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<document id="841576CAF701B1B6111003C811D82C34" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFB8FF89FF3DA2700070FEA9" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus emusorum Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="176" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="174" updateTime="1698611248703" updateUser="plazi">
<mods:mods id="C05FAF947F9286AC6CA4D55946A2E844" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D1C4CE9754C6643D237FACB84871139B">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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<mods:namePart id="3E20D67171ACE3D325C3BC5226897923">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="B1F4ABE65B0D6481F0DDDFA33E019A28">text</mods:typeOfResource>
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<mods:title id="B7966340BD8661E35F1FB77BE0873680">Zootaxa</mods:title>
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<mods:part id="4B9CC284A6EAC9B41A9EDBA245D82FC0">
<mods:date id="7783E043FD67E36EC5CCB047C2DAE0B1">2015</mods:date>
<mods:detail id="B2A10870B665091723865E287E309657" type="volume">
<mods:number id="1F2CE6F1206735593E74E7D1087F2E0E">4020</mods:number>
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<mods:classification id="55E9FB4B14D0D196D7E7304067A7DAA2">journal article</mods:classification>
<mods:identifier id="623E27F99ED7336859560729FC736BE0" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
<mods:identifier id="66704795D25D28DB5AF3A1D835BD7547" type="GBIF-Dataset">82b2f539-5d25-42d6-95c5-e8b16a353e12</mods:identifier>
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<mods:identifier id="63F5016145AEC739B4AE0832E351A925" type="Zenodo-Dep">237970</mods:identifier>
<mods:identifier id="D9AA9E6A65883A8E5A999EF2AAC7EC03" type="ZooBank">BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE</mods:identifier>
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<treatment id="172D87FDFFB8FF89FF3DA2700070FEA9" ID-DOI="http://doi.org/10.5281/zenodo.6121805" ID-GBIF-Taxon="119638521" ID-Zenodo-Dep="6121805" LSID="urn:lsid:plazi:treatment:172D87FDFFB8FF89FF3DA2700070FEA9" httpUri="http://treatment.plazi.org/id/172D87FDFFB8FF89FF3DA2700070FEA9" lastPageId="7" lastPageNumber="176" pageId="5" pageNumber="174">
<subSubSection id="D79E6560FFB8FF8BFF3DA270054EFA7A" pageId="5" pageNumber="174" type="nomenclature">
<paragraph id="9F3B36EBFFB8FF8BFF3DA270061AFA59" blockId="5.[151,579,1402,1462]" box="[151,579,1402,1429]" pageId="5" pageNumber="174">
<heading id="C4738187FFB8FF8BFF3DA270061AFA59" bold="true" box="[151,579,1402,1429]" fontSize="11" level="1" pageId="5" pageNumber="174" reason="1">
<emphasis id="ADF0EAF9FFB8FF8BFF3DA270061AFA59" bold="true" box="[151,579,1402,1429]" pageId="5" pageNumber="174">
<taxonomicName id="58844D68FFB8FF8BFF3DA2700587FA59" ID-CoL="9HXT9" authority="Bickel" authorityName="Bickel" box="[151,478,1402,1429]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="174" phylum="Arthropoda" rank="species" species="emusorum" status="sp. nov.">
<emphasis id="ADF0EAF9FFB8FF8BFF3DA27005D0FA58" bold="true" box="[151,393,1402,1428]" italics="true" pageId="5" pageNumber="174">Systenus emusorum</emphasis>
Bickel
</taxonomicName>
<taxonomicNameLabel id="B6C35782FFB8FF8BFE4FA271061AFA59" box="[485,579,1403,1429]" pageId="5" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9F3B36EBFFB8FF8BFF3DA297054EFA7A" blockId="5.[151,579,1402,1462]" box="[151,279,1437,1462]" pageId="5" pageNumber="174">
(
<figureCitation id="07BF2A6EFFB8FF8BFF35A29704BFFA7A" box="[159,230,1437,1462]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="5" pageNumber="174">Figs 2</figureCitation>
a, b)
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFB8FF8BFF3DA2EA0636F9D0" pageId="5" pageNumber="174" type="materials_examined">
<paragraph id="9F3B36EBFFB8FF8BFF3DA2EA0636F9D0" blockId="5.[151,1437,1504,2036]" pageId="5" pageNumber="174">
<emphasis id="ADF0EAF9FFB8FF8BFF3DA2EA0566FA35" bold="true" box="[151,319,1504,1529]" pageId="5" pageNumber="174">
<typeStatus id="403F8849FFB8FF8BFF3DA2EA048BFA35" box="[151,210,1504,1529]" pageId="5" pageNumber="174">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="403F8849FFB8FF8BFEE4A2EA05B5FA35" box="[334,492,1504,1529]" pageId="5" pageNumber="174" type="holotype">
<emphasis id="ADF0EAF9FFB8FF8BFEE4A2EA05B5FA35" bold="true" box="[334,492,1504,1529]" pageId="5" pageNumber="174">HOLOTYPE</emphasis>
</typeStatus>
♂,
<collectingCountry id="E793767BFFB8FF8BFDB0A2EA069AFA34" box="[538,707,1504,1528]" name="Costa Rica" pageId="5" pageNumber="174">COSTA RICA</collectingCountry>
:
<emphasis id="ADF0EAF9FFB8FF8BFD65A2EA0702FA35" bold="true" box="[719,859,1504,1529]" pageId="5" pageNumber="174">Guanacaste</emphasis>
:
<quantity id="587C9B0EFFB8FF8BFCC1A2EA07FFFA35" box="[875,934,1504,1529]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="174" unit="km" value="3.0">3 km</quantity>
SE Rio Naranjo,
<date id="EB3A102BFFB8FF8BFBD8A2EA0157FA34" box="[1138,1294,1504,1528]" pageId="5" pageNumber="174" value="1992-03-02" valueMax="1992-03-09" valueMin="1992-03-02">2—9.III.1992</date>
, F.D. Parker (
<collectionCode id="F995AE2EFFB8FF8BFF34A10E04AAF9D0" LSID="urn:lsid:biocol.org:col:34802" box="[158,243,1540,1564]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="5" pageNumber="174">LACM</collectionCode>
, ex.
<collectionCode id="F995AE2EFFB8FF8BFE8DA10E0520F9D0" box="[295,377,1540,1564]" pageId="5" pageNumber="174">EMUS</collectionCode>
;
<collectionCode id="F995AE2EFFB8FF8BFE2EA10E058CF9D0" LSID="urn:lsid:biocol.org:col:34802" box="[388,469,1540,1564]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="5" pageNumber="174">LACM</collectionCode>
Ent 329998).
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFB8FF88FF6DA12205EAFE8C" lastPageId="6" lastPageNumber="175" pageId="5" pageNumber="174" type="description">
<paragraph id="9F3B36EBFFB8FF8BFF6DA1220716F9A9" blockId="5.[151,1437,1504,2036]" pageId="5" pageNumber="174">
<emphasis id="ADF0EAF9FFB8FF8BFF6DA1220508F98D" bold="true" box="[199,337,1576,1601]" pageId="5" pageNumber="174">Description</emphasis>
.
<emphasis id="ADF0EAF9FFB8FF8BFEF5A12205C5F98D" bold="true" box="[351,412,1576,1601]" pageId="5" pageNumber="174">Male</emphasis>
: body length:
<quantity id="587C9B0EFFB8FF8BFD97A12206CCF98C" box="[573,661,1576,1600]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="174" unit="mm" value="2.3">2.3 mm</quantity>
; wing 2.4 x
<quantity id="587C9B0EFFB8FF8BFCB7A122072AF98C" box="[797,883,1576,1600]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="174" unit="mm" value="0.7">0.7 mm</quantity>
(
<figureCitation id="07BF2A6EFFB8FF8BFC2BA122079CF98C" box="[897,965,1576,1601]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="5" pageNumber="174">Fig. 2</figureCitation>
a); similar to
<taxonomicName id="58844D68FFB8FF8BFBF2A12300BBF98D" box="[1112,1250,1576,1601]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="174" phylum="Arthropoda" rank="species" species="eboritibia">
<emphasis id="ADF0EAF9FFB8FF8BFBF2A12300BBF98D" box="[1112,1250,1576,1601]" italics="true" pageId="5" pageNumber="174">S. eboritibia</emphasis>
</taxonomicName>
except as noted:
<emphasis id="ADF0EAF9FFB8FF8BFF3DA146048DF9A9" box="[151,212,1612,1637]" italics="true" pageId="5" pageNumber="174">Head</emphasis>
: antenna missing on unique specimen, but scape yellow.
</paragraph>
<paragraph id="9F3B36EBFFB8FF88FF6DA17A05EAFE8C" blockId="5.[151,1437,1504,2036]" lastBlockId="6.[151,1437,151,320]" lastPageId="6" lastPageNumber="175" pageId="5" pageNumber="174">
<emphasis id="ADF0EAF9FFB8FF8BFF6DA17A0541F945" box="[199,280,1648,1673]" italics="true" pageId="5" pageNumber="174">Thorax</emphasis>
: metepimeron distinctly yellow.
<emphasis id="ADF0EAF9FFB8FF8BFD27A17B069BF944" box="[653,706,1649,1672]" italics="true" pageId="5" pageNumber="174">Legs</emphasis>
: all coxae and remainder of legs yellow, except distal quarter of FIII and TIII dark brown, CI and CII with white anterior setae and shorter vestiture, CIII with strong while lateral seta at ½; major leg setae black; I: 2.9; 2.8; 1.2/ 0.7/ 0.4/ 0.2/ 0.3; leg I bare of major setation; TI with whitish vestiture on anterior face (MSSC); II: 3.7; 3.9; 2.0/ 1.2/ distal tarsomeres missing; TII with strong ad seta and weaker pd seta at ¼, pd seta only and also ventral seta at 2/3, and with apical ring of ad, pd, av and ventral seta; III: 3.8; 4.3; tarsomeres missing; TIII with 3 black dorsal setae from ¼ to 3/4, and with apical ring of dorsal, ad and av seta.
<emphasis id="ADF0EAF9FFB8FF8BFF7AA0430551F8AC" box="[208,264,1865,1888]" italics="true" pageId="5" pageNumber="174">Wing</emphasis>
: veins M and R4+5 subparallel (not bowed) beyond dm-cu crossvein; CuAx ratio: 0.8; lower calypter pale yellow with yellowish marginal setae; halter yellow.
<emphasis id="ADF0EAF9FFB8FF8BFD49A0640714F84B" box="[739,845,1902,1927]" italics="true" pageId="5" pageNumber="174">Abdomen</emphasis>
: tergite 1 yellowish; tergite 2 mostly yellow but with some metallic green iridescence; tergite 3 metallic bronze-green dorsally and yellowish lateral; tergites 4–7 mostly metallic green-bronze, shining with little pruinosity; posterior margin of tergite 1 with posterior row of long black setae, tergites otherwise covered with short black vestiture; segment 7 forming relatively shorter peduncle, with sternite 7 glabrous and tergite 7 covered with black vestiture; sternite 8 with white hairs; hypopygium (
<figureCitation id="07BF2A6EFFBBFF88FAEAA79D01DCFF7C" box="[1344,1413,151,176]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="6" pageNumber="175">Fig. 2</figureCitation>
b) dark brown with yellow surstylus and cercus; epandrium projected ventrally and apically downturned; surstylus basally squarish with apical digitiform projection on dorsal margin, and with curved digitiform projection arising dorsobasally; cercus elongate, thick basally, then narrowed and clavate distally, and covered with short yellow setae.
<emphasis id="ADF0EAF9FFBBFF88FF77A62D056DFE8C" bold="true" box="[221,308,295,320]" pageId="6" pageNumber="175">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<caption id="CBFB6663FFBBFF88FF3DA0BF000EF826" httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="6" pageNumber="175" targetBox="[167,1436,373,1951]" targetPageId="6">
<paragraph id="9F3B36EBFFBBFF88FF3DA0BF000EF826" blockId="6.[151,1436,1973,2026]" pageId="6" pageNumber="175">
<emphasis id="ADF0EAF9FFBBFF88FF3DA0BF0555F807" bold="true" box="[151,268,1973,1995]" pageId="6" pageNumber="175">FIGURE 2</emphasis>
.
<taxonomicName id="58844D68FFBBFF88FEB3A0BC0586F807" box="[281,479,1974,1995]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="emusorum" status="sp. nov.">
<emphasis id="ADF0EAF9FFBBFF88FEB3A0BC0586F807" box="[281,479,1974,1995]" italics="true" pageId="6" pageNumber="175">Systenus emusorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6C35782FFBBFF88FE4FA0BF066FF807" box="[485,566,1973,1995]" pageId="6" pageNumber="175" rank="species">
<emphasis id="ADF0EAF9FFBBFF88FE4FA0BF066FF807" bold="true" box="[485,566,1973,1995]" pageId="6" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype):
<emphasis id="ADF0EAF9FFBBFF88FD19A0BF0699F807" bold="true" box="[691,704,1973,1995]" pageId="6" pageNumber="175">a</emphasis>
. male habitus, left lateral;
<emphasis id="ADF0EAF9FFBBFF88FC67A0BF0782F807" bold="true" box="[973,987,1973,1995]" pageId="6" pageNumber="175">b</emphasis>
. hypopygium, left lateral.
<taxonomicName id="58844D68FFBBFF88FB42A0BC012EF807" box="[1256,1399,1974,1995]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="naranjensis" status="sp. nov.">
<emphasis id="ADF0EAF9FFBBFF88FB42A0BC012EF807" box="[1256,1399,1974,1995]" italics="true" pageId="6" pageNumber="175">S. naranjensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6C35782FFBBFF88FAD7A0BF049AF826" pageId="6" pageNumber="175" rank="species">
<emphasis id="ADF0EAF9FFBBFF88FAD7A0BF049AF826" bold="true" pageId="6" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype):
<emphasis id="ADF0EAF9FFBBFF88FEEAA0DE0512F826" bold="true" box="[320,331,2004,2026]" pageId="6" pageNumber="175">c</emphasis>
. male habitus, left lateral;
<emphasis id="ADF0EAF9FFBBFF88FDF0A0DE0631F826" bold="true" box="[602,616,2004,2026]" pageId="6" pageNumber="175">d</emphasis>
. hypopygium, left lateral;
<emphasis id="ADF0EAF9FFBBFF88FCDCA0DE07D8F826" bold="true" box="[886,897,2004,2026]" pageId="6" pageNumber="175">e</emphasis>
. male head, anterior.
</paragraph>
</caption>
<subSubSection id="D79E6560FFBAFF89FF6DA79D000DFED1" pageId="7" pageNumber="176" type="discussion">
<paragraph id="9F3B36EBFFBAFF89FF6DA79D000DFED1" blockId="7.[151,1437,151,357]" pageId="7" pageNumber="176">
<emphasis id="ADF0EAF9FFBAFF89FF6DA79D056DFF7C" bold="true" box="[199,308,151,176]" pageId="7" pageNumber="176">Remarks</emphasis>
.
<taxonomicName id="58844D68FFBAFF89FEEEA792067BFF7C" box="[324,546,152,176]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="176" phylum="Arthropoda" rank="species" species="emusorum">
<emphasis id="ADF0EAF9FFBAFF89FEEEA792067BFF7C" box="[324,546,152,176]" italics="true" pageId="7" pageNumber="176">Systenus emusorum</emphasis>
</taxonomicName>
is known from a single male taken at the Rio Naranjo site,
<collectingCountry id="E793767BFFBAFF89FB7CA792010BFF7C" box="[1238,1362,151,176]" name="Costa Rica" pageId="7" pageNumber="176">Costa Rica</collectingCountry>
in the month of March. The antennae are missing on the single known specimen. However, diagnostic characters include all coxae and legs mostly yellow except the distal quarter of both femur III and tibia III which are dark brown, and the metepimeron is yellow. Tibia I has ivory-colored vestiture on the anterior surface.
</paragraph>
</subSubSection>
<subSubSection id="D79E6560FFBAFF89FF6DA62D0070FEA9" pageId="7" pageNumber="176" type="etymology">
<paragraph id="9F3B36EBFFBAFF89FF6DA62D0070FEA9" blockId="7.[151,1437,151,357]" pageId="7" pageNumber="176">
<emphasis id="ADF0EAF9FFBAFF89FF6DA62D051FFE8C" bold="true" box="[199,326,295,320]" pageId="7" pageNumber="176">Etymology</emphasis>
. This species is named in honor of the staff at the Entomological Museum, Utah State University (acronym EMUS) for their work in building such a fine entomological collection.
</paragraph>
</subSubSection>
</treatment>
</document>
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