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<document id="FC39B8DB799973643799DE6FC8D64799" ID-CLB-Dataset="157781" ID-DOI="10.11646/zootaxa.5192.1.1" ID-GBIF-Dataset="8e2cc009-d580-4904-a0b7-9ed7a9e3ab9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138639" ID-ZooBank="236A609B-8A6A-47D7-9BB9-E2FDCA5C37E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664794801684" checkinUser="plazi" docAuthor="Kuwahara, Gregory K. &amp; Marshall, Stephen A." docDate="2022" docId="0F519B74FF87AD5CFF2C97DCFB83FC32" docLanguage="en" docName="zootaxa.5192.1.1.pdf" docOrigin="Zootaxa 5192 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5192.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Howickia cruspica Kuwahara &amp; Marshall 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="98" masterDocId="F368E30CFFD9AD3DFFBB9376FFA9FFFB" masterDocTitle="A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)" masterLastPageNumber="152" masterPageNumber="1" pageNumber="95" updateTime="1699386905592" updateUser="ExternalLinkService">
<mods:mods id="00611B52BBAA1F188121BDF438D92B9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B29A72BB397102A52E6C8EFC2808953">
<mods:title id="28ADC1510D48BCB9836954BA06A4167E">A revision of the Australian species of Howickia Richards (Diptera: Sphaeroceridae: Limosininae)</mods:title>
</mods:titleInfo>
<mods:name id="77B11F5A54FA21D897611B3A67ADC896" type="personal">
<mods:role id="0DC62F74EF9EAEC99BEF5D2F243B4FA8">
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<mods:namePart id="346ADA58B3A84306D46B2A048234D348">Kuwahara, Gregory K.</mods:namePart>
<mods:nameIdentifier id="021D79B52DC8631CD6451E7DF246A6E1" type="ORCID">0000-0002-0319-2672</mods:nameIdentifier>
<mods:nameIdentifier id="A2F526BA6429047D345781C79F26B3F6" type="email">gkuwahar@uoguelph.ca</mods:nameIdentifier>
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<mods:namePart id="0D7743052EBF57E44F92A5352838B9EB">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="225652D0D6EC0268FB9EB8191748D1E9" type="ORCID">0000-0002-5732-9718</mods:nameIdentifier>
<mods:nameIdentifier id="1B3DA11C0B9705E016BB04FA6B2BC497" type="email">samarsha@uoguelph.ca</mods:nameIdentifier>
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<mods:title id="550D985CC54E739B42719AEB1B6EA1C9">Zootaxa</mods:title>
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<mods:part id="A3B488D6709B915EB5D030B753AB4395">
<mods:date id="6AFDDDA2F80AB8D49FD43C326A8F699B">2022</mods:date>
<mods:detail id="B5C0564E8D4EDBCB6CB9AE7ED03ABEA0" type="pubDate">
<mods:number id="12B94C17B4339AAC0C6FDFC519143A11">2022-10-03</mods:number>
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<mods:detail id="3EA3955915DE435004E2897058425585" type="volume">
<mods:number id="9681A0BC7F493B481127F0E38F2D687D">5192</mods:number>
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<mods:end id="99F94C1DB6238CA006370F2DED37C0ED">152</mods:end>
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<mods:url id="36450B87C44E88807C1D14FCDFBE1FEB">http://dx.doi.org/10.11646/zootaxa.5192.1.1</mods:url>
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<mods:classification id="1588F1CDAA1396FE10ADECF14D0AD863">journal article</mods:classification>
<mods:identifier id="AD8288613656520F75F7386D27C57B63" type="CLB-Dataset">157781</mods:identifier>
<mods:identifier id="A6A4C3B0981A55AAF75D243B428D8CC2" type="DOI">10.11646/zootaxa.5192.1.1</mods:identifier>
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<treatment id="0F519B74FF87AD5CFF2C97DCFB83FC32" ID-DOI="http://doi.org/10.5281/zenodo.7138758" ID-GBIF-Taxon="202134585" ID-Zenodo-Dep="7138758" LSID="urn:lsid:plazi:treatment:0F519B74FF87AD5CFF2C97DCFB83FC32" httpUri="http://treatment.plazi.org/id/0F519B74FF87AD5CFF2C97DCFB83FC32" lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95">
<subSubSection id="CFE279E9FF87AD63FF2C97DCFE77FB3E" box="[151,478,1194,1221]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="87472A62FF87AD63FF2C97DCFE77FB3E" blockId="94.[151,478,1194,1257]" box="[151,478,1194,1221]" pageId="94" pageNumber="95">
<heading id="DC0F9D0EFF87AD63FF2C97DCFE77FB3E" bold="true" box="[151,478,1194,1221]" fontSize="11" level="2" pageId="94" pageNumber="95" reason="4">
<emphasis id="B58CF670FF87AD63FF2C97DCFE77FB3E" bold="true" box="[151,478,1194,1221]" pageId="94" pageNumber="95">
<taxonomicName id="40F851E1FF87AD63FF2C97DCFED0FB3E" ID-CoL="BD3KT" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[151,377,1194,1221]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="cruspica" status="sp. nov.">
<emphasis id="B58CF670FF87AD63FF2C97DCFED0FB3E" bold="true" box="[151,377,1194,1221]" italics="true" pageId="94" pageNumber="95">Howickia cruspica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEBF4B0BFF87AD63FE3B97DDFE77FB3E" box="[384,478,1195,1221]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FF87AD5CFF2C97B9FB64FE2A" lastPageId="97" lastPageNumber="98" pageId="94" pageNumber="95" type="description">
<paragraph id="87472A62FF87AD63FF2C97B9FEF1FB12" blockId="94.[151,478,1194,1257]" box="[151,344,1230,1257]" pageId="94" pageNumber="95">
(
<figureCitation id="1FC336E7FF87AD63FF2497B9FEF9FB12" box="[159,336,1230,1257]" captionStart="FIGURES 26.1–26.2" captionStartId="95.[152,270,1933,1957]" captionTargetBox="[170,1418,181,1880]" captionTargetId="figure-19@95.[170,1418,181,1909]" captionTargetPageId="95" captionText="FIGURES 26.1–26.2. Howickia cruspica sp. nov. 26.1, female habitus, lateral; 26.2, female head, anterolateral. Scale bars indicate 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7138805" httpUri="https://zenodo.org/record/7138805/files/figure.png" pageId="94" pageNumber="95">Figs. 26.1–26.5</figureCitation>
)
</paragraph>
<paragraph id="87472A62FF87AD63FF2C9660FDD4FACA" blockId="94.[151,1437,1302,2019]" box="[151,637,1302,1329]" pageId="94" pageNumber="95">
<emphasis id="B58CF670FF87AD63FF2C9660FE80FACB" bold="true" box="[151,297,1302,1328]" pageId="94" pageNumber="95">Description.</emphasis>
<emphasis id="B58CF670FF87AD63FE8B9661FE69FACA" box="[304,448,1303,1329]" italics="true" pageId="94" pageNumber="95">Body length:</emphasis>
Female
<quantity id="40008787FF87AD63FD9B9661FDD0FACA" box="[544,633,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="94" pageNumber="95" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
<paragraph id="87472A62FF87AD63FF7C964DFCFDF9D6" blockId="94.[151,1437,1302,2019]" pageId="94" pageNumber="95">
<emphasis id="B58CF670FF87AD63FF7C964DFEA4FAAE" box="[199,269,1339,1365]" italics="true" pageId="94" pageNumber="95">Head:</emphasis>
Yellow, areas between frontal vitta and orbital plates orange-yellow; frontal vitta striate and shiny, orbital plates microtrichose; gena yellow; antenna yellow, apically pale. Frontal width 1.5x interfrontal height, frontal vitta narrow. Two pairs of large interfrontal bristles preceded by a smaller pair (0.6x length of posterior pairs); 2 lateroclinate orbital bristles, anterior bristle 0.6x length of posterior bristle. Gena with large upcurved bristle and 2 smaller setae, finely and densely striate, shiny, lower third microtrichose; vibrissal angle with 2 very fine subvibrissal setae. Face shiny with a very small flattened triangle along ventral margin; palpus clavate with 1 large apical and 1 large preapical seta. Eye height 2.0x genal height.
</paragraph>
<paragraph id="87472A62FF87AD63FF7C9541FC23F91A" blockId="94.[151,1437,1302,2019]" pageId="94" pageNumber="95">
<emphasis id="B58CF670FF87AD63FF7C9541FE88F9AA" box="[199,289,1591,1617]" italics="true" pageId="94" pageNumber="95">Thorax:</emphasis>
Dark brown; scutum shiny despite microtrichia, posterolateral corners yellow-brown and with pale spot on posterior notopleural lobe; posterodorsal corners of anepisternum and katepisternum lighter brown, katatergite black. Two pairs of dorsocentral bristles (anterior pair 0.6x length of posterior pair); 6–7 rows of long, fine acrostichal setulae. Prosternum narrow, linear, posteriorly expanded. Scutellum microtrichose, subquadrate, its length 0.5x its width; 4 scutellar bristles long, apical pair 1.5x length of basal pair.
</paragraph>
<paragraph id="87472A62FF87AD63FF7C959DFD45F88A" blockId="94.[151,1437,1302,2019]" pageId="94" pageNumber="95">
<emphasis id="B58CF670FF87AD63FF7C959DFEAFF8FF" box="[199,262,1771,1796]" italics="true" pageId="94" pageNumber="95">Legs:</emphasis>
Brown, fore and mid femora caramel-brown and distally paler, hind femur dark brown, tibiae paler than femora; tarsi dirty-white. Mid basitarsus with a distinctly enlarged ventral seta; hind femur with a large curved apicoventral spur. Basal half of mid tibia with 5 dorsal bristles (3 anterodorsal and 2 posterodorsal), distal half with 3 dorsal bristles (2 anterodorsal and 1 posterodorsal).
</paragraph>
<paragraph id="87472A62FF87AD63FF7C940DFBC0F826" blockId="94.[151,1437,1302,2019]" pageId="94" pageNumber="95">
<emphasis id="B58CF670FF87AD63FF7C940DFEFCF86E" box="[199,341,1915,1941]" italics="true" pageId="94" pageNumber="95">Wing/halter:</emphasis>
Wings significantly reduced, extending to about midpoint of T4; veins reduced but all identifiable: CS2 0.5x length of CS3; R
<subScript id="1B7C2827FF87AD63FE0594C7FE7EF844" attach="left" box="[446,471,1969,1983]" fontSize="6" pageId="94" pageNumber="95">2+3</subScript>
sinuate, meeting costa above crossvein
<emphasis id="B58CF670FF87AD63FC2A94E9FC1FF843" box="[913,950,1951,1976]" italics="true" pageId="94" pageNumber="95">i-m</emphasis>
; R
<subScript id="1B7C2827FF87AD63FC6C94C7FC58F844" attach="left" box="[983,1009,1969,1983]" fontSize="6" pageId="94" pageNumber="95">4+5</subScript>
strongly upcurved, meeting costa well before wing tip; costa slightly overpassing R
<subScript id="1B7C2827FF87AD63FD3794A3FD0FF818" attach="left" box="[652,678,2005,2019]" fontSize="6" pageId="94" pageNumber="95">4+5</subScript>
. Halter slightly reduced and pure white.
</paragraph>
<caption id="D3877AEAFF86AD62FF2394FBFEEBF832" ID-DOI="http://doi.org/10.5281/zenodo.7138805" ID-Zenodo-Dep="7138805" httpUri="https://zenodo.org/record/7138805/files/figure.png" pageId="95" pageNumber="96" startId="95.[152,270,1933,1957]" targetBox="[170,1418,181,1880]" targetPageId="95">
<paragraph id="87472A62FF86AD62FF2394FBFEEBF832" blockId="95.[151,1436,1933,1993]" pageId="95" pageNumber="96">
<emphasis id="B58CF670FF86AD62FF2394FBFE2BF85E" bold="true" box="[152,386,1933,1957]" pageId="95" pageNumber="96">FIGURES 26.1–26.2.</emphasis>
<taxonomicName id="40F851E1FF86AD62FE3094FBFDE2F85E" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[395,587,1933,1957]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="cruspica" status="sp. nov.">
<emphasis id="B58CF670FF86AD62FE3094FBFDE2F85E" box="[395,587,1933,1957]" italics="true" pageId="95" pageNumber="96">Howickia cruspica</emphasis>
</taxonomicName>
<emphasis id="B58CF670FF86AD62FDEF94FBFD0FF85E" bold="true" box="[596,678,1933,1957]" pageId="95" pageNumber="96">
<taxonomicNameLabel id="AEBF4B0BFF86AD62FDEF94FBFD0FF85E" box="[596,678,1933,1957]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
26.1, female habitus, lateral; 26.2, female head, anterolateral. Scale bars indicate 0.5 mm.
</paragraph>
</caption>
<caption id="D3877AEAFFB9AD5DFF239422FB8DF86B" ID-DOI="http://doi.org/10.5281/zenodo.7138807" ID-Zenodo-Dep="7138807" httpUri="https://zenodo.org/record/7138807/files/figure.png" pageId="96" pageNumber="97" startId="96.[152,270,1876,1900]" targetBox="[165,1419,197,1853]" targetPageId="96">
<paragraph id="87472A62FFB9AD5DFF239422FB8DF86B" blockId="96.[151,1437,1876,1936]" pageId="96" pageNumber="97">
<emphasis id="B58CF670FFB9AD5DFF239422FE2BF897" bold="true" box="[152,386,1876,1900]" pageId="96" pageNumber="97">FIGURES 26.3–26.5.</emphasis>
<taxonomicName id="40F851E1FFB9AD5DFE309422FDE5F897" authority="Kuwahara &amp; Marshall, 2022" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[395,588,1876,1900]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="cruspica" status="sp. nov.">
<emphasis id="B58CF670FFB9AD5DFE309422FDE5F897" box="[395,588,1876,1900]" italics="true" pageId="96" pageNumber="97">Howickia cruspica</emphasis>
</taxonomicName>
<emphasis id="B58CF670FFB9AD5DFDEE9423FD01F896" bold="true" box="[597,680,1877,1901]" pageId="96" pageNumber="97">
<taxonomicNameLabel id="AEBF4B0BFFB9AD5DFDEE9423FD01F896" box="[597,680,1877,1901]" pageId="96" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
26.3, left wing; 26.4, female terminalia, dorsal; 26.5, female terminalia, ventral. Female cerci were excluded from ventral illustration. Scale bar indicates 0.5 mm.
</paragraph>
</caption>
<paragraph id="87472A62FFB8AD5CFF7C93E1FD87FF4A" blockId="97.[151,1437,151,969]" box="[199,558,151,177]" pageId="97" pageNumber="98">
<emphasis id="B58CF670FFB8AD5CFF7C93E1FEC6FF4A" box="[199,367,151,177]" italics="true" pageId="97" pageNumber="98">Male abdomen</emphasis>
: Male unknown.
</paragraph>
<paragraph id="87472A62FFB8AD5CFF7C93CDFB64FE2A" blockId="97.[151,1437,151,969]" pageId="97" pageNumber="98">
<emphasis id="B58CF670FFB8AD5CFF7C93CDFE3AFF2E" box="[199,403,187,213]" italics="true" pageId="97" pageNumber="98">Female abdomen:</emphasis>
Black, shiny despite dense microtrichia. T2–5 and S2–5 heavily sclerotized, sparsely longsetose; pleural membrane with sparse setae. Preabdomen slightly shorter than length of head + thorax. Postabdomen largely retracted into preabdomen, with only last few segments exposed, broad as far as can be seen and likely 3.0x length of T5. T6 and S6 hidden; T7 and S7 reduced to pairs of L-shaped plates, each bearing a posterior row of 2–3 setae. Medial plate of T8 appears to be absent or greatly reduced. Epiproct elongate, rhombic, centrally split into two equal lateral halves, each bearing a dorsal seta. Cercus elongate rectangular with 3–4 long sinuate setae (2 apical, 1 lateral and 1 dorsal) and several smaller setulae. S8 a broad, rectangular, posteriorly microtrichose plate with a posterior row of setae. Hypoproct broad but short, posteriorly microtrichose with a row of setae.
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB8AD5CFF7C92ACFEFCFDE2" pageId="97" pageNumber="98" type="materials_examined">
<paragraph id="87472A62FFB8AD5CFF7C92ACFEFCFDE2" blockId="97.[151,1437,151,969]" pageId="97" pageNumber="98">
<emphasis id="B58CF670FFB8AD5CFF7C92ACFE19FE0F" bold="true" box="[199,432,474,500]" pageId="97" pageNumber="98">Material examined.</emphasis>
<materialsCitation id="3790203FFFB8AD5CFE0C92ADFEFCFDE2" ID-GBIF-Occurrence="3920718383" collectingDate="1978-04-28" collectingDateMax="1978-05-07" collectingDateMin="1978-04-28" collectionCode="ANIC" collectorName="S. Peck" country="Australia" elevation="200" location="Cement Creek" municipality="Warburton" pageId="97" pageNumber="98" specimenCount="1" specimenCount-female="1" stateProvince="Victoria" typeStatus="holotype">
<typeStatus id="584394C0FFB8AD5CFE0C92ADFD8DFE0E" box="[439,548,475,501]" pageId="97" pageNumber="98" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="FFEF6AF2FFB8AD5CFD9792ADFD66FE0E" box="[556,719,475,501]" name="Australia" pageId="97" pageNumber="98">AUSTRALIA</collectingCountry>
:
<emphasis id="B58CF670FFB8AD5CFD6292ACFCEAFE0F" bold="true" box="[729,835,474,500]" pageId="97" pageNumber="98">
<collectingRegion id="453CE480FFB8AD5CFD6292ACFC97FE0F" box="[729,830,474,500]" country="Australia" name="Victoria" pageId="97" pageNumber="98">Victoria</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6723B018FFB8AD5CFCF192ADFC6DFE0E" box="[842,964,475,501]" pageId="97" pageNumber="98">Warburton</collectingMunicipality>
,
<location id="82277CB9FFB8AD5CFC7492ADFBDBFE0E" LSID="urn:lsid:plazi:treatment:0F519B74FF87AD5CFF2C97DCFB83FC32:82277CB9FFB8AD5CFC7492ADFBDBFE0E" box="[975,1138,475,501]" country="Australia" municipality="Warburton" name="Cement Creek" pageId="97" pageNumber="98" stateProvince="Victoria">Cement Creek</location>
,
<quantity id="40008787FFB8AD5CFBC692ADFB6FFE0E" box="[1149,1222,475,501]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="97" pageNumber="98" unit="m" value="200.0">
<elevation id="0CD5CD51FFB8AD5CFBC692ADFB6FFE0E" box="[1149,1222,475,501]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="97" pageNumber="98" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="F3460CA2FFB8AD5CFB6A92ADFAD2FE0E" box="[1233,1403,475,501]" pageId="97" pageNumber="98" value="1978-04-28" valueMax="1978-05-07" valueMin="1978-04-28">
<collectingDate id="E302F54AFFB8AD5CFB6A92ADFAD2FE0E" box="[1233,1403,475,501]" pageId="97" pageNumber="98" value="1978-04-28" valueMax="1978-05-07" valueMin="1978-04-28">28.iv–7.v.1978</collectingDate>
</date>
,
<collectorName id="2A0D4FB4FFB8AD5CFA3D92ADFF64FDE2" pageId="97" pageNumber="98">S. Peck</collectorName>
(
<specimenCount id="91FEE1EBFFB8AD5CFF5B9289FF5AFDE2" box="[224,243,511,537]" pageId="97" pageNumber="98" type="female">♀</specimenCount>
,
<collectionCode id="E1E9B2A7FFB8AD5CFF449289FEE0FDE2" box="[255,329,511,537]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="97" pageNumber="98">ANIC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB8AD5CFF7C9155FE68FD9A" pageId="97" pageNumber="98" type="etymology">
<paragraph id="87472A62FFB8AD5CFF7C9155FE68FD9A" blockId="97.[151,1437,151,969]" pageId="97" pageNumber="98">
<emphasis id="B58CF670FFB8AD5CFF7C9155FEE7FDC6" bold="true" box="[199,334,547,573]" pageId="97" pageNumber="98">Etymology.</emphasis>
This species name refers to the enlarged apicoventral hind tibial spur (Latin ‘crus’: leg; Latin ‘spica’: spike, dart, spear).
</paragraph>
</subSubSection>
<subSubSection id="CFE279E9FFB8AD5CFF7C911CFB83FC32" pageId="97" pageNumber="98" type="discussion">
<paragraph id="87472A62FFB8AD5CFF7C911CFC4AFCC2" blockId="97.[151,1437,151,969]" pageId="97" pageNumber="98">
<emphasis id="B58CF670FFB8AD5CFF7C911CFEE6FD7F" bold="true" box="[199,335,618,644]" pageId="97" pageNumber="98">Comments.</emphasis>
<taxonomicName id="40F851E1FFB8AD5CFEE2911DFD86FD7F" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[345,559,619,645]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="cruspica">
<emphasis id="B58CF670FFB8AD5CFEE2911DFD86FD7F" box="[345,559,619,645]" italics="true" pageId="97" pageNumber="98">Howickia cruspica</emphasis>
</taxonomicName>
is known from a single female specimen but is easily distinguishable from all other species of
<taxonomicName id="40F851E1FFB8AD5CFECC91F9FE4BFD52" authorityName="Richards" authorityYear="1951" box="[375,482,655,681]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFB8AD5CFECC91F9FE4BFD52" box="[375,482,655,681]" italics="true" pageId="97" pageNumber="98">Howickia</emphasis>
</taxonomicName>
. This species was included in
<taxonomicName id="40F851E1FFB8AD5CFCFA91F9FC05FD52" authorityName="Richards" authorityYear="1951" box="[833,940,655,681]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFB8AD5CFCFA91F9FC05FD52" box="[833,940,655,681]" italics="true" pageId="97" pageNumber="98">Howickia</emphasis>
</taxonomicName>
due to synapomorphies shared with known species of
<taxonomicName id="40F851E1FFB8AD5CFEB491C5FED3FD36" authorityName="Richards" authorityYear="1951" box="[271,378,691,717]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFB8AD5CFEB491C5FED3FD36" box="[271,378,691,717]" italics="true" pageId="97" pageNumber="98">Howickia</emphasis>
</taxonomicName>
(mid tibia with paired proximal and distal dorsal bristles, sinuate R
<subScript id="1B7C2827FFB8AD5CFBCE91B3FB27FD28" attach="left" box="[1141,1166,709,723]" fontSize="6" pageId="97" pageNumber="98">2+3</subScript>
, curved R
<subScript id="1B7C2827FFB8AD5CFABF91B3FAB7FD28" attach="left" box="[1284,1310,709,723]" fontSize="6" pageId="97" pageNumber="98">4+5</subScript>
, telescopic female terminalia with reduced postabdominal sclerites). Without a male specimen of
<taxonomicName id="40F851E1FFB8AD5CFBDB91A1FB4AFD0B" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[1120,1251,727,752]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="cruspica">
<emphasis id="B58CF670FFB8AD5CFBDB91A1FB4AFD0B" box="[1120,1251,727,752]" italics="true" pageId="97" pageNumber="98">H. cruspica</emphasis>
</taxonomicName>
, it is impossible to determine whether this species has the defining hypandrial characters of
<taxonomicName id="40F851E1FFB8AD5CFC63918DFBEAFCEE" authorityName="Richards" authorityYear="1951" box="[984,1091,763,789]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFB8AD5CFC63918DFBEAFCEE" box="[984,1091,763,789]" italics="true" pageId="97" pageNumber="98">Howickia</emphasis>
</taxonomicName>
, but the synapomorphies listed above provide some support for the placement of
<taxonomicName id="40F851E1FFB8AD5CFD7F9069FCEFFCC3" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[708,838,799,824]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="cruspica">
<emphasis id="B58CF670FFB8AD5CFD7F9069FCEFFCC3" box="[708,838,799,824]" italics="true" pageId="97" pageNumber="98">H. cruspica</emphasis>
</taxonomicName>
in this genus.
</paragraph>
<paragraph id="87472A62FFB8AD5CFF7C9035FB83FC32" blockId="97.[151,1437,151,969]" pageId="97" pageNumber="98">
<taxonomicName id="40F851E1FFB8AD5CFF7C9035FE30FCA7" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" box="[199,409,835,861]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="cruspica">
<emphasis id="B58CF670FFB8AD5CFF7C9035FE30FCA7" box="[199,409,835,861]" italics="true" pageId="97" pageNumber="98">Howickia cruspica</emphasis>
</taxonomicName>
has the largest wing of the three brachypterous species of
<taxonomicName id="40F851E1FFB8AD5CFB939035FB3AFCA6" authorityName="Richards" authorityYear="1951" box="[1064,1171,835,861]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFB8AD5CFB939035FB3AFCA6" box="[1064,1171,835,861]" italics="true" pageId="97" pageNumber="98">Howickia</emphasis>
</taxonomicName>
, and is the only species of
<taxonomicName id="40F851E1FFB8AD5CFF0C9011FE8BFC7A" authorityName="Richards" authorityYear="1951" box="[183,290,871,897]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B58CF670FFB8AD5CFF0C9011FE8BFC7A" box="[183,290,871,897]" italics="true" pageId="97" pageNumber="98">Howickia</emphasis>
</taxonomicName>
to have an enlarged apicoventral spur on the hind tibia. The only other brachypterous species known from VIC,
<taxonomicName id="40F851E1FFB8AD5CFEAD90FDFD37FC5E" authority="(Richards, 1973)" authorityName="Kuwahara &amp; Marshall" authorityYear="2022" baseAuthorityName="Kuwahara &amp; Marshall" baseAuthorityYear="2022" box="[278,670,907,933]" class="Insecta" family="Sphaeroceridae" genus="Howickia" kingdom="Animalia" order="Diptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="sabina">
<emphasis id="B58CF670FFB8AD5CFEAD90FDFE7CFC5E" box="[278,469,907,933]" italics="true" pageId="97" pageNumber="98">Howickia sabina</emphasis>
(
<bibRefCitation id="E3695793FFB8AD5CFE5D90FDFD3FFC5E" author="Richards, O. W." box="[486,662,907,933]" pageId="97" pageNumber="98" pagination="297 - 401" refId="ref94252" refString="Richards, O. W. (1973) The Sphaeroceridae (= Borboridae or Cypselidae; Diptera Cyclorrhapha) of the Australian region. Australian Journal of Zoology Supplemental Series, 22, 297 - 401. https: // doi. org / 10.1071 / AJZS 022" type="journal article" year="1973">Richards, 1973</bibRefCitation>
)
</taxonomicName>
, has a striped frons, a shorter gena, smaller wings, no apicoventral spur on the hind tibia, and only two dorsal setae on the basal half of the mid tibia.
</paragraph>
</subSubSection>
</treatment>
</document>
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