261 lines
26 KiB
XML
261 lines
26 KiB
XML
<document id="4A0F8D4BB82838BA5997E615AC8747A3" ENCODING="Cp1252" ID-CLB-Dataset="51824" ID-DOI="http://dx.doi.org/10.11646/zootaxa.3721.3.5" ID-GBIF-Dataset="d1a65891-6077-4227-aa28-98049e13ce30" checkinTime="1416762300902" checkinUser="christiana" docAuthor="GARY D. ALPERT" docDate="2013" docId="FE5FC8D22726EF334D9974E720448123" docLanguage="en" docName="zt03721p290_nurText.htm" docOrigin="Zootaxa 3721 (3)" docTitle="Tyrannomyrmex legatus ALPERT, 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="290" masterDocId="12B9756F83B94827C3D33695C9DB40DB" masterDocTitle="A new species of Tyrannomyrmex (Hymenoptera: Formicidae) from Sri Lanka" masterLastPageNumber="290" masterPageNumber="286" pageNumber="287" updateTime="1732856163590" updateUser="ExternalLinkService">
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<mods:mods id="84014E30758B358074D2EC788F3A1787" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A3479F62FC880E66F2C8934DAC733ACC">
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<mods:title id="0B9038FFD51AC3E06F63B1ED223D2084">A new species of Tyrannomyrmex (Hymenoptera: Formicidae) from Sri Lanka</mods:title>
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</mods:titleInfo>
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<mods:name id="6D5CEA502015159C60502697F2FD98AC" type="personal">
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<mods:role id="B8E0C5860A4A79FBB08992992F265889">
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<mods:roleTerm id="89F46B7DB4DDCDCA5CC713DDE966ACA9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="446DB2B9591767EE920FA80B51F05505">GARY D. ALPERT</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4E73AF15FF793AADE111CD65CAC0F47A">text</mods:typeOfResource>
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<mods:relatedItem id="526C86FF98208554190990C16105AA12" type="host">
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<mods:titleInfo id="3682924640A65A6D2547FB57BA05FFD4">
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<mods:title id="83E4F5BD0A17E7CCD8B3BE462C29581B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4C474E489FAFFF20EACE0CF76F0F67AD">
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<mods:date id="38460517C563D6C88A15D2CE07C11804">2013</mods:date>
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<mods:detail id="4DA09617C3805C28CFC5D3D6587A6295" type="volume">
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<mods:number id="288868EB7ADBB9B51168D341058D52F2">3721</mods:number>
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</mods:detail>
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<mods:detail id="D763DBE80B54E46984A68DFD33666327" type="issue">
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<mods:number id="44CBC7F6ACB01D75EA4032B7FBB4D2A1">3</mods:number>
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</mods:detail>
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<mods:extent id="1FA11345980C5645FD3803FE5A216D87" unit="page">
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<mods:start id="7D7FE036A66E4E30337EFC6EBB457750">286</mods:start>
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<mods:end id="07EC84F0972B127CF5BCA00A4AAC10FB">290</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="610ABF7CC7304275D3FEA133170F8042">journal article</mods:classification>
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<mods:identifier id="90A88966806F1AD46563A2EA262E3294" type="DOI">http://dx.doi.org/10.11646/zootaxa.3721.3.5</mods:identifier>
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</mods:mods>
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<treatment id="FE5FC8D22726EF334D9974E720448123" ID-DOI="http://doi.org/10.5281/zenodo.5640953" ID-GBIF-Taxon="114926881" ID-Zenodo-Dep="5640953" LSID="urn:lsid:plazi:treatment:FE5FC8D22726EF334D9974E720448123" httpUri="http://treatment.plazi.org/id/FE5FC8D22726EF334D9974E720448123" lastPageId="4" lastPageNumber="290" pageId="1" pageNumber="287" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="DEFD1B85F22AE84A3F92430760CB568F" pageId="1" pageNumber="287" type="nomenclature">
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<paragraph id="ABFF64E26325B19079F54FEC2EFBAF24" pageId="1" pageNumber="287" type="mainText">
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<taxonomicName id="7FBC68B1723EBDB505D432D91BE7FE49" ID-CoL="5BSZ" genus="Tyrannomyrmex" pageId="1" pageNumber="287" rank="species" species="legatus">Tyrannomyrmex legatus</taxonomicName>
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<taxonomicNameLabel id="3B679C8C5AA73C98D4776DA35F3C5347" pageId="1" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="062B90338C42148417536A3BCE42B308" pageId="1" pageNumber="287" type="description">
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<paragraph id="FB87FF92E1393C5B4E8A037E907B9DE3" pageId="1" pageNumber="287" type="mainText">(Figs. 1-3)</paragraph>
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</subSubSection>
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<subSubSection id="06AEA38B18AB7C4EC8573E058851A743" pageId="1" pageNumber="287" type="materials_examined">
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<paragraph id="F36A60D9ED1209AFEE4B75FB906E2510" pageId="1" pageNumber="287" type="mainText">
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<materialsCitation id="8BF0AC5198DF34B899789884847F5B18" ID-GBIF-Occurrence="3396408301" collectingDate="2013-01-01" collectingDateMax="2013-12-31" collectingDateMin="2013-01-01" collectingMethod="CTFS Plot Winkler sacks 15 - 18" collectionCode="MCZ" collectorName="N. R. Gunawardene" country="Sri Lanka" elevation="432 m- 571 m," httpUri="http://www.antweb.org/specimen/CASENT0106177" latitude="6.413717" location="Sinharaja Forest Reserve" longitude="80.41652" specimenCode="number 35624, CASENT 0106177, ANTC 4038" specimenCount="1" stateProvince="Sinharaja Forest Reserve" typeStatus="holotype">
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<typeStatus id="F58F185CEEDB2F828602984D39390AEB">Holotype</typeStatus>
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worker.
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<collectingCountry id="023CC08AECB95B54E9AB0694D6B59083" name="Sri Lanka" value="Sri Lanka">Sri Lanka</collectingCountry>
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,
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<collectingRegion id="59EA803AF4E36759FEB18B7315CF547E">Sinharaja Forest Reserve</collectingRegion>
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,
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<geoCoordinate id="FA551F0F57185906BEB7CFFDEDD2A339" direction="north" orientation="latitude" precision="0" value="6.411617">N 06° 24.697'</geoCoordinate>
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-
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<geoCoordinate id="A92802AC6AA4FE6CF5E57311F5D1BA14" direction="north" orientation="latitude" precision="0" value="6.413717">N 06° 24.823'</geoCoordinate>
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;
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<geoCoordinate id="4F4430E0C697AB449C9507D367D9B75A" direction="east" orientation="longitude" precision="0" value="80.41872">E80° 25.123'</geoCoordinate>
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-
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<geoCoordinate id="BA23E0929C6C4D653EA0347B3D152662" direction="east" orientation="longitude" precision="0" value="80.41652">E 80° 24.991'</geoCoordinate>
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,
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<elevation id="D977146C4A4F5B6C78E699D4694051A9">
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432m-
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<quantity id="32CA55139CDADC4F1F3117AA0BCAB2AD" metricMagnitude="2" metricUnit="m" metricValue="5.71" unit="m" value="571.0">571m</quantity>
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,
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</elevation>
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<date id="3A6DCEBEB4ECC25849AE66C215684D27" value="2006-03-25" valueMax="2006-04-07" valueMin="2006-03-25">
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<collectingDate id="0180CD5CFFD0D657CC490C2FB75A8992" value="2006-03-25" valueMax="2006-04-07" valueMin="2006-03-25">25 MAR-07APR 2006</collectingDate>
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</date>
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,
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<collectorName id="88537A7922A8B181BBB0F3F67B442C42">
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<taxonomicNameLabel id="837DC82086DD8A4FB64D0FB2A8904BF7" pageId="1" pageNumber="287" rank="subSpecies">N.R.</taxonomicNameLabel>
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Gunawardene
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</collectorName>
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,
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<collectingMethod id="F4A321C3F859FE51390A8AAA76A1B381">CTFS Plot Winkler sacks 15-18</collectingMethod>
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.
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<collectionCode id="F3768B05E51925BBAE2520D73D3AF21B" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZ</collectionCode>
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<typeStatus id="5080F709FCFFE85FD8086F40447E3AE1">holotype</typeStatus>
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<specimenCode id="EFB7912EE01C5EFAD2A1F5BC84F27A95">number 35624</specimenCode>
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(
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<specimenCode id="D3E0D911D1CAC89749F86EF642BC4E8C">CASENT0106177</specimenCode>
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,
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<specimenCode id="EF0EC26539B14CF14298329E5DC7BB05">ANTC4038</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="80A9E3E1DF1D6A2150BA302815D46F57" pageId="1" pageNumber="287" type="mainText">
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<typeStatus id="9C5996F1A0A72DC7D4D10F21F3FC238D">Holotype</typeStatus>
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deposited in the Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA. The specimen and collection labels have been imaged and placed on AntWiki (http://www.antwiki.org).
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</paragraph>
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</subSubSection>
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<subSubSection id="7869B1D9C1A5FE7882AC25DABF554479" lastPageId="3" lastPageNumber="289" pageId="1" pageNumber="287" type="description">
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<paragraph id="6C21C44FF62CE229F3D7E975AC0D641D" pageId="1" pageNumber="287" type="mainText">Worker measurements. HW 0.52, HL 0.72, EL 0.05, SL 0.54, PrW 0.47, ML 0.99, PL 0.35, PPL 0.22, PW</paragraph>
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<paragraph id="673F61F7F0ED5FB474D74C7D26695900" pageId="1" pageNumber="287" type="mainText">0.29, PPW 0.28, GL 0.76, TL 3.04, CI 72, SI 104.</paragraph>
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<paragraph id="9F945AB4C15E7564FA7DEED1043A0644" lastPageId="2" lastPageNumber="288" pageId="1" pageNumber="287" type="mainText">
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Description of worker. General appearance as in Fig. 1. Head in full face view longer than wide, sides and posterior margins rounded, sides slightly tapering anteriorly. Mandibles triangular, masticatory margin edentate except for two blunt apical teeth, subapical tooth close to and slightly smaller than apical tooth. When mandibles closed, gap at mastigatory margin v-shaped. Palp formula 2, 2. Clypeal anterior margin protrudes above mandibles and is thickened, bearing several straight, stiff hairs, clypeus posteriorly narrowly inserted between frontal lobes. Clypeus lacking carinae, bearing a few irregular shaped foveolae. Frontal lobes present and short but prominent, rounded and obscuring antennal sockets. Antennae 11-segmented with weakly defined three-segmented club, apical segment being largest and entire club equal in length to rest of funiculus, excluding pedicel. Scape shallowly curved at base, relatively short, not reaching posterior margin of head. Eyes small and irregular, composed of 4 or 5 poorly defined ommatidia and situated slightly anterior to midlength of head. Mesosoma without promesonotal or metanotal suture, forming a slightly convex outline in side view. In dorsal view mesosoma is widest in promesonotal area, decreasing in width posteriorly. Anteroventral corner of pronotum rounded. Propodeum armed with two very small, triangular denticles. Propodeal lobes broad and rounded. Petiole without clearly differentiated peduncle, in side view node somewhat thickened and rounded, anterior face less rounded than posterior face, in dorsal view longer than wide. Anteroventral lobe or projection present. Postpetiole wider than long; in dorsal view anterior margin concave and posterior margin convex. First gastral tergite covered with microreticulum fading well before half of its length, remainder of gaster smooth. Middle and hind legs without tibial spurs. All surfaces of the head, mesosoma, petiole and postpetiole completely covered with numerous large round foveolae almost touching
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<pageBreakToken id="B903911F8FE04CAA54C328C1BDAF4D92" pageId="2" pageNumber="288" start="start">each</pageBreakToken>
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other. Space between foveolae smooth and shiny. Long, stiff, erect, white hairs emerging from the center of each foveola. Abundant erect hairs present on body appendages, including antennal scape as well as funiculus; hairs present on all leg segments and on both dorsal and ventral surfaces of mandibles; those on ventral surface not differing visibly in structure from rest of pilosity. Body color reddish, appendages a lighter reddish color.
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</paragraph>
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<caption id="C4A38B7F7B44F88CC8A563EB389ED1C1" pageId="2" pageNumber="288">
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<paragraph id="0FD7B0B36868FDA4977824386952A002" pageId="2" pageNumber="288" type="caption">
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FIGURES 1-3.
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<taxonomicName id="B7F85194C1BDB73E9D50FCF6A727D698" genus="Tyrannomyrmex" pageId="2" pageNumber="288" rank="species" species="legatus">Tyrannomyrmex legatus</taxonomicName>
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,
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<typeStatus id="0229E029756AE46BEBB10BEE52720DAA">holotype</typeStatus>
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worker; 1-lateral view; 2-dorsal view; 3-frontal view.
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</paragraph>
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</caption>
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<paragraph id="4C29A114A2ACB3F54E361B4F242C3600" pageId="3" pageNumber="289" type="mainText">
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<pageBreakToken id="AF53DB3FD3C3DD64F296D911BC37C892" pageId="3" pageNumber="289" start="start">Gyne</pageBreakToken>
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and male: Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="F2A95047DD562BCCBAABC77614ED8CFD" pageId="3" pageNumber="289" type="etymology">
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<paragraph id="3EC5DCE7D897C3FC87D6FC78A558A6AC" pageId="3" pageNumber="289" type="mainText">
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Etymology. "
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<taxonomicName id="3E085026A13349AC7CCE786AFB81DCBF" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">legatus</taxonomicName>
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" refers to a military commander appointed by the Roman Senate.
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</paragraph>
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</subSubSection>
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<subSubSection id="4E6D61DF909FA1830473E560F9296E4E" pageId="3" pageNumber="289" type="diagnosis">
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<paragraph id="30886B7F57D5DC06450CC2F9FA9E96C8" pageId="3" pageNumber="289" type="mainText">
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Diagnosis.
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<taxonomicName id="7866E0990DF4B8325362CC236EC6BA6C" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">Tyrannomyrmex legatus</taxonomicName>
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is most easily distinguished from
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<taxonomicName id="FCDA12CE8B7730A2B24440DDF83DE70A" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="rex">T. rex</taxonomicName>
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and
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<taxonomicName id="6F14C62A651F348C7A7ED4AB9FEFB491" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="dux">T. dux</taxonomicName>
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by differences in pilosity, sculpture and the shape of the petiole and postpetiole.
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<taxonomicName id="4C1BF9F169244EF1E6000B07603A087C" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="rex">T. rex</taxonomicName>
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is almost lacking pilosity on the mesosomal dorsum, while the whole dorsal surface is covered with long erect hairs in
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<taxonomicName id="50FFD2307E7332CBFE168E60D127F642" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">T. legatus</taxonomicName>
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and
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<taxonomicName id="6E4FBC8366F3DD8EA890D2DDAD7EFDF1" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="dux">T. dux</taxonomicName>
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. The foveolation is weaker in
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<taxonomicName id="B3258F1ECFA3942A7CFE37EEFE3371D6" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="rex">T. rex</taxonomicName>
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, especially on the mesosoma where the foveae on the mesosoma are small with most interspaces equal or wider than their diameter.
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<taxonomicName id="60E2BB19EFA2638CB320E1944D48CC1F" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">Tyrannomyrmex legatus</taxonomicName>
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can be most easily separated from
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<taxonomicName id="9881EB01371D0DAEBB6CC2798104FD8D" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="dux">T. dux</taxonomicName>
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by the shape of the petiole, which is much more robust in the former. In lateral view, the peduncle of the petiole is not clearly differentiated, with an abrupt anterior slope of the node. There is also a conspicuous antereoventral projection of the petiole in
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<taxonomicName id="529BA6AECF1A8C1E52C28FC24097FE1F" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">T. legatus</taxonomicName>
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which is absent in
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<taxonomicName id="9308D557EB2F9115223D97AF031B28C8" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="dux">T. dux</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="4F58ADD21EE0354806892330ED1CB93A" pageId="3" pageNumber="289" type="distribution">
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<paragraph id="0A9E058916045C563359CC473B0C262B" pageId="3" pageNumber="289" type="mainText">Distribution and Habits. This single specimen was collected in lowland dipterocarp forest (undisturbed, unlogged), near a stream at the bottom of a slope in the drier period of the year.</paragraph>
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</subSubSection>
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<subSubSection id="A43ECFFCE7965BB4949DA897CBCEA104" lastPageId="4" lastPageNumber="290" pageId="3" pageNumber="289" type="discussion">
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<paragraph id="2DF3159163EF925C5DBAB78D29935CF3" pageId="3" pageNumber="289" type="mainText">Discussion</paragraph>
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<paragraph id="7F5C4A3C89B8121ABF523735FDC72762" pageId="3" pageNumber="289" type="mainText">
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The small eyes, edentate mandibles, and close similarity among the workers of all three
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<taxonomicName id="634596FE3EFECC0A6FCD42FEE0369411" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="genus">Tyrannomyrmex</taxonomicName>
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species strongly suggest that they may also be similar ecologically, and that they are probably subterranean and predaceous. While the three known worker specimens have been taken in leaf litter samples, the rarity of collections suggests that
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<taxonomicName id="DC02CFF9BFCABF19EA65C21F14B8CD68" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="genus">Tyrannomyrmex</taxonomicName>
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species may both nest and forage in the deeper soil horizons, and that foragers may only occasionally enter the leaf litter layers closer to the surface.
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</paragraph>
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<paragraph id="0F017F78EE7570A915D8B1EE24AB3351" pageId="3" pageNumber="289" type="mainText">
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So far, all species of
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<taxonomicName id="78EA6B910B60E2DF313F0DD8CA7FF44A" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="genus">Tyrannomyrmex</taxonomicName>
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occur in tropical Old World forests. The wide distribution range from India and Sri Lanka in the west to peninsular Malaysia and perhaps the Philippine archipelago in the east suggests that more species may be discovered as we improve our ability to sample deeper soil microhabitats. New probe techniques for sampling below the leaf litter layer hold promise for discovering more of this subterranean ant community (Ryder-Wilkie et al. 2007).
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</paragraph>
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<paragraph id="B4107F83B463F0E66CCB53FC02796053" pageId="3" pageNumber="289" type="mainText">
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Based upon the additional analysis of
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<taxonomicName id="F1071099FE5DFA9ECF7AFB7824418A55" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">T. legatus</taxonomicName>
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, the following morphological characters for the genus are summarized below:
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</paragraph>
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<paragraph id="35B35CDAF28BF3461A667205FE73B375" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="80BEBA40EEAD48F84D10DBF7C6172845" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Mandibles with two teeth in the masticatory border, apical and smaller subapical. No teeth on the basal margin of the mandible.
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</paragraph>
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<paragraph id="DA43660CF4F939C5275F5E2C61B8FFA4" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="0BE50C8243CD01C271D9F81EDF77E79D" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Inner ventral margin of masticatory border of mandibles with setae. Setae can be normal or modified.
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</paragraph>
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<paragraph id="75CAF4722C6E51005A5F10A3D218B3B7" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="C522C0919AB7A69A2BAA17B3D51FA90A" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Clypeus devoid of carinae. Foveolae may be present.
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</paragraph>
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<paragraph id="50C196CAB69DAF70604C448F71F27F21" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="0311BE6F107332B67BE8784AF093B4FE" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Palpal formula 2, 2.
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</paragraph>
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<paragraph id="801558B5A7E75BCECA49EA63EE15BE3C" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="D792B3D20CB5EE9056A8206C79C3C671" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Compound eyes small, reduced to a few ommatidia.
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</paragraph>
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<paragraph id="8771281C7F9B2B92DA86108B029FDF43" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="1BC92A74501556490527DEA0D0D26424" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Antennae 11-segmented with an ill-defined 3-segmented club.
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</paragraph>
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<paragraph id="5EC899756FBFEA9E58CAA38C3FEA30EE" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="C54D061A39AAA01451E264BDC7C45670" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Frontal carinae and antennal scrobes absent.
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</paragraph>
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<paragraph id="34278BEE6155FD09E7C9A6801873CBAB" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="156DE36EE504EB9C2F54187FFD09628C" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Mesosoma without promesonotal suture.
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</paragraph>
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<paragraph id="82C86E80DE2D4C0F9438C4F580CB8686" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="73A5108F018C6377CA5B725E37F4B467" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
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Propodeal lobes large and round.
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</paragraph>
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<paragraph id="587026DB49358F2596AA5F31AA8E28AA" pageId="3" pageNumber="289" type="mainText">
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<normalizedToken id="04301872844FD99C393903AB080E9FF6" originalValue="•" pageId="3" pageNumber="289">*</normalizedToken>
|
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Sting large and robust.
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</paragraph>
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<paragraph id="E202DB440E4D4235812C71E1A354488D" pageId="3" pageNumber="289" type="mainText">
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<taxonomicName id="10597E0BEFC588847246E79757A90D0C" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="legatus">Tyrannomyrmex legatus</taxonomicName>
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has its sting extruded and it is comparable in length with
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|
<taxonomicName id="6700A4F46052EF56AD72DFF72B3FA709" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="dux">T. dux</taxonomicName>
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|
. It is possible that a fully extruded sting could appear to be longer as in
|
|
<taxonomicName id="0C3CCE617518998A610CB4B811921A96" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="species" species="rex">T. rex</taxonomicName>
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. The palp formula is 2,2 and each segment is rather short. The setae on the ventral margin of the mandibles are normal. There is no median seta on the anterior margin of the clypeus, but rather a series of evenly spaced setae along the entire margin.
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</paragraph>
|
|
<paragraph id="986A204DB2A3D9F251A9FA20E0B289BF" pageId="3" pageNumber="289" type="mainText">
|
|
<normalizedToken id="4BC219FC9520A7D238457CF42854A311" originalValue="Fernández" pageId="3" pageNumber="289">Fernandez</normalizedToken>
|
|
(2003) provisionally concluded that
|
|
<taxonomicName id="344A5A71F8133DBF7EEE4D20C2DE9729" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="genus">Tyrannomyrmex</taxonomicName>
|
|
is a distinct and isolated genus within the subfamily
|
|
<taxonomicName id="E33E71B0B227C6E0CDFA6220DDEEA8FD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2258" lsidName-HNS="Myrmicinae Lepeletier" pageId="3" pageNumber="289" rank="subFamily" subFamily="Myrmicinae">Myrmicinae</taxonomicName>
|
|
with possible affinities to either the Adelomyrmex-genus group or the tribe
|
|
<taxonomicName id="BEDC5A91A5E4760FB80078B0CD686E4D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2312" lsidName-HNS="Solenopsidini Forel" pageId="3" pageNumber="289" rank="tribe" tribe="Solenopsidini">Solenopsidini</taxonomicName>
|
|
. In a comprehensive treatment of the
|
|
<taxonomicName id="A30F8F868DA36E9785F355E6CD3420C7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:152" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Formicidae Latreille" order="Hymenoptera" pageId="3" pageNumber="289" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
|
, Bolton (2003) lists in Appendix 2, a set of character states for all ant genera. Antennomere count, presence and
|
|
<typeStatus id="2C25A7FA3F5F827FD98ECBD9772A5DEA">type</typeStatus>
|
|
of antennal club, palp formula, total dental count and spur formula are scored for each genus. Using this set of characters, a matrix was constructed which forms the foundation of a LUCID Key to the Extant Ant Genera of the World at http://gap.entclub.org/index.html.
|
|
<taxonomicName id="CF6AAC93846E104EE64975DA29CFD940" genus="Tyrannomyrmex" pageId="3" pageNumber="289" rank="genus">Tyrannomyrmex</taxonomicName>
|
|
and the genus
|
|
<taxonomicName id="EFAED6E5D3862C4D1C8896D1D0695183" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="289" phylum="Arthropoda" rank="genus">Monomorium</taxonomicName>
|
|
uniquely share the same character states in this key.
|
|
</paragraph>
|
|
<paragraph id="EFDEA89E7423DBD42FBFF1DCCFF3CD16" pageId="4" pageNumber="290" type="mainText">
|
|
<pageBreakToken id="AE9D47D703D16046F9BA53F097D7CA6B" pageId="4" pageNumber="290" start="start">Bolton</pageBreakToken>
|
|
(2003) constructs a diagnosis for the tribe
|
|
<taxonomicName id="A342FBF0419E9DBF223C8F2A2FE6079B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2312" lsidName-HNS="Solenopsidini Forel" pageId="4" pageNumber="290" rank="tribe" tribe="Solenopsidini">Solenopsidini</taxonomicName>
|
|
and lists a set of characters that encompass
|
|
<taxonomicName id="974AB2C6BE27D65E1266A75FD688B59B" genus="Tyrannomyrmex" pageId="4" pageNumber="290" rank="genus">Tyrannomyrmex</taxonomicName>
|
|
along with 17 other genera including
|
|
<taxonomicName id="A7295ACE77A577D73ECE05001439C5F1" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="290" phylum="Arthropoda" rank="genus">Monomorium</taxonomicName>
|
|
. Specifically all members have mandibles subtriangular to triangular and short; total dental count 2-6; anterior clypeal margin with or without an isolated stout median seta; clypeus bicarinate and usually distinct; frontal lobes small and narrow, with straight to convex outer margins; antennal scrobes and frontal carinae usually absent; dorsal mesosoma without promesonotal suture; metatibial spur simple to absent; propodeal lobes broad and rounded; tergite of abdominal segment IV broadly overlapping sternite on ventral surface of gaster; head and dorsal mesosoma usually smooth or weakly sculptured; antenna with 7-12 segments, with a 2, 3, or 4-segmented club.
|
|
<taxonomicName id="CA530F8AF60A792387A5E2452A6BF81C" genus="Tyrannomyrmex" pageId="4" pageNumber="290" rank="genus">Tyrannomyrmex</taxonomicName>
|
|
falls well within this schema including its strong surface sculpture which is shared with a number of Australian
|
|
<taxonomicName id="D814ADCD6D1ABC83D5A263B19521D046" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="290" phylum="Arthropoda" rank="genus">Monomorium</taxonomicName>
|
|
species, some species of the
|
|
<taxonomicName id="97A6D031577E79180219A6FB2EC8968F" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="290" phylum="Arthropoda" rank="species" species="scabriceps">Monomorium scabriceps</taxonomicName>
|
|
group, some
|
|
<taxonomicName id="845593925AF9AEC5FCCE5221C4E373CA" class="Insecta" family="Formicidae" genus="Oxyepoecus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="290" phylum="Arthropoda" rank="genus">Oxyepoecus</taxonomicName>
|
|
,
|
|
<taxonomicName id="042AF1FD4A900AAFFB12E9E8468D7DEF" class="Insecta" family="Formicidae" genus="Mayriella" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="290" phylum="Arthropoda" rank="genus">Mayriella</taxonomicName>
|
|
and some major workers of
|
|
<taxonomicName id="070DE2DF7E25BEB63250641CBC37D910" class="Insecta" family="Formicidae" genus="Oligomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="290" phylum="Arthropoda" rank="genus">Oligomyrmex</taxonomicName>
|
|
(Bolton, 2003).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |