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<document id="6BA4253C6940EC8B81A27F7FFE1D3DB2" ID-CLB-Dataset="157781" ID-DOI="10.11646/zootaxa.5192.1.1" ID-GBIF-Dataset="8e2cc009-d580-4904-a0b7-9ed7a9e3ab9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138639" ID-ZooBank="236A609B-8A6A-47D7-9BB9-E2FDCA5C37E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664794801684" checkinUser="plazi" docAuthor="Kuwahara, Gregory K. &amp; Marshall, Stephen A." docDate="2022" docId="0F519B74FFB7AD4DFF2C93E1FB90FEE6" docLanguage="en" docName="zootaxa.5192.1.1.pdf" docOrigin="Zootaxa 5192 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5192.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Howickia dealata Kuwahara &amp; Marshall 2022, comb. nov." docType="treatment" docVersion="6" lastPageNumber="113" masterDocId="F368E30CFFD9AD3DFFBB9376FFA9FFFB" masterDocTitle="A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)" masterLastPageNumber="152" masterPageNumber="1" pageNumber="111" updateTime="1699386905592" updateUser="ExternalLinkService">
<mods:mods id="1312E02EB08931E182410E07B4CC4881" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="61B54C2F53D60B17F1B4E65D6E3105D7">A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)</mods:title>
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<mods:namePart id="E56E2CBF055ECB0F5B0699555AF51439">Kuwahara, Gregory K.</mods:namePart>
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<mods:namePart id="4F70B0E6FBAA5DF4190E70B68B07D989">Marshall, Stephen A.</mods:namePart>
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<mods:date id="201E144FC92FB1A22BC0BE3D324A962F">2022</mods:date>
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<mods:number id="BDF95A08D2F93F6DB30A83EE9A5EA67E">2022-10-03</mods:number>
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<mods:number id="489F098D01C8DC3E9957B9A8226C0BC5">5192</mods:number>
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<treatment id="0F519B74FFB7AD4DFF2C93E1FB90FEE6" ID-DOI="http://doi.org/10.5281/zenodo.7138790" ID-GBIF-Taxon="202134573" ID-Zenodo-Dep="7138790" LSID="urn:lsid:plazi:treatment:0F519B74FFB7AD4DFF2C93E1FB90FEE6" httpUri="http://treatment.plazi.org/id/0F519B74FFB7AD4DFF2C93E1FB90FEE6" lastPageId="112" lastPageNumber="113" pageId="110" pageNumber="111">
<subSubSection id="CFE279E9FFB7AD53FF2C93E1FD2CFF4A" box="[151,645,151,178]" pageId="110" pageNumber="111" type="nomenclature">
<paragraph id="87472A62FFB7AD53FF2C93E1FD2CFF4A" blockId="110.[151,645,151,214]" box="[151,645,151,178]" pageId="110" pageNumber="111">
<heading id="DC0F9D0EFFB7AD53FF2C93E1FD2CFF4A" bold="true" box="[151,645,151,178]" fontSize="11" level="2" pageId="110" pageNumber="111" reason="4">
<emphasis id="B58CF670FFB7AD53FF2C93E1FD2CFF4A" bold="true" box="[151,645,151,178]" pageId="110" pageNumber="111">
<taxonomicName id="40F851E1FFB7AD53FF2C93E1FE5EFF4A" ID-CoL="BD3KW" authority="(Richards)" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" baseAuthorityName="Kuwahara &amp; Marshall" baseAuthorityYear="2022" box="[151,503,151,178]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="dealata" status="comb. nov.">
<emphasis id="B58CF670FFB7AD53FF2C93E1FEC3FF49" bold="true" box="[151,362,151,178]" italics="true" pageId="110" pageNumber="111">Howickia dealata</emphasis>
(Richards)
</taxonomicName>
<taxonomicNameLabel id="AEBF4B0BFFB7AD53FE4593E1FD2CFF4A" box="[510,645,151,177]" pageId="110" pageNumber="111" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB7AD53FF2C93CDFEF1FF2D" box="[151,344,187,214]" pageId="110" pageNumber="111" type="description">
<paragraph id="87472A62FFB7AD53FF2C93CDFEF1FF2D" blockId="110.[151,645,151,214]" box="[151,344,187,214]" pageId="110" pageNumber="111">
(
<figureCitation id="1FC336E7FFB7AD53FF2493CDFEF9FF2D" box="[159,336,187,214]" captionStart="FIGURES 32.1–32.5" captionStartId="111.[152,270,1943,1967]" captionTargetBox="[156,1431,181,1902]" captionTargetId="figure-19@111.[156,1431,181,1919]" captionTargetPageId="111" captionText="FIGURES 32.1–32.5. Howickia dealata (Richards). 32.1, male habitus, lateral; 32.2, male head, anterior; 32.3, epiproct and cercus, dorsal; 32.4, S8 and hypoproct, ventral; 32.5, spermathecae. Body scale bar indicates 0.25 mm, head bar indicates 0.1 mm. Photo credit: G. Thompson." figureDoi="http://doi.org/10.5281/zenodo.7138839" httpUri="https://zenodo.org/record/7138839/files/figure.png" pageId="110" pageNumber="111">Figs. 32.1–32.5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB7AD53FF2C9276FB91FEE3" box="[151,1080,255,280]" pageId="110" pageNumber="111" type="reference_group">
<paragraph id="87472A62FFB7AD53FF2C9276FB91FEE3" blockId="110.[151,1080,255,280]" box="[151,1080,255,280]" pageId="110" pageNumber="111">
<treatmentCitationGroup id="A7E80D4CFFB7AD53FF2C9276FB91FEE3" box="[151,1080,255,280]" pageId="110" pageNumber="111">
<treatmentCitation id="06590C73FFB7AD53FF2C9276FDE1FEE3" author="Richards, O. W." box="[151,584,256,280]" page="393" pageId="110" pageNumber="111" year="1973">
<taxonomicName id="40F851E1FFB7AD53FF2C9276FDE1FEE3" ID-CoL="6S27V" authority="Richards, 1973: 393" authorityName="Richards" authorityPageNumber="393" authorityYear="1973" box="[151,584,256,280]" class="Insecta" family="Sphaeroceridae" genus="Monteithiana" kingdom="Animalia" order="Diptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="dealata">
<emphasis id="B58CF670FFB7AD53FF2C9276FEDBFEE3" box="[151,370,256,280]" italics="true" pageId="110" pageNumber="111">Monteithiana dealata</emphasis>
<bibRefCitation id="E3695793FFB7AD53FEC39276FDE1FEE3" author="Richards, O. W." box="[376,584,256,280]" pageId="110" pageNumber="111" pagination="297 - 401" refId="ref94252" refString="Richards, O. W. (1973) The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australian region. Australian Journal of Zoology Supplemental Series, 22, 297 - 401. https: // doi. org / 10.1071 / AJZS 022" type="journal article" year="1973">Richards, 1973: 393</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="06590C73FFB7AD53FDE99276FC8BFEEC" author="Marshall, S. A." box="[594,802,255,280]" page="604" pageId="110" pageNumber="111" year="1989">
<bibRefCitation id="E3695793FFB7AD53FDE99276FC8BFEEC" author="Marshall, S. A." box="[594,802,255,280]" pageId="110" pageNumber="111" pagination="601 - 607" refId="ref93510" refString="Marshall, S. A. (1989) 96. Family Sphaeroceridae. In: Evenhuis, N. L. (Ed.), Catalog of Diptera of the Australasian and Oceanic Regions. Bishop Museum Press and E. J. Brill, Honolulu, pp. 601 - 607." type="book chapter" year="1989">Marshall, 1989: 604</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="06590C73FFB7AD53FC969276FB9DFEE3" author="Rohacek, J. &amp; Marshall, S. A. &amp; Norrbom, A. L. &amp; Buck, M. &amp; Quiros, D. I. &amp; Smith, I." box="[813,1076,256,280]" page="172" pageId="110" pageNumber="111" year="2001">
<bibRefCitation id="E3695793FFB7AD53FC969276FB9DFEE3" author="Rohacek, J. &amp; Marshall, S. A. &amp; Norrbom, A. L. &amp; Buck, M. &amp; Quiros, D. I. &amp; Smith, I." box="[813,1076,256,280]" pageId="110" pageNumber="111" refId="ref94357" refString="Rohacek, J., Marshall, S. A., Norrbom, A. L., Buck, M., Quiros, D. I. &amp; Smith, I. (2001) World Catalog of Sphaeroceridae. Slezske zemske Museum, Opava, 415 pp." type="book" year="2001">
Roháček
<emphasis id="B58CF670FFB7AD53FC379276FC16FEE3" box="[908,959,256,280]" italics="true" pageId="110" pageNumber="111">et al.</emphasis>
, 2001: 172
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB7AD53FF2C9233FD93FA1B" pageId="110" pageNumber="111" type="description">
<paragraph id="87472A62FFB7AD53FF2C9233FB76FE9B" blockId="110.[151,1437,325,2008]" box="[151,1247,325,352]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF2C9233FE80FEA4" bold="true" box="[151,297,325,351]" pageId="110" pageNumber="111">Description.</emphasis>
<emphasis id="B58CF670FFB7AD53FE8B9230FE69FE9B" box="[304,448,326,352]" italics="true" pageId="110" pageNumber="111">Body length:</emphasis>
<quantity id="40008787FFB7AD53FE7D9230FDE6FE9B" box="[454,591,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="110" pageNumber="111" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 mm</quantity>
, males smaller than females (as stated by
<bibRefCitation id="E3695793FFB7AD53FB9E9230FB7AFE9B" author="Richards, O. W." box="[1061,1235,326,352]" pageId="110" pageNumber="111" pagination="297 - 401" refId="ref94252" refString="Richards, O. W. (1973) The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australian region. Australian Journal of Zoology Supplemental Series, 22, 297 - 401. https: // doi. org / 10.1071 / AJZS 022" type="journal article" year="1973">Richards, 1973</bibRefCitation>
).
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C921CFB55FDC3" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C921CFEA4FE7F" box="[199,269,362,388]" italics="true" pageId="110" pageNumber="111">Head:</emphasis>
Golden-yellow, upper half of frontal vitta, orbital plates and behind dark brown, lower half of frontal vitta pale yellow; orbital plates and ocellar triangle microtrichose; gena brownish-yellow; antenna yellow, apically darkened. Frontal width 1.4x interfrontal height, frontal vitta narrow. Two pairs of large interfrontal bristles preceded by a smaller pair of setae; 2 lateroclinate orbital bristles, anterior bristle 0.7x length of posterior bristle. Gena with strong but fine upcurved bristle and 4–5 fine upcurved setae, finely striate, shiny, lower third microtrichose; vibrissal angle with 1 subvibrissal seta. Palpus clavate with 2 large apical setae. Eye height 3.2x genal height.
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C9134FE1FFD33" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C9134FE88FDA7" box="[199,289,578,604]" italics="true" pageId="110" pageNumber="111">Thorax:</emphasis>
Black; scutum shiny despite fine brown microtrichia, posterior edge slightly reddish. One pair of dorsocentral bristles; 7–8 rows of short, stout acrostichal setulae. Prosternum broad, anteriorly acute. Scutellum shiny but heavily microtrichose, black, semicircular, its length 0.3x its width; 4 scutellar bristles long, basal pair 1.3x length of apical pair.
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C91A5FBE0FCCF" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C91A5FEAFFD17" box="[199,262,723,748]" italics="true" pageId="110" pageNumber="111">Legs:</emphasis>
Dark brown, femora proximally and tibiae proximally and distally lighter; tarsi yellow-brown. Male mid tibia with a moderate apicoventral bristle. Basal half of mid tibia with 2 dorsal bristles (1 anterodorsal and 1 posterodorsal), distal half with 2 dorsal bristles (1 anterodorsal and 1 posterodorsal).
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C9048FE51FCA3" blockId="110.[151,1437,325,2008]" box="[199,504,830,856]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C9048FEFCFCA3" box="[199,341,830,856]" italics="true" pageId="110" pageNumber="111">Wing/halter:</emphasis>
Halter absent.
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C9014FB00FBAF" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C9014FEC6FC87" box="[199,367,866,892]" italics="true" pageId="110" pageNumber="111">Male abdomen</emphasis>
: Black, heavily sclerotized, shiny, postabdomen slightly paler. T2–5 and S2–4 uniformly longsetose along entire surface. S5 broad but short, densely setose with posteromedial margin broadly desclerotized and bearing a row of 6 short, flattened teeth. Epandrium large, hemispherical, and densely long-setose; cercus semicircular with a large sinuate seta and several smaller setae. Hypandrium medially strongly curved; ventral and posterodorsal lobes present but undescribed (see comments). Surstylus subtriangular with dense setation along posterior surface and a stout apicoventral thorn-like seta. Postgonite elongate, tapered and slightly curved. Phallapodeme slightly curved. Distiphallus apparently largely membranous with two lateral, well-sclerotized plates.
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C9728FD93FA1B" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C9728FE3EFB83" box="[199,407,1118,1144]" italics="true" pageId="110" pageNumber="111">Female abdomen:</emphasis>
Black, heavily sclerotized, shiny despite microtrichia. T2–5 and S2–5 wide, uniformly long-setose along entire surface with large posterolateral setae; pleural membrane bare. Preabdomen subequal in length to head + thorax. Postabdomen retracted into preabdomen and hardened, making most of the postabdomen impossible to examine. Epiproct shield-like, medially split into 2 halves and dorsomedially bearing a pair of setae. Cercus short, subtriangular with 5 long sinuate setae (2 apical, 2 basolateral, 1 dorsal and 1 mesial) and several smaller setulae. S6 and S7 (as could be seen) relatively well-developed, present as broad, rectangular, microtrichose plates bearing a posterior row of setae. S8 elongate, ovoid with a preapical pair of setae surrounded by a patch of microtrichia. Hypoproct broad, shield-like, posteriorly microtrichose with a preapical row of setae. Spermathecae stout, cylindrical and strongly grooved; stem short (0.2x length of spermatheca) and connected to a bulbous basal expansion (appearing as a swollen disk with a sphere within the center); paired spermathecae both connected to the apical portion of the basal expansion.
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB7AD53FF7C969CFDADF927" pageId="110" pageNumber="111" type="materials_examined">
<paragraph id="87472A62FFB7AD53FF7C969CFBC6F9D3" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C969CFED3F9F8" bold="true" box="[199,378,1513,1540]" pageId="110" pageNumber="111">Type material.</emphasis>
<materialsCitation id="3790203FFFB7AD53FE3D969CFE47F9D3" ID-GBIF-Occurrence="3920718546" collectingDate="1967-02-02" collectingDateMax="1967-02-04" collectingDateMin="1967-02-02" collectionCode="QMBA" collectorName="G. Monteith" country="Australia" location="Hellyer Gorge" pageId="110" pageNumber="111" specimenCount="1" specimenCount-male="1" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="584394C0FFB7AD53FE3D969CFE5AF9FF" box="[390,499,1514,1540]" pageId="110" pageNumber="111" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="FFEF6AF2FFB7AD53FDBB969CFD0AF9FF" box="[512,675,1514,1540]" name="Australia" pageId="110" pageNumber="111">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FFB7AD53FD09969FFC86F9F8" bold="true" box="[690,815,1513,1539]" pageId="110" pageNumber="111">
<collectingRegion id="453CE480FFB7AD53FD09969FFC83F9F8" box="[690,810,1513,1539]" country="Australia" name="Tasmania" pageId="110" pageNumber="111">Tasmania</collectingRegion>
:
</emphasis>
<location id="82277CB9FFB7AD53FC80969CFC4AF9FF" LSID="urn:lsid:plazi:treatment:0F519B74FFB7AD4DFF2C93E1FB90FEE6:82277CB9FFB7AD53FC80969CFC4AF9FF" box="[827,995,1514,1540]" country="Australia" name="Hellyer Gorge" pageId="110" pageNumber="111" stateProvince="Tasmania">Hellyer Gorge</location>
,
<locationDeviation id="96F53340FFB7AD53FC48969CFB76F9FF" box="[1011,1247,1514,1540]" location="Waratah" pageId="110" pageNumber="111">
<quantity id="40008787FFB7AD53FC48969CFBFDF9FF" box="[1011,1108,1514,1540]" metricMagnitude="4" metricUnit="m" metricValue="2.41" pageId="110" pageNumber="111" unit="km" value="24.1">24.1 km</quantity>
N Waratah
</locationDeviation>
,
<date id="F3460CA2FFB7AD53FB54969CFAD8F9FF" box="[1263,1393,1514,1540]" pageId="110" pageNumber="111" value="1967-02-02" valueMax="1967-02-04" valueMin="1967-02-02">
<collectingDate id="E302F54AFFB7AD53FB54969CFAD8F9FF" box="[1263,1393,1514,1540]" pageId="110" pageNumber="111" value="1967-02-02" valueMax="1967-02-04" valueMin="1967-02-02">2–4.ii.1967</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FFB7AD53FA3A969CFEA8F9D3" pageId="110" pageNumber="111">G. Monteith</collectorName>
, T.7126 (
<specimenCount id="91FEE1EBFFB7AD53FED19578FE28F9D3" box="[362,385,1550,1576]" pageId="110" pageNumber="111" type="male">♂</specimenCount>
,
<collectionCode id="E1E9B2A7FFB7AD53FE369578FE4FF9D3" box="[397,486,1550,1576]" pageId="110" pageNumber="111">QMBA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3790203FFFB7AD53FE439578FBC5F9D3" ID-GBIF-Occurrence="3920718645" box="[504,1132,1550,1576]" collectionCode="QMBA" country="Australia" pageId="110" pageNumber="111" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="584394C0FFB7AD53FE439578FDC9F9D3" box="[504,608,1550,1576]" pageId="110" pageNumber="111" type="holotype">Holotype</typeStatus>
photos provided by Geoff Thompson,
<collectionCode id="E1E9B2A7FFB7AD53FBAF9578FBC5F9D3" box="[1044,1132,1550,1576]" pageId="110" pageNumber="111">QMBA</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C9544FEB5F98B" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<materialsCitation id="3790203FFFB7AD53FF7C9544FB50F9B7" ID-GBIF-Occurrence="3920718775" box="[199,1273,1586,1613]" collectionCode="UQIC, BMNH" country="Australia" pageId="110" pageNumber="111" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Tasmania" typeStatus="paratype">
<typeStatus id="584394C0FFB7AD53FF7C9544FE91F9B7" box="[199,312,1586,1612]" pageId="110" pageNumber="111" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="FFEF6AF2FFB7AD53FEF89544FE4FF9B7" box="[323,486,1586,1612]" name="Australia" pageId="110" pageNumber="111">AUSTRALIA</collectingCountry>
: same data as holotype (
<specimenCount id="91FEE1EBFFB7AD53FD459544FC85F9B7" box="[766,812,1586,1613]" pageId="110" pageNumber="111" type="male">5 ♂</specimenCount>
,
<specimenCount id="91FEE1EBFFB7AD53FC819544FCCCF9B7" box="[826,869,1586,1612]" pageId="110" pageNumber="111" type="female">6 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFB7AD53FCCE9544FC10F9B7" box="[885,953,1586,1612]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="110" pageNumber="111">UQIC</collectionCode>
&amp;
<collectionCode id="E1E9B2A7FFB7AD53FC5B9544FB90F9B7" box="[992,1081,1586,1612]" pageId="110" pageNumber="111">BMNH</collectionCode>
). Types in
<collectionCode id="E1E9B2A7FFB7AD53FB0E9544FB50F9B7" box="[1205,1273,1586,1612]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34515" lsid="urn:lsid:biocol.org:col:34515" name="University of Queensland Insect Collection" pageId="110" pageNumber="111">UQIC</collectionCode>
</materialsCitation>
now stored at the
<collectionCode id="E1E9B2A7FFB7AD53FF7B9520FEB1F98B" box="[192,280,1622,1648]" pageId="110" pageNumber="111">QMBA</collectionCode>
.
</paragraph>
<paragraph id="87472A62FFB7AD53FF7C950FFDADF927" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C950FFE11F968" bold="true" box="[199,440,1657,1683]" pageId="110" pageNumber="111">Material examined.</emphasis>
<collectingCountry id="FFEF6AF2FFB7AD53FE7F950CFDCEF96F" box="[452,615,1658,1684]" name="Australia" pageId="110" pageNumber="111">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FFB7AD53FDC2950FFD5FF968" bold="true" box="[633,758,1657,1683]" pageId="110" pageNumber="111">
<collectingRegion id="453CE480FFB7AD53FDC2950FFD58F968" box="[633,753,1657,1683]" country="Australia" name="Tasmania" pageId="110" pageNumber="111">Tasmania</collectingRegion>
:
</emphasis>
Mount Field National Park, near Lake Fenton, 42˚41’S 146˚37’E,
<quantity id="40008787FFB7AD53FEAB95E8FEC1F943" box="[272,360,1694,1720]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="110" pageNumber="111" unit="m" value="1000.0">1000 m</quantity>
,
<collectionCode id="E1E9B2A7FFB7AD53FEC895E8FE1EF943" box="[371,439,1694,1720]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="110" pageNumber="111">ANIC</collectionCode>
Berlesate No. 667, litter under
<taxonomicName id="40F851E1FFB7AD53FCA595E8FC59F943" box="[798,1008,1694,1720]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="110" pageNumber="111" phylum="Tracheophyta" rank="species" species="gunnii">
<emphasis id="B58CF670FFB7AD53FCA595E8FC59F943" box="[798,1008,1694,1720]" italics="true" pageId="110" pageNumber="111">Nothofagus gunnii</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="40F851E1FFB7AD53FBAE95E8FAB5F943" box="[1045,1308,1694,1720]" class="Pinopsida" family="Cupressaceae" genus="Athrotaxis" kingdom="Plantae" order="Pinales" pageId="110" pageNumber="111" phylum="Tracheophyta" rank="species" species="selaginoides">
<emphasis id="B58CF670FFB7AD53FBAE95E8FAB5F943" box="[1045,1308,1694,1720]" italics="true" pageId="110" pageNumber="111">Athrotaxis selaginoides</emphasis>
</taxonomicName>
,
<date id="F3460CA2FFB7AD53FA9095E8FA3EF943" box="[1323,1431,1694,1720]" pageId="110" pageNumber="111" value="1980-01-31">31.i.1980</date>
, Lawrence &amp; Weir (
<specimenCount id="91FEE1EBFFB7AD53FECA95B4FE30F927" box="[369,409,1730,1756]" pageId="110" pageNumber="111" type="female">1 ♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFB7AD53FE1C95B4FE51F927" box="[423,504,1730,1756]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="110" pageNumber="111">DEBU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB7AD4DFF7C9593FB90FEE6" lastPageId="112" lastPageNumber="113" pageId="110" pageNumber="111" type="discussion">
<paragraph id="87472A62FFB7AD53FF7C9593FEFDF823" blockId="110.[151,1437,325,2008]" pageId="110" pageNumber="111">
<emphasis id="B58CF670FFB7AD53FF7C9593FEE6F904" bold="true" box="[199,335,1765,1791]" pageId="110" pageNumber="111">Comments.</emphasis>
The redescription of male terminalia is based largely on Richards’ (1973) description and illustrations. The only apparent male specimen (identified previously) available for this work was missing its abdomen, but a written description of the postabdomen by a previous student noted a clearly tripartite hypandrium but without further details. The only female specimen available for examination had its postabdomen retracted, making it impossible to evert the postabdomen without risking damaging or destroying the postabdomen, even after clearing, and so could not be illustrated. Some features were visible through the preabdominal sclerites and are described above.
</paragraph>
<caption id="D3877AEAFFB6AD52FF2394E1FE4FF803" ID-DOI="http://doi.org/10.5281/zenodo.7138839" ID-Zenodo-Dep="7138839" httpUri="https://zenodo.org/record/7138839/files/figure.png" pageId="111" pageNumber="112" startId="111.[152,270,1943,1967]" targetBox="[156,1431,181,1902]" targetPageId="111">
<paragraph id="87472A62FFB6AD52FF2394E1FE4FF803" blockId="111.[151,1437,1943,2040]" pageId="111" pageNumber="112">
<emphasis id="B58CF670FFB6AD52FF2394E1FE28F854" bold="true" box="[152,385,1943,1967]" pageId="111" pageNumber="112">FIGURES 32.1–32.5.</emphasis>
<taxonomicName id="40F851E1FFB6AD52FE3294EEFD1AF84B" authority="(Richards)" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" baseAuthorityName="Kuwahara &amp; Marshall" baseAuthorityYear="2022" box="[393,691,1944,1968]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="dealata">
<emphasis id="B58CF670FFB6AD52FE3294EEFD94F84B" box="[393,573,1944,1968]" italics="true" pageId="111" pageNumber="112">Howickia dealata</emphasis>
(Richards)
</taxonomicName>
. 32.1, male habitus, lateral; 32.2, male head, anterior; 32.3, epiproct and cercus, dorsal; 32.4, S8 and hypoproct, ventral; 32.5, spermathecae. Body scale bar indicates 0.25 mm, head bar indicates 0.1 mm. Photo credit: G. Thompson.
</paragraph>
</caption>
<paragraph id="87472A62FFA9AD4DFF7C93E1FB90FEE6" blockId="112.[151,1437,151,285]" pageId="112" pageNumber="113">
One of two species originally described as
<taxonomicName id="40F851E1FFA9AD4DFD1393E1FC96FF4A" authorityName="Richards" authorityYear="1973" box="[680,831,151,177]" class="Insecta" family="Sphaeroceridae" genus="Monteithiana" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFA9AD4DFD1393E1FC96FF4A" box="[680,831,151,177]" italics="true" pageId="112" pageNumber="113">Monteithiana</emphasis>
</taxonomicName>
,
<taxonomicName id="40F851E1FFA9AD4DFCF693E1FC6DFF4A" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" baseAuthorityName="Kuwahara &amp; Marshall" baseAuthorityYear="2022" box="[845,964,151,177]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="dealata">
<emphasis id="B58CF670FFA9AD4DFCF693E1FC6DFF4A" box="[845,964,151,177]" italics="true" pageId="112" pageNumber="113">H. dealata</emphasis>
</taxonomicName>
, has mid tibial chaetotaxy similar to other wingless
<taxonomicName id="40F851E1FFA9AD4DFEB993CDFEC4FF2E" authorityName="Richards" authorityYear="1951" box="[258,365,187,213]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFA9AD4DFEB993CDFEC4FF2E" box="[258,365,187,213]" italics="true" pageId="112" pageNumber="113">Howickia</emphasis>
</taxonomicName>
(paired bristles at one-third and four-fifths length), stout apical seta on the surstylus, male S5 with a posteromedial row of flattened teeth, and the tripartite hypandrium that defines
<taxonomicName id="40F851E1FFA9AD4DFB3D93A9FB58FF02" authorityName="Richards" authorityYear="1951" box="[1158,1265,223,249]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFA9AD4DFB3D93A9FB58FF02" box="[1158,1265,223,249]" italics="true" pageId="112" pageNumber="113">Howickia</emphasis>
</taxonomicName>
. We here treat
<taxonomicName id="40F851E1FFA9AD4DFF2C9275FE87FEE6" authorityName="Richards" authorityYear="1973" box="[151,302,259,285]" class="Insecta" family="Sphaeroceridae" genus="Monteithiana" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFA9AD4DFF2C9275FE87FEE6" box="[151,302,259,285]" italics="true" pageId="112" pageNumber="113">Monteithiana</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="40F851E1FFA9AD4DFE489275FDF7FEE6" authorityName="Richards" authorityYear="1951" box="[499,606,259,285]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFA9AD4DFE489275FDF7FEE6" box="[499,606,259,285]" italics="true" pageId="112" pageNumber="113">Howickia</emphasis>
</taxonomicName>
(see comments on
<taxonomicName id="40F851E1FFA9AD4DFC8D9275FB9CFEE6" authority="(Richards))" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" baseAuthorityName="Kuwahara &amp; Marshall" baseAuthorityYear="2022" box="[822,1077,259,285]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="cynthia">
<emphasis id="B58CF670FFA9AD4DFC8D9275FC03FEE6" box="[822,938,259,285]" italics="true" pageId="112" pageNumber="113">H. cynthia</emphasis>
(Richards))
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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