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<document id="262E034C798C29BF7A12B4E6FAC0B4CD" ID-DOI="10.11646/zootaxa.3852.3.1" ID-GBIF-Dataset="957e5d0f-c452-4d45-85cf-24f138ab2359" ID-ISSN="1175-5326" ID-Zenodo-Dep="229938" ID-ZooBank="13B15A57-576A-4617-9D8A-7F27D970315A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460883855845" checkinUser="plazi" docAuthor="Couri, Márcia &amp; Pont, Adrian C." docDate="2014" docId="03FF87844601FFDBFF5EFD376AD2B42B" docLanguage="en" docName="zt03852p320.pdf" docOrigin="Zootaxa 3852 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenosia duomaculata Couri &amp; Pont, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="305" masterDocId="FFC6FFFC4600FFDFFFC9FFCB6E55B616" masterDocTitle="New species of Afrotropical Muscidae (Diptera: Muscoidea)" masterLastPageNumber="320" masterPageNumber="301" pageNumber="302" updateTime="1698358705595" updateUser="plazi">
<mods:mods id="E3482CF5CF7369F828052C6A2F0076E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5579A5DA9E003808A76ADBC487675C93">
<mods:title id="505B9456D6290E0C6E4DC691FF383058">New species of Afrotropical Muscidae (Diptera: Muscoidea)</mods:title>
</mods:titleInfo>
<mods:name id="0B10EC272208924733CF7A5A75FDAE28" type="personal">
<mods:role id="15068E249C0E6C767C1AC29AB37D1FE7">
<mods:roleTerm id="3BC900C675FDB025DD80EDB18E6EC9DD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="450442CCB47A7D4586CAAE4C6A3A2A94">Couri, Márcia</mods:namePart>
</mods:name>
<mods:name id="10EBF0B80EFE0C0364886B160E39514B" type="personal">
<mods:role id="46939A1D732C3718647A76C9FC181070">
<mods:roleTerm id="A0EF0FE4C1DDFE215F771AE6B6515BA5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5061A8FD1B6967BD236097AEDAD1B34F">Pont, Adrian C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="AA27F73477F5A16CF0053C2C915219BF">text</mods:typeOfResource>
<mods:relatedItem id="8C172A1757D3397C7B0581668FF7B05D" type="host">
<mods:titleInfo id="70666A1F0F3BD57D3818A3FBE424C579">
<mods:title id="2A73CA938FECD601344C8FD1C0B6CD9B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AACE0CCB9A7BEEAF4C151194CE0C7D88">
<mods:date id="F5FA53D6F058D7E4E54F2A27DB80FC3D">2014</mods:date>
<mods:detail id="AEA413A676D7E5639F83104C5D41EEAA" type="volume">
<mods:number id="313CFB3D3AC97BE19E10E51FD4B8F0EC">3852</mods:number>
</mods:detail>
<mods:detail id="44415589EB681562290B60331CE2EB3C" type="issue">
<mods:number id="CDA660E3B2A7C2AE62FCC05E974592F1">3</mods:number>
</mods:detail>
<mods:extent id="D313695F4826DE66BAC31E8A616F5230" unit="page">
<mods:start id="47F80667BB65F4709248D8250E26C33A">301</mods:start>
<mods:end id="D1B3B84AB8F0B6C64BD326EED8B0E66A">320</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B73C46B9FD9036DF911A2448EBED3C4F">journal article</mods:classification>
<mods:identifier id="A70FAD5841DA8145306B6652220FF785" type="DOI">10.11646/zootaxa.3852.3.1</mods:identifier>
<mods:identifier id="DB5C17E9278D27045D8A1D65953A8C23" type="GBIF-Dataset">957e5d0f-c452-4d45-85cf-24f138ab2359</mods:identifier>
<mods:identifier id="69A46BAC99A537087D0F18BB7465FCF4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="384D1FF99F10A7A1E2796160D32068C8" type="Zenodo-Dep">229938</mods:identifier>
<mods:identifier id="29D8B43AD76B36AD12332AB99641848D" type="ZooBank">13B15A57-576A-4617-9D8A-7F27D970315A</mods:identifier>
</mods:mods>
<treatment id="03FF87844601FFDBFF5EFD376AD2B42B" ID-DOI="http://doi.org/10.5281/zenodo.6137874" ID-GBIF-Taxon="119585850" ID-Zenodo-Dep="6137874" LSID="urn:lsid:plazi:treatment:03FF87844601FFDBFF5EFD376AD2B42B" httpUri="http://treatment.plazi.org/id/03FF87844601FFDBFF5EFD376AD2B42B" lastPageId="4" lastPageNumber="305" pageId="1" pageNumber="302">
<subSubSection id="C34C65194601FFDEFF5EFD376F4EB52E" pageId="1" pageNumber="302" type="nomenclature">
<paragraph id="8BE936924601FFDEFF5EFD376C55B503" blockId="1.[151,512,764,824]" box="[151,512,764,789]" pageId="1" pageNumber="302">
<heading id="D0A181FE4601FFDEFF5EFD376C55B503" bold="true" box="[151,512,764,789]" fontSize="10" level="2" pageId="1" pageNumber="302" reason="2">
<emphasis id="B922EA804601FFDEFF5EFD376C55B503" bold="true" box="[151,512,764,789]" pageId="1" pageNumber="302">
<taxonomicName id="4C564D114601FFDEFF5EFD376FCAB503" ID-CoL="9J2HP" box="[151,415,764,789]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="duomaculata" status="sp. nov.">
<emphasis id="B922EA804601FFDEFF5EFD376FCAB503" bold="true" box="[151,415,764,789]" italics="true" pageId="1" pageNumber="302">Coenosia duomaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4601FFDEFE6EFD376C55B503" box="[423,512,764,789]" pageId="1" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936924601FFDEFF5EFCD46F4EB52E" blockId="1.[151,512,764,824]" box="[151,283,799,824]" pageId="1" pageNumber="302">
<figureCitation id="136D2A174601FFDEFF5EFCD46E89B52E" box="[151,220,799,824]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,264,1956,1979]" captionTargetBox="[196,1429,467,1915]" captionTargetId="figure@2.[173,1458,445,1939]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1. Coenosia duomaculata sp. nov., notum, dorsal view; 2. Coenosia nigromaculata sp. nov., notum, dorsal view; 3. Helina harrisorum sp. nov., notum, dorsal view; 4. Helina ferfriniorum sp. nov., notum, dorsal view." httpUri="https://zenodo.org/record/229939/files/figure.png" pageId="1" pageNumber="302">Figs 1</figureCitation>
,
<figureCitation id="136D2A174601FFDEFF23FCD46F4DB52E" box="[234,280,799,824]" captionStart="FIGURES 5 – 12" captionStartId="3.[151,264,1482,1505]" captionTargetBox="[191,1414,197,1445]" captionTargetId="figure@3.[151,1436,175,1461]" captionTargetPageId="3" captionText="FIGURES 5 – 12. Coenosia duomaculata sp. nov.: 5. ovipositor, dorsal view; 6. ovipositor, ventral view and spermathecae; 7. Egg. Coenosia nigromaculata sp. nov.: 8. ovipositor, dorsal view; 9. ovipositor, ventral view and spermathecae. Coenosia fragilis sp. nov.: 10. sternite 5, dorsal view; 11. cercal plate and surstyli, dorsal view; 12. cercal plate, surstylus and aedeagus, lateral view." httpUri="https://zenodo.org/record/229940/files/figure.png" pageId="1" pageNumber="302">5–7</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65194601FFDEFF5EFCAC6D8EB342" pageId="1" pageNumber="302" type="materials_examined">
<paragraph id="8BE936924601FFDEFF5EFCAC6F92B5DE" blockId="1.[151,1436,871,2013]" pageId="1" pageNumber="302">
<emphasis id="B922EA804601FFDEFF5EFCAC6F12B596" bold="true" box="[151,327,871,896]" pageId="1" pageNumber="302">
<typeStatus id="54ED88304601FFDEFF5EFCAC6E87B596" box="[151,210,871,896]" pageId="1" pageNumber="302">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED88304601FFDEFE86FCAC6FE2B596" box="[335,439,871,896]" pageId="1" pageNumber="302" type="holotype">Holotype</typeStatus>
♀,
<collectingCountry id="F34176024601FFDEFE2CFCA36C1CB596" box="[485,585,872,896]" name="Kenya" pageId="1" pageNumber="302">KENYA</collectingCountry>
: Rift Valley Prov., Timboroa Forest (compt. 9), 00˚04.092′N, 35˚30.909′E,
<quantity id="4CAE9B774601FFDEFF5EFC476EA8B5B2" box="[151,253,908,933]" metricMagnitude="3" metricUnit="m" metricValue="2.628" pageId="1" pageNumber="302" unit="m" value="2628.0">2628 m</quantity>
, Malaise traps, indigenous Afromontane forest,
<date id="FFE810524601FFDEFCBBFC476A44B5B3" box="[882,1041,908,933]" pageId="1" pageNumber="302" value="2011-04-14" valueMax="2011-04-16" valueMin="2011-04-14">14–16.iv.2011</date>
(A.H. &amp; M.K. Kirk-Spriggs),
<collectionCode id="ED47AE574601FFDEFF5EFC7B6EBDB5DE" box="[151,232,944,968]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)32170 (
<collectionCode id="ED47AE574601FFDEFEABFC7B6FEEB5DE" box="[354,443,944,968]" httpUri="http://grbio.org/cool/dhtn-7yaw" name="National Museum of Kenya" pageId="1" pageNumber="302">NMKE</collectionCode>
).
</paragraph>
<paragraph id="8BE936924601FFDEFF0EFC1E6D8EB342" blockId="1.[151,1436,871,2013]" pageId="1" pageNumber="302">
<typeStatus id="54ED88304601FFDEFF0EFC1E6F62B5FB" box="[199,311,981,1005]" pageId="1" pageNumber="302" type="paratype">Paratypes</typeStatus>
, 74♀.
<collectingCountry id="F34176024601FFDEFE42FC1F6FBAB5FA" box="[395,495,980,1004]" name="Kenya" pageId="1" pageNumber="302">KENYA</collectingCountry>
: same data as
<typeStatus id="54ED88304601FFDEFD6BFC1F6D52B5FB" box="[674,775,980,1005]" pageId="1" pageNumber="302" type="holotype">holotype</typeStatus>
, except:
<collectionCode id="ED47AE574601FFDEFCBDFC1F6D92B5FB" box="[884,967,980,1005]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)31868, 31875, 31896, 31915, 31920, 31921, 31925, 31927, 31932, 32134, 32135, 32136, 32138, 32139, 32140, 32144, 32145, 32147, 32148, 32150, 32151, 32152, 32153, 32154, 32155, 32156, 32157, 32158, 32159, 32160, 32161, 32162, 32163, 32164, 32165, 32166, 32167, 32169, 32171, 32172, 32173, 32174, 32175, 32176, 32177, 32178, 32179, 32180, 32182, 32183, 32184, 32185, 32187, 32189, 32190, 32191, 32192, 32193, 32195, 32198, 32199, 32200 (all
<collectionCode id="ED47AE574601FFDEFB09FBAF6B41B26A" box="[1216,1300,1124,1148]" pageId="1" pageNumber="302">BMSA</collectionCode>
); 5♀, same, except:
<collectionCode id="ED47AE574601FFDEFF26FB436F17B2B6" box="[239,322,1160,1184]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)32203, 32204, 32207, 32209, 32211 (all
<collectionCode id="ED47AE574601FFDEFCE6FB436DD2B2B6" box="[815,903,1160,1184]" httpUri="http://grbio.org/cool/dhtn-7yaw" name="National Museum of Kenya" pageId="1" pageNumber="302">NMKE</collectionCode>
); 1♀, same, except:
<collectionCode id="ED47AE574601FFDEFBA2FB436AE8B2B6" box="[1131,1213,1160,1184]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)32143 (
<collectionCode id="ED47AE574601FFDEFAFFFB436BC5B2B6" box="[1334,1424,1160,1184]" pageId="1" pageNumber="302">BMNH</collectionCode>
); 2♀, same, except:
<collectionCode id="ED47AE574601FFDEFEBDFB676F93B2D2" box="[372,454,1196,1220]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)32149, 32196 (both
<collectionCode id="ED47AE574601FFDEFD12FB676D7EB2D2" box="[731,811,1196,1220]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="302">MNRJ</collectionCode>
).
<collectingCountry id="F34176024601FFDEFC88FB676DF0B2D2" box="[833,933,1196,1220]" name="Kenya" pageId="1" pageNumber="302">KENYA</collectingCountry>
: Rift Valley Prov., Nakuru/Eldoret road at: 00˚07.414′N, 35˚27.822′E,
<quantity id="4CAE9B774601FFDEFE04FB046C73B2F1" box="[461,550,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="2.5889999999999995" pageId="1" pageNumber="302" unit="m" value="2589.0">2589 m</quantity>
, general sweeping, mixed forest,
<date id="FFE810524601FFDEFC6EFB046A49B2FE" box="[935,1052,1231,1256]" pageId="1" pageNumber="302" value="2011-04-13">13.iv.2011</date>
(A.H. &amp; M.K. Kirk-Spriggs, 1♀,
<collectionCode id="ED47AE574601FFDEFF5EFB3F6EBCB31B" box="[151,233,1268,1293]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)32621 (
<collectionCode id="ED47AE574601FFDEFEADFB3F6FECB31A" box="[356,441,1268,1292]" pageId="1" pageNumber="302">BMSA</collectionCode>
).
<collectingCountry id="F34176024601FFDEFE04FB3E6C04B31A" box="[461,593,1269,1292]" name="Burundi" pageId="1" pageNumber="302">BURUNDI</collectingCountry>
: Kayanza Prov., Parc National de la Kibira, Rwegura Sector, 02˚55.320′S, 29˚30.067′E,
<quantity id="4CAE9B774601FFDEFEFCFAD36FDAB339" box="[309,399,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="2.2369999999999997" pageId="1" pageNumber="302" unit="m" value="2237.0">2237 m</quantity>
,
<date id="FFE810524601FFDEFE6DFADC6C1CB326" box="[420,585,1303,1328]" pageId="1" pageNumber="302" value="2010-11-21" valueMax="2010-11-26" valueMin="2010-11-21">21–26.xi.2010</date>
, 1♀,
<collectionCode id="ED47AE574601FFDEFD5BFAD36CB6B326" box="[658,739,1304,1328]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)28092 (
<collectionCode id="ED47AE574601FFDEFCAAFAD36DE1B326" box="[867,948,1304,1328]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="302">MNRJ</collectionCode>
); Malaise traps, indigenous Afromontane forest,
<date id="FFE810524601FFDEFF2DFAF76FD3B343" box="[228,390,1340,1365]" pageId="1" pageNumber="302" value="2010-11-25" valueMax="2010-11-26" valueMin="2010-11-25">25–26.xi.2010</date>
(A.H. Kirk-Spriggs), 2♀,
<collectionCode id="ED47AE574601FFDEFD67FAF76CAAB343" box="[686,767,1340,1365]" pageId="1" pageNumber="302">BMSA</collectionCode>
(D)24706 (
<collectionCode id="ED47AE574601FFDEFCB3FAF76D85B342" box="[890,976,1340,1364]" pageId="1" pageNumber="302">BMSA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C65194601FFDEFF0EFA946A51B01E" pageId="1" pageNumber="302" type="diagnosis">
<paragraph id="8BE936924601FFDEFF0EFA946A51B01E" blockId="1.[151,1436,871,2013]" pageId="1" pageNumber="302">
<emphasis id="B922EA804601FFDEFF0EFA946F14B36E" bold="true" box="[199,321,1375,1400]" pageId="1" pageNumber="302">Diagnosis.</emphasis>
Palpus yellow; prescutum with 2 well-separated brown maculae; anterior dorsal part of pleura yellow in ground colour, wholly pale dusted. The new species is morphologically close to
<taxonomicName id="4C564D114601FFDEFB44FA4E6BC2B38B" authority="Emden" authorityName="Emden" box="[1165,1431,1412,1437]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="poecilotarsis">
<emphasis id="B922EA804601FFDEFB44FA4E6AF5B38A" box="[1165,1184,1413,1436]" italics="true" pageId="1" pageNumber="302">C</emphasis>
.
<emphasis id="B922EA804601FFDEFB67FA4F6B6AB38B" box="[1198,1343,1412,1437]" italics="true" pageId="1" pageNumber="302">poecilotarsis</emphasis>
Emden
</taxonomicName>
, but can be easily separated by the presence in
<taxonomicName id="4C564D114601FFDEFD7FFA636D3EB3D6" box="[694,875,1447,1472]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="poecilotarsis">
<emphasis id="B922EA804601FFDEFD7FFA636C9CB3A9" box="[694,713,1448,1471]" italics="true" pageId="1" pageNumber="302">C</emphasis>
.
<emphasis id="B922EA804601FFDEFD13FA6C6D3EB3D6" box="[730,875,1447,1472]" italics="true" pageId="1" pageNumber="302">poecilotarsis</emphasis>
</taxonomicName>
of a black presutural transverse fascia, which is undivided although deeply incised posteriorly, the black palpus and a black patch covering the lower edge of postpronotal lobe, upper anterior corner of anepisternum and anterior spiracle.
</paragraph>
</subSubSection>
<subSubSection id="C34C65194601FFDBFF0EF9DF6F0CB756" lastPageId="4" lastPageNumber="305" pageId="1" pageNumber="302" type="description">
<paragraph id="8BE936924601FFDEFF0EF9DF6EB5B166" blockId="1.[151,1436,871,2013]" pageId="1" pageNumber="302">
<emphasis id="B922EA804601FFDEFF0EF9DF6F0CB03B" bold="true" box="[199,345,1556,1581]" pageId="1" pageNumber="302">Description.</emphasis>
<emphasis id="B922EA804601FFDEFEAAF9DF6C19B03B" box="[355,588,1556,1581]" italics="true" pageId="1" pageNumber="302">General colouration</emphasis>
. Ground colour brown, pale grey dusted; head with frons velvety black, brown pruinose (viewed from certain angles); fronto-orbital plate, face and gena pale grey pruinose; antenna with scape and pedicel dark yellow, with grey pruinosity and postpedicel dark brown, arista brown; palpus yellow; notum (
<figureCitation id="136D2A174601FFDEFF3AF9B46F6CB08E" box="[243,313,1663,1688]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,264,1956,1979]" captionTargetBox="[196,1429,467,1915]" captionTargetId="figure@2.[173,1458,445,1939]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1. Coenosia duomaculata sp. nov., notum, dorsal view; 2. Coenosia nigromaculata sp. nov., notum, dorsal view; 3. Helina harrisorum sp. nov., notum, dorsal view; 4. Helina ferfriniorum sp. nov., notum, dorsal view." httpUri="https://zenodo.org/record/229939/files/figure.png" pageId="1" pageNumber="302">Fig. 1</figureCitation>
) with prescutum with 2 well-separated brown maculae; postpronotal lobe yellow; scutum and scutellum each with transverse brown fascia in basal ½; pleura pale dusted, with yellowish dusting in some areas; anterior spiracle yellow; wing with very faint brown longitudinal infuscation along costal vein; calypters white; haltere pale yellow, with whitish knob; legs with coxae yellow, pale dusted; femora yellow, brown on dorsal surface; tibiae yellow and tarsi brown (some specimens with femur and tibia brown with yellow areas); abdomen pale dusted with some yellowish areas and elliptical brown lateral maculae on all tergites; tergites 3–5 with median fascia.
</paragraph>
<paragraph id="8BE936924601FFDEFF0EF8B66DDDB182" blockId="1.[151,1436,871,2013]" box="[199,904,1916,1941]" pageId="1" pageNumber="302">
♀.
<emphasis id="B922EA804601FFDEFF23F8B66FD9B182" box="[234,396,1917,1940]" italics="true" pageId="1" pageNumber="302">Measurements</emphasis>
. Body length:
<quantity id="4CAE9B774601FFDEFDFAF8B76CD9B182" box="[563,652,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="302" unit="mm" value="5.3">5.3 mm</quantity>
; wing length:
<quantity id="4CAE9B774601FFDEFCE2F8B76DD1B182" box="[811,900,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="302" unit="mm" value="6.2">6.2 mm</quantity>
.
</paragraph>
<paragraph id="8BE936924601FFDDFF0EF86B6B57B6C3" blockId="1.[151,1436,871,2013]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<emphasis id="B922EA804601FFDEFF0EF86B6F51B1AF" box="[199,260,1952,1977]" italics="true" pageId="1" pageNumber="302">Head</emphasis>
. Eye bare; frons at vertex ⅓ head width; 3–4 pairs of inclinate frontal setae, first and third from lunule shorter, and 1 pair of reclinate orbital setae; medial vertical seta extremely long; ocellar setae short; antenna inserted just above mid-level of eye, with postpedicel about 2.5 × length of pedicel; arista with very fine and short pubescence, longest setulae barely longer than basal width of arista; palpus with long ventral setulae.
</paragraph>
<paragraph id="8BE936924602FFDDFF0EFF146CF8B791" blockId="2.[151,1436,151,392]" pageId="2" pageNumber="303">
<emphasis id="B922EA804602FFDDFF0EFF146F4DB6EE" box="[199,280,223,248]" italics="true" pageId="2" pageNumber="303">Thorax</emphasis>
. Dorsocentrals 1 + 3; acrostichal setulae in 2 sparse irregular rows, prescutellar pair undifferentiated; 1 postpronotal; 1 postsutural intra-alar developed (the posterior); 1 postsutural supra-alar; 2 postalars; prealar absent; notopleuron with 2 setae similar in size; anepisternum with a series of 3 long and 2 short setae; katepisternals 1 + 1 + 1, arranged in an equilateral triangle. Scutellum with 1 long basal and 1 long apical pairs of setae, similar in length, with a few ground setulae close to apex.
</paragraph>
<caption id="DF29661A4602FFDDFF5EF86F6ABAB1CF" httpUri="https://zenodo.org/record/229939/files/figure.png" pageId="2" pageNumber="303" targetBox="[196,1429,467,1915]" targetPageId="2">
<paragraph id="8BE936924602FFDDFF5EF86F6ABAB1CF" blockId="2.[151,1436,1956,2009]" pageId="2" pageNumber="303">
<emphasis id="B922EA804602FFDDFF5EF86F6F05B1AC" bold="true" box="[151,336,1956,1979]" pageId="2" pageNumber="303">FIGURES 1–4. 1</emphasis>
.
<taxonomicName id="4C564D114602FFDDFEA9F86E6C19B1AC" box="[352,588,1956,1978]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="duomaculata" status="sp. nov.">
<emphasis id="B922EA804602FFDDFEA9F86E6C19B1AC" box="[352,588,1956,1978]" italics="true" pageId="2" pageNumber="303">Coenosia duomaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4602FFDDFD9DF86E6CF2B1AC" box="[596,679,1957,1978]" pageId="2" pageNumber="303" rank="species">
<emphasis id="B922EA804602FFDDFD9DF86E6CF2B1AC" bold="true" box="[596,679,1957,1978]" pageId="2" pageNumber="303">sp. nov.</emphasis>
</taxonomicNameLabel>
, notum, dorsal view;
<emphasis id="B922EA804602FFDDFC44F86E6DCCB1AC" bold="true" box="[909,921,1957,1978]" pageId="2" pageNumber="303">2</emphasis>
.
<taxonomicName id="4C564D114602FFDDFC61F86E6AF1B1AC" box="[936,1188,1956,1978]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="nigromaculata" status="sp. nov.">
<emphasis id="B922EA804602FFDDFC61F86E6AF1B1AC" box="[936,1188,1956,1978]" italics="true" pageId="2" pageNumber="303">Coenosia nigromaculata</emphasis>
</taxonomicName>
<emphasis id="B922EA804602FFDDFB64F86E6B53B1AC" bold="true" box="[1197,1286,1957,1978]" pageId="2" pageNumber="303">
<taxonomicNameLabel id="A21157FB4602FFDDFB64F86E6B54B1AC" box="[1197,1281,1957,1978]" pageId="2" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
notum, dorsal view;
<emphasis id="B922EA804602FFDDFF1CF8086EB4B1CE" bold="true" box="[213,225,1987,2008]" pageId="2" pageNumber="303">3</emphasis>
.
<taxonomicName id="4C564D114602FFDDFF26F8086FF8B1CF" box="[239,429,1987,2009]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="harrisorum" status="sp. nov.">
<emphasis id="B922EA804602FFDDFF26F8086FF8B1CF" box="[239,429,1987,2009]" italics="true" pageId="2" pageNumber="303">Helina harrisorum</emphasis>
</taxonomicName>
<emphasis id="B922EA804602FFDDFE7DF80F6C51B1CF" bold="true" box="[436,516,1988,2009]" pageId="2" pageNumber="303">
<taxonomicNameLabel id="A21157FB4602FFDDFE7DF80F6C51B1CF" box="[436,516,1988,2009]" pageId="2" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, notum, dorsal view;
<emphasis id="B922EA804602FFDDFD28F8086CB8B1CE" bold="true" box="[737,749,1987,2008]" pageId="2" pageNumber="303">4</emphasis>
.
<taxonomicName id="4C564D114602FFDDFD32F8086D96B1CF" box="[763,963,1987,2009]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="ferfriniorum" status="sp. nov.">
<emphasis id="B922EA804602FFDDFD32F8086D96B1CF" box="[763,963,1987,2009]" italics="true" pageId="2" pageNumber="303">Helina ferfriniorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4602FFDDFC00F80F6A4CB1CF" box="[969,1049,1988,2009]" pageId="2" pageNumber="303" rank="species">
<emphasis id="B922EA804602FFDDFC00F80F6A4CB1CF" bold="true" box="[969,1049,1988,2009]" pageId="2" pageNumber="303">sp. nov.</emphasis>
</taxonomicNameLabel>
, notum, dorsal view.
</paragraph>
</caption>
<caption id="DF29661A4603FFDCFF5EFA016F46B02B" httpUri="https://zenodo.org/record/229940/files/figure.png" pageId="3" pageNumber="304" targetBox="[191,1414,197,1445]" targetPageId="3">
<paragraph id="8BE936924603FFDCFF5EFA016F46B02B" blockId="3.[151,1436,1482,1597]" pageId="3" pageNumber="304">
<emphasis id="B922EA804603FFDCFF5EFA016F1DB3F6" bold="true" box="[151,328,1482,1505]" pageId="3" pageNumber="304">FIGURES 5–12.</emphasis>
<taxonomicName id="4C564D114603FFDCFE87FA006C6CB3F7" box="[334,569,1483,1505]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="duomaculata" status="sp. nov.">
<emphasis id="B922EA804603FFDCFE87FA006C6CB3F7" box="[334,569,1483,1505]" italics="true" pageId="3" pageNumber="304">Coenosia duomaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4603FFDCFD88FA006CC7B3F6" box="[577,658,1483,1504]" pageId="3" pageNumber="304" rank="species">
<emphasis id="B922EA804603FFDCFD88FA006CC7B3F6" bold="true" box="[577,658,1483,1504]" pageId="3" pageNumber="304">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="B922EA804603FFDCFD68FA006CF8B3F6" bold="true" box="[673,685,1483,1504]" pageId="3" pageNumber="304">5</emphasis>
. ovipositor, dorsal view;
<emphasis id="B922EA804603FFDCFC7BFA006DEBB3F6" bold="true" box="[946,958,1483,1504]" pageId="3" pageNumber="304">6</emphasis>
. ovipositor, ventral view and spermathecae;
<emphasis id="B922EA804603FFDCFA40FA006BC0B3F6" bold="true" box="[1417,1429,1483,1504]" pageId="3" pageNumber="304">7</emphasis>
. Egg.
<taxonomicName id="4C564D114603FFDCFF06FA216F98B3E9" box="[207,461,1513,1535]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="nigromaculata" status="sp. nov.">
<emphasis id="B922EA804603FFDCFF06FA216F98B3E9" box="[207,461,1513,1535]" italics="true" pageId="3" pageNumber="304">Coenosia nigromaculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4603FFDCFE1FFA216C7FB3E9" box="[470,554,1514,1535]" pageId="3" pageNumber="304" rank="species">
<emphasis id="B922EA804603FFDCFE1FFA216C7FB3E9" bold="true" box="[470,554,1514,1535]" pageId="3" pageNumber="304">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="B922EA804603FFDCFDF5FA226C1DB3E8" bold="true" box="[572,584,1513,1534]" pageId="3" pageNumber="304">8</emphasis>
. ovipositor, dorsal view;
<emphasis id="B922EA804603FFDCFC9EFA226D31B3E8" bold="true" box="[855,868,1513,1534]" pageId="3" pageNumber="304">9</emphasis>
. ovipositor, ventral view and spermathecae.
<taxonomicName id="4C564D114603FFDCFAF5FA216EB5B008" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="fragilis" status="sp. nov.">
<emphasis id="B922EA804603FFDCFAF5FA216EB5B008" italics="true" pageId="3" pageNumber="304">Coenosia fragilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4603FFDCFF2EF9C36F6DB00B" box="[231,312,1544,1565]" pageId="3" pageNumber="304" rank="species">
<emphasis id="B922EA804603FFDCFF2EF9C36F6DB00B" bold="true" box="[231,312,1544,1565]" pageId="3" pageNumber="304">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="B922EA804603FFDCFE8FF9C36F0AB00B" bold="true" box="[326,351,1544,1565]" pageId="3" pageNumber="304">10</emphasis>
. sternite 5, dorsal view;
<emphasis id="B922EA804603FFDCFD95F9C36C27B00B" bold="true" box="[604,626,1544,1565]" pageId="3" pageNumber="304">11</emphasis>
. cercal plate and surstyli, dorsal view;
<emphasis id="B922EA804603FFDCFBCDF9C36A48B00B" bold="true" box="[1028,1053,1544,1565]" pageId="3" pageNumber="304">12</emphasis>
. cercal plate, surstylus and aedeagus, lateral view.
</paragraph>
</caption>
<paragraph id="8BE936924603FFDCFF0EF9AE6A58B18B" blockId="3.[151,1436,1636,2024]" pageId="3" pageNumber="304">
<emphasis id="B922EA804603FFDCFF0EF9AE6EA9B06A" box="[199,252,1637,1660]" italics="true" pageId="3" pageNumber="304">Legs</emphasis>
. Fore femur with rows of posteroventral, posterior and posterodorsal setae, those in posterodorsal row stronger; fore tibia with 1 posterior median dorsal seta and 1 dorsal preapical, apical setae on posteroventral and posterior surfaces long; mid femur with 4–5 anterodorsal setae; posteroventral surface with closely spaced setae, without anteroventrals; 2 posterior preapicals; mid tibia with 1 median posterodorsal seta and 1 submedian anterodorsal; 1 dorsal preapical and anterodorsal, anterior, anteroventral, ventral and posteroventral apical setae, the last three stronger; hind femur with a row of about 4–5 well-separated anterodorsal setae and 1 preapical posterodorsal, 2 fine and long posteroventrals in basal ⅓, without anteroventrals, except for seta near base; hind tibia with strong anterior, anterodorsal and posterodorsal setae inserted proximally in median ⅓, but not at same level, 1 posterodorsal and 1 anterodorsal preapicals and 1 strong ventral apical.
</paragraph>
<paragraph id="8BE936924603FFDCFF0EF8626C63B1FE" blockId="3.[151,1436,1636,2024]" pageId="3" pageNumber="304">
<emphasis id="B922EA804603FFDCFF0EF8626EAAB1D6" box="[199,255,1961,1984]" italics="true" pageId="3" pageNumber="304">Wing</emphasis>
. Veins bare; veins
<emphasis id="B922EA804603FFDCFE1DF8626FBEB1D6" box="[468,491,1961,1984]" italics="true" pageId="3" pageNumber="304">M</emphasis>
and
<emphasis id="B922EA804603FFDCFDE8F8626C19B1D3" box="[545,588,1961,1989]" italics="true" pageId="3" pageNumber="304">R4+5</emphasis>
parallel at wing-tip; vein
<emphasis id="B922EA804603FFDCFCA4F8626D86B1D3" box="[877,979,1961,1989]" italics="true" pageId="3" pageNumber="304">A1+CuA2</emphasis>
very short; lower calypter slightly larger than upper calypter,
<emphasis id="B922EA804603FFDCFEB5F81B6FC2B1F1" box="[380,407,2000,2023]" italics="true" pageId="3" pageNumber="304">ca</emphasis>
1.5 × as long.
</paragraph>
<paragraph id="8BE936924604FFDBFF0EFF5C6F88B6C3" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="305">
<emphasis id="B922EA804604FFDBFF0EFF5C6F64B6A6" box="[199,305,151,176]" italics="true" pageId="4" pageNumber="305">Abdomen</emphasis>
. Syntergite 1 + 2 and tergites 3–5 each with 1 lateral pair of setae; tergites 4 and 5 with 2 additional median setae; sternite 1 bare.
</paragraph>
<paragraph id="8BE936924604FFDBFF0EFF146F1FB70A" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="305">
<taxonomicName id="4C564D114604FFDBFF0EFF146F14B6EE" box="[199,321,223,248]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="genus">
<emphasis id="B922EA804604FFDBFF0EFF146F14B6EE" box="[199,321,223,248]" italics="true" pageId="4" pageNumber="305">Terminalia</emphasis>
</taxonomicName>
. Ovipositor long, with microtrichia on both sides (
<figureCitation id="136D2A174604FFDBFCBDFF146D8FB6EE" box="[884,986,223,248]" captionStart="FIGURES 5 – 12" captionStartId="3.[151,264,1482,1505]" captionTargetBox="[191,1414,197,1445]" captionTargetId="figure@3.[151,1436,175,1461]" captionTargetPageId="3" captionText="FIGURES 5 – 12. Coenosia duomaculata sp. nov.: 5. ovipositor, dorsal view; 6. ovipositor, ventral view and spermathecae; 7. Egg. Coenosia nigromaculata sp. nov.: 8. ovipositor, dorsal view; 9. ovipositor, ventral view and spermathecae. Coenosia fragilis sp. nov.: 10. sternite 5, dorsal view; 11. cercal plate and surstyli, dorsal view; 12. cercal plate, surstylus and aedeagus, lateral view." httpUri="https://zenodo.org/record/229940/files/figure.png" pageId="4" pageNumber="305">Figs 5–6</figureCitation>
); spermathecae pear-shaped as in
<figureCitation id="136D2A174604FFDBFA9AFF146BC3B6EE" box="[1363,1430,223,248]" captionStart="FIGURES 5 – 12" captionStartId="3.[151,264,1482,1505]" captionTargetBox="[191,1414,197,1445]" captionTargetId="figure@3.[151,1436,175,1461]" captionTargetPageId="3" captionText="FIGURES 5 – 12. Coenosia duomaculata sp. nov.: 5. ovipositor, dorsal view; 6. ovipositor, ventral view and spermathecae; 7. Egg. Coenosia nigromaculata sp. nov.: 8. ovipositor, dorsal view; 9. ovipositor, ventral view and spermathecae. Coenosia fragilis sp. nov.: 10. sternite 5, dorsal view; 11. cercal plate and surstyli, dorsal view; 12. cercal plate, surstylus and aedeagus, lateral view." httpUri="https://zenodo.org/record/229940/files/figure.png" pageId="4" pageNumber="305">Fig. 6</figureCitation>
; egg as in
<figureCitation id="136D2A174604FFDBFEC8FECF6F13B70A" box="[257,326,260,285]" captionStart="FIGURES 5 – 12" captionStartId="3.[151,264,1482,1505]" captionTargetBox="[191,1414,197,1445]" captionTargetId="figure@3.[151,1436,175,1461]" captionTargetPageId="3" captionText="FIGURES 5 – 12. Coenosia duomaculata sp. nov.: 5. ovipositor, dorsal view; 6. ovipositor, ventral view and spermathecae; 7. Egg. Coenosia nigromaculata sp. nov.: 8. ovipositor, dorsal view; 9. ovipositor, ventral view and spermathecae. Coenosia fragilis sp. nov.: 10. sternite 5, dorsal view; 11. cercal plate and surstyli, dorsal view; 12. cercal plate, surstylus and aedeagus, lateral view." httpUri="https://zenodo.org/record/229940/files/figure.png" pageId="4" pageNumber="305">Fig. 7</figureCitation>
.
</paragraph>
<paragraph id="8BE936924604FFDBFF0EFEE36F0CB756" blockId="4.[151,1436,151,573]" box="[199,345,295,320]" pageId="4" pageNumber="305">♂ Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34C65194604FFDBFF0EFE876B7AB773" box="[199,1327,332,357]" pageId="4" pageNumber="305" type="distribution">
<paragraph id="8BE936924604FFDBFF0EFE876B7AB773" blockId="4.[151,1436,151,573]" box="[199,1327,332,357]" pageId="4" pageNumber="305">
<emphasis id="B922EA804604FFDBFF0EFE876F35B773" bold="true" box="[199,352,332,357]" pageId="4" pageNumber="305">Distribution.</emphasis>
<collectingCountry id="F34176024604FFDBFEAEFE866FE5B773" box="[359,432,333,357]" name="Kenya" pageId="4" pageNumber="305">Kenya</collectingCountry>
(Rift Valley forest, Mau Escarpment) and
<collectingCountry id="F34176024604FFDBFC47FE876DBFB773" box="[910,1002,332,357]" name="Burundi" pageId="4" pageNumber="305">Burundi</collectingCountry>
(Parc National de la Kibira).
</paragraph>
</subSubSection>
<subSubSection id="C34C65194604FFDBFF0EFEA46CDAB7D9" pageId="4" pageNumber="305" type="etymology">
<paragraph id="8BE936924604FFDBFF0EFEA46CDAB7D9" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="305">
<emphasis id="B922EA804604FFDBFF0EFEA46F13B79E" bold="true" box="[199,326,367,392]" pageId="4" pageNumber="305">Etymology</emphasis>
. The specific epithet
<taxonomicName id="4C564D114604FFDBFD97FEA46CA5B79E" box="[606,752,367,392]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="duomaculata">
<emphasis id="B922EA804604FFDBFD97FEA46CA5B79E" box="[606,752,367,392]" italics="true" pageId="4" pageNumber="305">duomaculata</emphasis>
</taxonomicName>
is derived from the Latin
<emphasis id="B922EA804604FFDBFB86FEA46A2CB79E" box="[1103,1145,367,392]" italics="true" pageId="4" pageNumber="305">duo</emphasis>
(“two”) and
<emphasis id="B922EA804604FFDBFAE0FEA46BC9B79E" box="[1321,1436,367,392]" italics="true" pageId="4" pageNumber="305">maculatus</emphasis>
(“maculate”), and refers to the two well-separated brown maculae present on the prescutum. It is an adjective in agreement with the feminine noun
<taxonomicName id="4C564D114604FFDBFDD7FE736CDDB7D9" box="[542,648,440,463]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B922EA804604FFDBFDD7FE736CDDB7D9" box="[542,648,440,463]" italics="true" pageId="4" pageNumber="305">Coenosia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65194604FFDBFF0EFE176AD2B42B" pageId="4" pageNumber="305" type="discussion">
<paragraph id="8BE936924604FFDBFF0EFE176AD2B42B" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="305">
<emphasis id="B922EA804604FFDBFF0EFE176F61B7E3" bold="true" box="[199,308,476,501]" pageId="4" pageNumber="305">Remarks</emphasis>
. The
<typeStatus id="54ED88304604FFDBFEB0FE166FFCB7E3" box="[377,425,477,501]" pageId="4" pageNumber="305">type</typeStatus>
material was compared by ACP with the
<typeStatus id="54ED88304604FFDBFC59FE176DA7B7E3" box="[912,1010,476,501]" pageId="4" pageNumber="305" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C564D114604FFDBFBD5FE166A85B7E3" box="[1052,1232,476,501]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="poecilotarsis">
<emphasis id="B922EA804604FFDBFBD5FE166A7AB7E2" box="[1052,1071,477,500]" italics="true" pageId="4" pageNumber="305">C</emphasis>
.
<emphasis id="B922EA804604FFDBFBF6FE176A85B7E3" box="[1087,1232,476,501]" italics="true" pageId="4" pageNumber="305">poecilotarsis</emphasis>
</taxonomicName>
in the BMNH. A second female of the same species named by Emden and also examined by ACP, also has these characters, but the femora are entirely black and the tibiae are mostly black with basal parts red to yellowish.
</paragraph>
</subSubSection>
</treatment>
</document>
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