasis>
|
||
</taxonomicName>
|
||
spp. (unidentified).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A66544951A427A73F2F8EDFD33C054" lastPageId="23" lastPageNumber="244" pageId="20" pageNumber="241" type="discussion">
|
||
<paragraph id="9B0336CF951A427B73F2F8EDFE04C2BF" blockId="20.[151,1437,1329,2040]" lastBlockId="22.[151,1437,151,825]" lastPageId="22" lastPageNumber="243" pageId="20" pageNumber="241">
|
||
<emphasis id="A9C8EADD951A427973F2F8EDFEEFC52D" bold="true" box="[199,315,1834,1860]" pageId="20" pageNumber="241">Remarks.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951A42797270F8EEFDE5C52A" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[325,561,1833,1859]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951A42797270F8EEFDE5C52A" box="[325,561,1833,1859]" italics="true" pageId="20" pageNumber="241">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951A4279710FF8EDFDAFC52D" bold="true" box="[570,635,1834,1860]" pageId="20" pageNumber="241">
|
||
<taxonomicNameLabel id="B2FB57A6951A4279710FF8EDFDAFC52D" box="[570,635,1834,1860]" pageId="20" pageNumber="241" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="5CBC4D4C951A4279706DF8EDFC25C52A" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[856,1009,1833,1860]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="arcophilus">
|
||
<emphasis id="A9C8EADD951A4279706DF8EDFC25C52A" box="[856,1009,1833,1860]" italics="true" pageId="20" pageNumber="241">A. arcophilus</emphasis>
|
||
</taxonomicName>
|
||
and a group of four nominal species closely allied to
|
||
<emphasis id="A9C8EADD951A42797278F889FE09C50E" box="[333,477,1869,1896]" italics="true" pageId="20" pageNumber="241">
|
||
<taxonomicName id="5CBC4D4C951A42797278F889FE0DC50E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[333,473,1869,1896]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="palaensis">A. palaensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
This group includes
|
||
<taxonomicName id="5CBC4D4C951A427971F2F889FBD6C501" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[711,1026,1869,1896]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="rhombifer">
|
||
<emphasis id="A9C8EADD951A427971F2F889FC81C50E" box="[711,853,1869,1896]" italics="true" pageId="20" pageNumber="241">A. rhombifer</emphasis>
|
||
(
|
||
<bibRefCitation id="FF2D4B3E951A42797056F889FC2EC501" author="Burger, O." box="[867,1018,1869,1896]" pageId="20" pageNumber="241" pagination="361 - 390" refId="ref12768" refString="Burger, O. (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5CBC4D4C951A42797738F889FAEDC501" authority="Campos, 2001" authorityName="Campos" authorityYear="2001" box="[1037,1337,1870,1896]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="guinotae">
|
||
<emphasis id="A9C8EADD951A42797738F889FB5AC50E" box="[1037,1166,1870,1896]" italics="true" pageId="20" pageNumber="241">A. guinotae</emphasis>
|
||
Campos, 2001
|
||
</taxonomicName>
|
||
, and two species currently in
|
||
<taxonomicName id="5CBC4D4C951A4279724BF8B6FDD0C5E2" authorityName="Bosc" authorityYear="1802" box="[382,516,1905,1931]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A9C8EADD951A4279724BF8B6FDD0C5E2" box="[382,516,1905,1931]" italics="true" pageId="20" pageNumber="241">Pinnotheres</emphasis>
|
||
</taxonomicName>
|
||
but clearly also referable to
|
||
<emphasis id="A9C8EADD951A42797064F8B6FC05C5E2" box="[849,977,1905,1931]" italics="true" pageId="20" pageNumber="241">
|
||
<taxonomicName id="5CBC4D4C951A42797064F8B6FC19C5E2" authorityName="Manning" authorityYear="1993" box="[849,973,1905,1931]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="genus">Arcotheres</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="5CBC4D4C951A427970EFF8B5FA9AC5E5" authority="(Tesch, 1918)" authorityName="Komai & Kawai & Ng" authorityYear="2020" baseAuthorityName="Tesch" baseAuthorityYear="1918" box="[986,1358,1905,1932]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="onychodactylus" status="comb. nov.">
|
||
<emphasis id="A9C8EADD951A427970EFF8B5FB7DC5E2" box="[986,1193,1905,1932]" italics="true" pageId="20" pageNumber="241">A. onychodactylus</emphasis>
|
||
(
|
||
<bibRefCitation id="FF2D4B3E951A4279778FF8B6FA92C5E5" author="Tesch, J. J." box="[1210,1350,1905,1932]" pageId="20" pageNumber="241" pagination="149 - 295" refId="ref15162" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2FB57A6951A42797662F8B6FF13C5D9" pageId="20" pageNumber="241" rank="species">comb. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="5CBC4D4C951A42797235F851FD67C5D9" authority="(Dai & Song, 1986)" authorityName="Komai & Kawai & Ng" authorityYear="2020" baseAuthorityName="Dai & Song" baseAuthorityYear="1986" box="[256,691,1941,1968]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="paralatissimus" status="comb. nov.">
|
||
<emphasis id="A9C8EADD951A42797235F851FE12C5C6" box="[256,454,1941,1968]" italics="true" pageId="20" pageNumber="241">A. paralatissimus</emphasis>
|
||
(
|
||
<bibRefCitation id="FF2D4B3E951A427972E2F852FD7FC5D9" author="Dai, A. - Y. & Song, Y. - Z." box="[471,683,1941,1968]" pageId="20" pageNumber="241" pagination="54 - 62" refId="ref13037" refString="Dai, A. - Y. & Song, Y. - Z. (1986) Intertidal crabs from Beibu Gulf of Guangxi. Transactions of the Chinese Crustacean Society, 54 - 62, pl. 1." type="book chapter" year="1986">Dai & Song, 1986</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2FB57A6951A42797189F852FCEEC5D9" box="[700,826,1941,1968]" pageId="20" pageNumber="241" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(PKLN & S.T. Ahyong, unpublished data). All differ from
|
||
<taxonomicName id="5CBC4D4C951A427973E1F87DFE89C5BA" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[212,349,1977,2004]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="ocularius">
|
||
<emphasis id="A9C8EADD951A427973E1F87DFE89C5BA" box="[212,349,1977,2004]" italics="true" pageId="20" pageNumber="241">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
in the same features as discussed for
|
||
<taxonomicName id="5CBC4D4C951A427971CAF87DFC5CC5BA" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[767,904,1977,2004]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD951A427971CAF87DFC5CC5BA" box="[767,904,1977,2004]" italics="true" pageId="20" pageNumber="241">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
, below. In addition, Japanese specimens identified as “
|
||
<taxonomicName id="5CBC4D4C951A427973CDF81AFE1FC59E" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[248,459,2013,2039]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="alcocki">
|
||
<emphasis id="A9C8EADD951A427973CDF81AFE1FC59E" box="[248,459,2013,2039]" italics="true" pageId="20" pageNumber="241">Arcotheres alcocki</emphasis>
|
||
</taxonomicName>
|
||
” by
|
||
<bibRefCitation id="FF2D4B3E951A42797130F81AFCFDC591" author="Takeda, M. & Konishi, K." box="[517,809,2013,2040]" pageId="20" pageNumber="241" pagination="137 - 144" refId="ref15115" refString="Takeda, M. & Konishi, K. (1988) Redescription of Pinnotheres alcocki Rathbun, 1909, a commensal pea crab new to Japan. Bulletin of the National Science Museum, Series A, 14 (3), 137 - 144." type="journal article" year="1988">Takeda & Konishi (1988)</bibRefCitation>
|
||
actually represent
|
||
<taxonomicName id="5CBC4D4C951A42797736F819FB5AC59E" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1027,1166,2013,2040]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="241" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD951A42797736F819FB5AC59E" box="[1027,1166,2013,2040]" italics="true" pageId="20" pageNumber="241">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
after re-examination of the material (NSMT-Cr 9363). Additional specimens of this taxon from
|
||
<collectingCountry id="E3AB765F9518427B70F1FF50FBD0C2D8" box="[964,1028,151,177]" name="Japan" pageId="22" pageNumber="243">Japan</collectingCountry>
|
||
were also obtained for comparisons (see Material and methods).
|
||
</paragraph>
|
||
<caption id="CFC36647951B427873A2F889FDCAC59E" ID-DOI="http://doi.org/10.5281/zenodo.4401493" ID-Zenodo-Dep="4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="21" pageNumber="242" startId="21.[151,250,1870,1894]" targetBox="[175,1414,188,1838]" targetPageId="21">
|
||
<paragraph id="9B0336CF951B427873A2F889FDCAC59E" blockId="21.[151,1436,1870,2039]" pageId="21" pageNumber="242">
|
||
<emphasis id="A9C8EADD951B427873A2F889FEF4C50E" bold="true" box="[151,288,1870,1895]" pageId="21" pageNumber="242">FIGURE 15.</emphasis>
|
||
<taxonomicName id="5CBC4D4C951B42787213F888FE2EC50E" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[294,506,1871,1895]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="242" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD951B42787213F888FE2EC50E" box="[294,506,1871,1895]" italics="true" pageId="21" pageNumber="242">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD951B42787135F888FDEDC50E" bold="true" box="[512,569,1871,1895]" pageId="21" pageNumber="242">
|
||
<taxonomicNameLabel id="B2FB57A6951B42787135F888FDEDC50E" box="[512,569,1871,1895]" pageId="21" pageNumber="242" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B0336CF9518427B73F2FF18FE20C150" blockId="22.[151,1437,151,825]" pageId="22" pageNumber="243">
|
||
In
|
||
<taxonomicName id="5CBC4D4C9518427B73D0FF18FEBAC290" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[229,366,223,249]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B73D0FF18FEBAC290" box="[229,366,223,249]" italics="true" pageId="22" pageNumber="243">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B7241FF18FE67C290" bold="true" box="[372,435,223,249]" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B7241FF18FE67C290" box="[372,435,223,249]" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the carapace is distinctly more strongly chitinised and calcified, with the cuticle stiff and relatively rigid (versus cuticle distinctly less chitinised and softer, even in fresh specimens of
|
||
<taxonomicName id="5CBC4D4C9518427B7796FEC4FAFFC374" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1187,1323,259,285]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B7796FEC4FAFFC374" box="[1187,1323,259,285]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
); eyes are clearly visible in the dorsal view in
|
||
<taxonomicName id="5CBC4D4C9518427B7110FEE0FD7AC328" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[549,686,295,321]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B7110FEE0FD7AC328" box="[549,686,295,321]" italics="true" pageId="22" pageNumber="243">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B7181FEE0FD27C328" bold="true" box="[692,755,295,321]" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B7181FEE0FD27C328" box="[692,755,295,321]" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="03872A4A9518427B7037FEE0FC97C328" box="[770,835,295,321]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure-17@13.[163,1425,181,1919]" captionTargetPageId="13" captionText="FIGURE 7. Arcotheres ocularius n. sp. A, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952, Fiji; B–E, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji.A, B, overall habitus in dorsal view; C, carapace, dorsal view; D, frontal view of cephalothorax; E, right chela, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401477" httpUri="https://zenodo.org/record/4401477/files/figure.png" pageId="22" pageNumber="243">Fig. 7</figureCitation>
|
||
A–C, 8A) (versus eyes are not visible or occasionally, only the edge of the eye is visible in
|
||
<taxonomicName id="5CBC4D4C9518427B711AFE8CFD6CC30C" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[559,696,331,357]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B711AFE8CFD6CC30C" box="[559,696,331,357]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="FF2D4B3E9518427B71F1FE8CFC72C30C" author="Ahyong, S. T. & Ng, P. K. L." box="[708,934,331,357]" pageId="22" pageNumber="243" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
|
||
: fig. 9A; PKLN & S.T. Ahyong, unpublished data); the maxilliped 3 dactylus reaches to or extends beyond the tip of the propodus in
|
||
<taxonomicName id="5CBC4D4C9518427B7751FEA8FB39C3E0" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[1124,1261,367,393]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B7751FEA8FB39C3E0" box="[1124,1261,367,393]" italics="true" pageId="22" pageNumber="243">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B77C6FEA8FAE6C3E0" bold="true" box="[1267,1330,367,393]" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B77C6FEA8FAE6C3E0" box="[1267,1330,367,393]" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="03872A4A9518427B7674FEA8FF1CC3C4" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="22" pageNumber="243">Fig. 8C, F, G</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9518427B73E7FE54FED7C3C7" box="[210,259,403,430]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="22" pageNumber="243">16A</figureCitation>
|
||
) (versus the maxilliped 3 dactylus is always shorter, never reaching the tip of the propodus in
|
||
<taxonomicName id="5CBC4D4C9518427B7638FE54FA41C3C4" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1293,1429,403,429]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B7638FE54FA41C3C4" box="[1293,1429,403,429]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="FF2D4B3E9518427B738EFE70FE75C3B8" author="Ahyong, S. T. & Ng, P. K. L." box="[187,417,439,465]" pageId="22" pageNumber="243" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
|
||
: fig. 9C); the female P4 merus and propodus are proportionately longer in
|
||
<taxonomicName id="5CBC4D4C9518427B77C1FE70FAAAC3B8" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[1268,1406,439,465]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B77C1FE70FAAAC3B8" box="[1268,1406,439,465]" italics="true" pageId="22" pageNumber="243">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B76B3FE70FF6CC39C" bold="true" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B76B3FE70FF6CC39C" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="03872A4A9518427B73FCFE1CFECAC39C" box="[201,286,475,501]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="22" pageNumber="243">Fig. 9L</figureCitation>
|
||
) (versus proportionately shorter in
|
||
<taxonomicName id="5CBC4D4C9518427B719BFE1CFCECC39C" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[686,824,475,501]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B719BFE1CFCECC39C" box="[686,824,475,501]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="FF2D4B3E9518427B7070FE1CFBFFC39C" author="Ahyong, S. T. & Ng, P. K. L." box="[837,1067,475,501]" pageId="22" pageNumber="243" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
|
||
: fig. 9A); the male telson is subpentagonal or subhexagonal in
|
||
<taxonomicName id="5CBC4D4C9518427B72C0FE38FDAAC070" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[501,638,511,537]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B72C0FE38FDAAC070" box="[501,638,511,537]" italics="true" pageId="22" pageNumber="243">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B71B1FE38FD17C070" bold="true" box="[644,707,511,537]" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B71B1FE38FD17C070" box="[644,707,511,537]" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="03872A4A9518427B71E7FE38FC90C070" box="[722,836,511,537]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="22" pageNumber="243">Figs. 13H</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9518427B7065FE39FCA8C070" box="[848,892,510,537]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="22" pageNumber="243">15F</figureCitation>
|
||
) (versus semicircular in
|
||
<taxonomicName id="5CBC4D4C9518427B77A4FE38FACEC070" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1169,1306,511,537]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B77A4FE38FACEC070" box="[1169,1306,511,537]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="FF2D4B3E9518427B7679FE38FE85C054" author="Takeda, M. & Konishi, K." pageId="22" pageNumber="243" pagination="137 - 144" refId="ref15115" refString="Takeda, M. & Konishi, K. (1988) Redescription of Pinnotheres alcocki Rathbun, 1909, a commensal pea crab new to Japan. Bulletin of the National Science Museum, Series A, 14 (3), 137 - 144." type="journal article" year="1988">Takeda & Konishi 1988</bibRefCitation>
|
||
: fig. 3C, as
|
||
<taxonomicName id="5CBC4D4C9518427B72EDFDE4FD91C054" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[472,581,547,573]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="alcocki">
|
||
<emphasis id="A9C8EADD9518427B72EDFDE4FD91C054" box="[472,581,547,573]" italics="true" pageId="22" pageNumber="243">P. alcocki</emphasis>
|
||
</taxonomicName>
|
||
); and the G1 has the tip subtruncate (
|
||
<figureCitation id="03872A4A9518427B70D8FDE4FBBAC054" box="[1005,1134,547,573]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="22" pageNumber="243">Figs. 13I, J</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9518427B774FFDE5FB1AC054" box="[1146,1230,546,573]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="22" pageNumber="243">15G, H</figureCitation>
|
||
) (versus the tip of the G1 has an elongated slender projection in
|
||
<taxonomicName id="5CBC4D4C9518427B7197FD80FCF8C008" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[674,812,583,609]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B7197FD80FCF8C008" box="[674,812,583,609]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="FF2D4B3E9518427B7055FD80FBA7C008" author="Takeda, M. & Konishi, K." box="[864,1139,583,609]" pageId="22" pageNumber="243" pagination="137 - 144" refId="ref15115" refString="Takeda, M. & Konishi, K. (1988) Redescription of Pinnotheres alcocki Rathbun, 1909, a commensal pea crab new to Japan. Bulletin of the National Science Museum, Series A, 14 (3), 137 - 144." type="journal article" year="1988">Takeda & Konishi 1988</bibRefCitation>
|
||
: fig. 3D, E, as
|
||
<taxonomicName id="5CBC4D4C9518427B762BFD80FA58C008" baseAuthorityName="Rathbun" baseAuthorityYear="1909" box="[1310,1420,583,609]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="alcocki">
|
||
<emphasis id="A9C8EADD9518427B762BFD80FA58C008" box="[1310,1420,583,609]" italics="true" pageId="22" pageNumber="243">P. alcocki</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="5CBC4D4C9518427B73A2FDACFE54C0EC" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[151,384,619,645]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B73A2FDACFE54C0EC" box="[151,384,619,645]" italics="true" pageId="22" pageNumber="243">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B72B2FDACFE11C0EC" bold="true" box="[391,453,619,645]" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B72B2FDACFE11C0EC" box="[391,453,619,645]" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="5CBC4D4C9518427B72CEFDACFD57C0EC" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[507,643,619,645]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B72CEFDACFD57C0EC" box="[507,643,619,645]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
are both associated with
|
||
<taxonomicName id="5CBC4D4C9518427B70A8FDACFC2BC0EC" box="[925,1023,619,645]" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcoida" pageId="22" pageNumber="243" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A9C8EADD9518427B70A8FDACFC2BC0EC" box="[925,1023,619,645]" italics="true" pageId="22" pageNumber="243">Anadara</emphasis>
|
||
</taxonomicName>
|
||
spp., but
|
||
<taxonomicName id="5CBC4D4C9518427B775EFDACFB20C0EC" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1131,1268,619,645]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B775EFDACFB20C0EC" box="[1131,1268,619,645]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
is also reliably known from
|
||
<taxonomicName id="5CBC4D4C9518427B7219FD48FE44C0C0" box="[300,400,655,681]" class="Bivalvia" family="Arcidae" genus="Barbatia" kingdom="Animalia" order="Arcoida" pageId="22" pageNumber="243" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A9C8EADD9518427B7219FD48FE44C0C0" box="[300,400,655,681]" italics="true" pageId="22" pageNumber="243">Barbatia</emphasis>
|
||
</taxonomicName>
|
||
species (also
|
||
<taxonomicName id="5CBC4D4C9518427B711AFD48FD5EC0C3" authorityName="Lamarck" authorityYear="1809" box="[559,650,655,682]" class="Bivalvia" family="Arcidae" kingdom="Animalia" order="Arcoida" pageId="22" pageNumber="243" phylum="Mollusca" rank="family">Arcidae</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="FF2D4B3E9518427B7194FD48FC6CC0C0" author="Takeda, M. & Konishi, K." box="[673,952,655,681]" pageId="22" pageNumber="243" pagination="137 - 144" refId="ref15115" refString="Takeda, M. & Konishi, K. (1988) Redescription of Pinnotheres alcocki Rathbun, 1909, a commensal pea crab new to Japan. Bulletin of the National Science Museum, Series A, 14 (3), 137 - 144." type="journal article" year="1988">Takeda & Konishi 1988</bibRefCitation>
|
||
; Campos 2001;
|
||
<bibRefCitation id="FF2D4B3E9518427B7747FD48FA43C0C0" author="Watanabe, T. & Henmi, Y." box="[1138,1431,655,681]" pageId="22" pageNumber="243" pagination="217 - 223" refId="ref15379" refString="Watanabe, T. & Henmi, Y. (2008) Morphological development of the commensal pea crab (Arcotheres sp.) in the laboratory reared specimens. Journal of Crustacean Biology, 29 (2), 217 - 223. https: // doi. org / 10.1651 / 08 - 3071 R. 1" type="journal article" year="2008">Watanabe & Henmi 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="FF2D4B3E9518427B73A2FD74FE92C0A4" author="Watanabe, T." box="[151,326,691,717]" pageId="22" pageNumber="243" pagination="45 - 50" refId="ref15351" refString="Watanabe, T. (2013) Host specificity and life history of pinnotherid crabs. Cancer, 22, 45 - 50. [in Japanese]" type="journal article" year="2013">Watanabe 2013</bibRefCitation>
|
||
; PKLN & S.T. Ahyong, unpublished data).
|
||
<bibRefCitation id="FF2D4B3E9518427B7019FD74FBC6C0A4" author="Schmitt, W. L. & Mccain, J. C. & Davidson, E." box="[812,1042,691,717]" pageId="22" pageNumber="243" refId="ref14912" refString="Schmitt, W. L., Mccain, J. C. & Davidson, E. (1973) Decapoda I. Brachyura I. Fam. Pinnotheridae. Crustaceorum Catalogus. Vol. 3. Dr. W. Junk, Den Haag, 160 pp." type="book" year="1973">
|
||
Schmitt
|
||
<emphasis id="A9C8EADD9518427B70BFFD74FC6FC0A4" box="[906,955,691,717]" italics="true" pageId="22" pageNumber="243">et al</emphasis>
|
||
. (1973)
|
||
</bibRefCitation>
|
||
also listed
|
||
<taxonomicName id="5CBC4D4C9518427B77A5FD74FAB9C0A7" authorityName="Linnaeus" authorityYear="1767" box="[1168,1389,691,718]" class="Bivalvia" family="Mactridae" genus="Mactra" kingdom="Animalia" order="Veneroida" pageId="22" pageNumber="243" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A9C8EADD9518427B77A5FD74FB36C0A5" box="[1168,1250,691,716]" italics="true" pageId="22" pageNumber="243">Mactra</emphasis>
|
||
(Mactridae)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5CBC4D4C9518427B73A2FD10FE5DC09B" box="[151,393,727,754]" class="Bivalvia" family="Placunidae" genus="Placuna" kingdom="Animalia" order="Pectinoida" pageId="22" pageNumber="243" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A9C8EADD9518427B73A2FD10FF20C098" box="[151,244,727,753]" italics="true" pageId="22" pageNumber="243">Placuna</emphasis>
|
||
(Placunidae)
|
||
</taxonomicName>
|
||
as host bivalves for
|
||
<taxonomicName id="5CBC4D4C9518427B7145FD10FD2DC098" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[624,761,727,753]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="palaensis">
|
||
<emphasis id="A9C8EADD9518427B7145FD10FD2DC098" box="[624,761,727,753]" italics="true" pageId="22" pageNumber="243">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
but these records are probably incorrect and the pinnotherid species was probably misidentified. For
|
||
<taxonomicName id="5CBC4D4C9518427B7169FD3CFD6DC17C" box="[604,697,763,789]" class="Bivalvia" family="Placunidae" genus="Placuna" kingdom="Animalia" order="Pectinoida" pageId="22" pageNumber="243" phylum="Mollusca" rank="genus">
|
||
<emphasis id="A9C8EADD9518427B7169FD3CFD6DC17C" box="[604,697,763,789]" italics="true" pageId="22" pageNumber="243">Placuna</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="E3AB765F9518427B71CBFD3CFCA6C17C" box="[766,882,763,789]" name="Singapore" pageId="22" pageNumber="243">Singapore</collectingCountry>
|
||
, at least,
|
||
<bibRefCitation id="FF2D4B3E9518427B70EFFD3CFB9AC17C" author="Ng, P. K. L." box="[986,1102,763,789]" pageId="22" pageNumber="243" pagination="474 - 485" refId="ref14331" refString="Ng, P. K. L. (2018) Arcotheres placunicola, a new species of pea crab (Crustacea: Brachyura: Pinnotheridae) from the windowpane shell, Placuna ephippium Philipsson, 1788 (Placunidae) in Singapore. Raffles Bulletin of Zoology, 66, 474 - 485." type="journal article" year="2018">Ng (2018)</bibRefCitation>
|
||
showed that the species present is
|
||
<taxonomicName id="5CBC4D4C9518427B73EFFCD8FE25C150" authority="Ng, 2018" authorityName="Ng" authorityYear="2018" box="[218,497,799,825]" class="Bivalvia" family="Arcidae" genus="Anadara" kingdom="Animalia" order="Arcoida" pageId="22" pageNumber="243" phylum="Mollusca" rank="species" species="placunicola">
|
||
<emphasis id="A9C8EADD9518427B73EFFCD8FEA9C150" box="[218,381,799,825]" italics="true" pageId="22" pageNumber="243">A. placunicola</emphasis>
|
||
<bibRefCitation id="FF2D4B3E9518427B72B1FCD8FE25C150" author="Ng, P. K. L." box="[388,497,799,825]" pageId="22" pageNumber="243" pagination="474 - 485" refId="ref14331" refString="Ng, P. K. L. (2018) Arcotheres placunicola, a new species of pea crab (Crustacea: Brachyura: Pinnotheridae) from the windowpane shell, Placuna ephippium Philipsson, 1788 (Placunidae) in Singapore. Raffles Bulletin of Zoology, 66, 474 - 485." type="journal article" year="2018">Ng, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="CFC366479518427B73A2F8AAFBE5C598" ID-DOI="http://doi.org/10.5281/zenodo.4401497" ID-Zenodo-Dep="4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="22" pageNumber="243" startId="22.[151,250,1901,1925]" targetBox="[180,1407,855,1867]" targetPageId="22">
|
||
<paragraph id="9B0336CF9518427B73A2F8AAFBE5C598" blockId="22.[151,1436,1901,2033]" pageId="22" pageNumber="243">
|
||
<emphasis id="A9C8EADD9518427B73A2F8AAFECBC5EC" bold="true" box="[151,287,1901,1925]" pageId="22" pageNumber="243">FIGURE 16.</emphasis>
|
||
<taxonomicName id="5CBC4D4C9518427B7210F8AAFE23C5EC" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[293,503,1901,1925]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="243" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9518427B7210F8AAFE23C5EC" box="[293,503,1901,1925]" italics="true" pageId="22" pageNumber="243">Arcotheres ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9518427B72C8F8A9FDE1C5EF" bold="true" box="[509,565,1902,1926]" pageId="22" pageNumber="243">
|
||
<taxonomicNameLabel id="B2FB57A69518427B72C8F8A9FDE1C5EF" box="[509,565,1902,1926]" pageId="22" pageNumber="243" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B0336CF9519427A73F2FF50FB7AC3C7" blockId="23.[151,1437,151,610]" pageId="23" pageNumber="244">
|
||
Compared to
|
||
<taxonomicName id="5CBC4D4C9519427A726BFF50FE27C2D8" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[350,499,151,177]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="arcophilus">
|
||
<emphasis id="A9C8EADD9519427A726BFF50FE27C2D8" box="[350,499,151,177]" italics="true" pageId="23" pageNumber="244">A. arcophilus</emphasis>
|
||
</taxonomicName>
|
||
, in which the eyes are also distinctly visible in the dorsal view, in
|
||
<taxonomicName id="5CBC4D4C9519427A77E5FF50FA8CC2D8" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[1232,1368,151,177]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9519427A77E5FF50FA8CC2D8" box="[1232,1368,151,177]" italics="true" pageId="23" pageNumber="244">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9519427A766BFF50FA48C2D8" bold="true" box="[1374,1436,151,177]" pageId="23" pageNumber="244">
|
||
<taxonomicNameLabel id="B2FB57A69519427A766BFF50FA48C2D8" box="[1374,1436,151,177]" pageId="23" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the maxilliped 3 dactylus reaches to or extends beyond the tip of the propodus (
|
||
<figureCitation id="03872A4A9519427A7722FF7CFB6DC2BC" box="[1047,1209,187,213]" captionStart="FIGURE 8" captionStartId="14.[151,250,1828,1852]" captionTargetBox="[161,1426,190,1794]" captionTargetId="figure-20@14.[151,1436,181,1805]" captionTargetPageId="14" captionText="FIGURE 8. Arcotheres ocularius n. sp. A–E, holotype, ovigerous female (9.5×10.7 mm), CBM-ZC 15952; F, paratype, ovigerous female (9.7×10.9 mm), ZRC 2019.1877, Fiji; G, non-type, female of post-hard stage V (8.4×9.4 mm), ZRC 2019.1025, Lombok. A, carapace and eyes, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right maxilliped 3, outer view; D, anterior part of thoracic sternum, ventral view; E, pleomere 6 and telson, ventral view; F, G, left maxilliped 3, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401479" httpUri="https://zenodo.org/record/4401479/files/figure.png" pageId="23" pageNumber="244">Figs. 8C, F, G</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9519427A77F0FF7CFB22C2BF" box="[1221,1270,187,214]" captionStart="FIGURE 16" captionStartId="22.[151,250,1901,1925]" captionTargetBox="[180,1407,855,1867]" captionTargetId="figure-370@22.[173,1414,849,1878]" captionTargetPageId="22" captionText="FIGURE 16. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954.A, right maxilliped 3, outer view; B, left chela, outer view; C, merus of left cheliped, ventral view; D, right P2, dactylus and propodus, dorsal view; E, right P3, dactylus and propodus, dorsal view; F, right P4 (shorter side), dactylus and propodus, dorsal view; G, left P4 (longer side), dactylus and propodus, dorsal view; H, right P5, dactylus and propodus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4401497" httpUri="https://zenodo.org/record/4401497/files/figure.png" pageId="23" pageNumber="244">16A</figureCitation>
|
||
) (versus never reaching the tip of the propodus in
|
||
<taxonomicName id="5CBC4D4C9519427A7116FF18FD6DC290" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[547,697,223,249]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="arcophilus">
|
||
<emphasis id="A9C8EADD9519427A7116FF18FD6DC290" box="[547,697,223,249]" italics="true" pageId="23" pageNumber="244">A. arcophilus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="FF2D4B3E9519427A71DEFF18FC1BC290" author="Ahyong, S. T. & Ng, P. K. L." box="[747,975,223,249]" pageId="23" pageNumber="244" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
|
||
: fig. 2C), the female P4 merus is proportionately longer in
|
||
<taxonomicName id="5CBC4D4C9519427A7258FEC4FE23C374" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[365,503,259,285]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9519427A7258FEC4FE23C374" box="[365,503,259,285]" italics="true" pageId="23" pageNumber="244">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9519427A72CBFEC4FDEAC374" bold="true" box="[510,574,259,285]" pageId="23" pageNumber="244">
|
||
<taxonomicNameLabel id="B2FB57A69519427A72CBFEC4FDEAC374" box="[510,574,259,285]" pageId="23" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="03872A4A9519427A717BFEC4FD77C374" box="[590,675,259,285]" captionStart="FIGURE 9" captionStartId="15.[151,250,1866,1890]" captionTargetBox="[206,1379,190,1831]" captionTargetId="figure-17@15.[197,1390,181,1842]" captionTargetPageId="15" captionText="FIGURE 9. Arcotheres ocularius n. sp., holotype, ovigerous female (10.2×11.8 mm), CBM-ZC 15952. A, left chela, outer view; B, chela and carpus of left cheliped, dorsal view; C, distal part of fixed finger of left chela, outer view; D, same, ventral view; E, merus of left cheliped, ventral view; F, right P2, dorsal view; G, same, dactylus; H, right P3, dorsal view; I, same, dactylus; J, right P4 (longer side), dorsal view; K, same, dactylus; L, left P4 (shorter side), dorsal view; M, same, dactylus; N, right P5, dorsal view; O, same, dactylus." figureDoi="http://doi.org/10.5281/zenodo.4401481" httpUri="https://zenodo.org/record/4401481/files/figure.png" pageId="23" pageNumber="244">Fig. 9L</figureCitation>
|
||
) (versus proportionately shorter in
|
||
<taxonomicName id="5CBC4D4C9519427A7707FEC4FB1DC374" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1074,1225,259,285]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="arcophilus">
|
||
<emphasis id="A9C8EADD9519427A7707FEC4FB1DC374" box="[1074,1225,259,285]" italics="true" pageId="23" pageNumber="244">A. arcophilus</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="FF2D4B3E9519427A77CEFEC4FF07C328" author="Ahyong, S. T. & Ng, P. K. L." pageId="23" pageNumber="244" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
|
||
: fig. 2A), the male telson is subpentagonal or subhexagonal in
|
||
<taxonomicName id="5CBC4D4C9519427A709CFEE0FBE0C328" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[937,1076,295,321]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9519427A709CFEE0FBE0C328" box="[937,1076,295,321]" italics="true" pageId="23" pageNumber="244">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2FB57A69519427A7708FEE0FBAFC328" box="[1085,1147,295,321]" pageId="23" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="03872A4A9519427A77B9FEE0FAD4C328" box="[1164,1280,295,321]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="23" pageNumber="244">Figs. 13H</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9519427A7638FEE1FAEEC328" box="[1293,1338,294,321]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="23" pageNumber="244">15F</figureCitation>
|
||
) (versus semicircular in
|
||
<taxonomicName id="5CBC4D4C9519427A7272FE8CFE0AC30C" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[327,478,331,357]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="arcophilus">
|
||
<emphasis id="A9C8EADD9519427A7272FE8CFE0AC30C" box="[327,478,331,357]" italics="true" pageId="23" pageNumber="244">A. arcophilus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="FF2D4B3E9519427A72D9FE8CFD06C30C" author="Ahyong, S. T. & Ng, P. K. L." box="[492,722,331,357]" pageId="23" pageNumber="244" pagination="191 - 226" refId="ref12598" refString="Ahyong, S. T. & Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura. Raffles Bulletin of Zoology, Supplement 16, 191 - 226." type="journal article" year="2007">Ahyong & Ng 2007</bibRefCitation>
|
||
: fig. 2E), and the G1 is C-shaped with the tip subtruncate in
|
||
<taxonomicName id="5CBC4D4C9519427A76B1FE8CFED5C3E0" authority="Komai & Kawai & Ng, 2020" authorityName="Komai & Kawai & Ng" authorityYear="2020" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="ocularius" status="sp. nov.">
|
||
<emphasis id="A9C8EADD9519427A76B1FE8CFED5C3E0" italics="true" pageId="23" pageNumber="244">A. ocularius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A9C8EADD9519427A7232FEA8FE91C3E0" bold="true" box="[263,325,367,393]" pageId="23" pageNumber="244">
|
||
<taxonomicNameLabel id="B2FB57A69519427A7232FEA8FE91C3E0" box="[263,325,367,393]" pageId="23" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="03872A4A9519427A7261FEA8FE06C3E0" box="[340,466,367,393]" captionStart="FIGURE 13" captionStartId="19.[151,250,1850,1874]" captionTargetBox="[161,1425,191,1815]" captionTargetId="figure-17@19.[151,1436,181,1826]" captionTargetPageId="19" captionText="FIGURE 13. Arcotheres ocularius n. sp., paratype, male of hard stage (5.0×4.6 mm), CBM-ZC 15957. A, carapace with eyes and antennae, dorsal view; B, frontal part of carapace and cephalic appendages, anterior view; C, right MXP3, outer view; D, thoracic sternites 3 and 4 with telson; E, left exposed part of thoracic sternites 5–8, ventral view; F, sterno-pleonal cavity with right G1 and G2, ventral view; G, left exposed part of thoracic sternite 8, posterior view (perpendicular); H, pleon and telson, ventral view; I, left G1, outer view; J, same, distal part, inner view; K, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401489" httpUri="https://zenodo.org/record/4401489/files/figure.png" pageId="23" pageNumber="244">Figs. 13I, J</figureCitation>
|
||
,
|
||
<figureCitation id="03872A4A9519427A72E8FEA9FDFBC3E0" box="[477,559,366,393]" captionStart="FIGURE 15" captionStartId="21.[151,250,1870,1894]" captionTargetBox="[175,1414,188,1838]" captionTargetId="figure-17@21.[166,1422,181,1847]" captionTargetPageId="21" captionText="FIGURE 15. Arcotheres ocularius n. sp., paratype, male of post-hard stage (6.8×6.2 mm), CBM-ZC 15954. A, habitus, dorsal view; B, thoracic sternites 3 and 4, ventral view; C, left exposed part of thoracic sternites 5–8, ventral view; D, left exposed part of thoracic sternite 8, posterior view (perpendicular); E, sterno-pleonal cavity with proximal part of right G1 and right G2 preserved (left G1 and G2 dissected), ventral view; F, pleon and telson, ventral view; G, left G1, outer view; H, same, distal part, inner view; I, left G2, outer view." figureDoi="http://doi.org/10.5281/zenodo.4401493" httpUri="https://zenodo.org/record/4401493/files/figure.png" pageId="23" pageNumber="244">15G, H</figureCitation>
|
||
) (versus G1 more slender and elongate, being only broadly arcuate with the tip possessing an elongate projection in
|
||
<taxonomicName id="5CBC4D4C9519427A7106FE54FD1DC3C4" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[563,713,403,429]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="arcophilus">
|
||
<emphasis id="A9C8EADD9519427A7106FE54FD1DC3C4" box="[563,713,403,429]" italics="true" pageId="23" pageNumber="244">A. arcophilus</emphasis>
|
||
</taxonomicName>
|
||
; PKLN & S.T. Ahyong, unpublished data).
|
||
</paragraph>
|
||
<paragraph id="9B0336CF9519427A73F2FE70FD33C054" blockId="23.[151,1437,151,610]" pageId="23" pageNumber="244">
|
||
In an ongoing genetic study of Asian pinnotherids, the Fijian and present Lombok specimens cluster closely with each other, distinct from what is here defined as
|
||
<taxonomicName id="5CBC4D4C9519427A71DBFE1CFCA3C39C" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[750,887,475,501]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" sensu="stricto" species="palaensis">
|
||
<emphasis id="A9C8EADD9519427A71DBFE1CFCA3C39C" box="[750,887,475,501]" italics="true" pageId="23" pageNumber="244">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2FB57A69519427A704BFE1CFC6CC39C" box="[894,952,475,501]" pageId="23" pageNumber="244" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
on the basis of the 16S mitochondrial gene (Tsang Ling-Ming, pers. comm.). Lombok actually has both species, with another Lombok female specimen (ZRC 2018.762) clustering with
|
||
<taxonomicName id="5CBC4D4C9519427A728AFDE4FD9CC054" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[447,584,547,573]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" sensu="stricto" species="palaensis">
|
||
<emphasis id="A9C8EADD9519427A728AFDE4FD9CC054" box="[447,584,547,573]" italics="true" pageId="23" pageNumber="244">A. palaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B2FB57A69519427A717AFDE4FD5DC054" box="[591,649,547,573]" pageId="23" pageNumber="244" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
instead.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A665449519427A73F2FD80FAA3C008" box="[199,1399,583,610]" pageId="23" pageNumber="244" type="etymology">
|
||
<paragraph id="9B0336CF9519427A73F2FD80FAA3C008" blockId="23.[151,1437,151,610]" box="[199,1399,583,610]" pageId="23" pageNumber="244">
|
||
<emphasis id="A9C8EADD9519427A73F2FD80FE9AC008" bold="true" box="[199,334,583,609]" pageId="23" pageNumber="244">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="5CBC4D4C9519427A7170FD80FD7BC008" authorityName="Komai & Kawai & Ng" authorityYear="2020" box="[581,687,583,609]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="244" phylum="Arthropoda" rank="species" species="ocularius">
|
||
<emphasis id="A9C8EADD9519427A7170FD80FD7BC008" box="[581,687,583,609]" italics="true" pageId="23" pageNumber="244">ocularius</emphasis>
|
||
</taxonomicName>
|
||
” (= optic) refers to the dorsally visible eyes in the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |