338 lines
36 KiB
XML
338 lines
36 KiB
XML
<document id="332BEF4CA47239D3CA411CC80D757A01" ID-DOI="10.5281/zenodo.195826" ID-GBIF-Dataset="71d372a2-19b7-4578-95b7-95bef047abd9" ID-ISSN="1175-5326" ID-Zenodo-Dep="195826" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460234808519" checkinUser="plazi" docAuthor="Renoult, Julien P., Geniez, Philippe, Bacquet, Paul, Guillaume, Claude P. & Crochet, Pierre-Andre" docDate="2010" docId="03F235737E28FFA3FF27C4F72758F84F" docLanguage="en" docName="zt02500p068.pdf" docOrigin="Zootaxa 2500" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Podarcis liolepis Boulenger 1905" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFCB4D0B7E2EFFA4FFB0C262233DFFBD" masterDocTitle="Systematics of the Podarcis hispanicus - complex (Sauria, Lacertidae) II: the valid name of the north-eastern Spanish form" masterLastPageNumber="68" masterPageNumber="58" pageNumber="64" updateTime="1698239255788" updateUser="plazi">
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<mods:titleInfo id="5AA5DCE93D822A566D51A47EFBF050B1">
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<mods:title id="3FE11893504918417D10BF8918604802">Systematics of the Podarcis hispanicus - complex (Sauria, Lacertidae) II: the valid name of the north-eastern Spanish form</mods:title>
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<mods:name id="113A2051307827BC02C75E4238BF9C14" type="personal">
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<mods:namePart id="8B6B9262E99C5D6383AED6BB76144192">Renoult, Julien P.</mods:namePart>
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<mods:roleTerm id="C4D63BE2CD4FD7808AA325EF65E6A8AB">Author</mods:roleTerm>
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<mods:namePart id="325E5676C27380D2B1728996749F32E5">Geniez, Philippe</mods:namePart>
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</mods:name>
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<mods:name id="CFEA86DEA87B1B133FF71C7C69382AC2" type="personal">
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<mods:roleTerm id="7CA17C30A432C4471752FEE5B20E5A39">Author</mods:roleTerm>
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<mods:namePart id="DDA7C7CC186AEC496EF8E34C4B7757DA">Bacquet, Paul</mods:namePart>
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<mods:name id="965D4206C385377267DC78ACCBF1134C" type="personal">
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<mods:roleTerm id="0680A7D5BE0530709DC21F1E41038B65">Author</mods:roleTerm>
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<mods:namePart id="846DC2DDFE833588249EB5CA0BA3208A">Guillaume, Claude P.</mods:namePart>
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</mods:name>
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<mods:name id="219C0594CF5971A272144361C6061524" type="personal">
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<mods:namePart id="0C6ABFF1A60D65F846784F284D3F110D">Crochet, Pierre-Andre</mods:namePart>
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<mods:typeOfResource id="EEBA82CF6A558285FE88DFA609EC2394">text</mods:typeOfResource>
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<mods:title id="1F43866F6495BB49E54FD0176A07AA3C">Zootaxa</mods:title>
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<mods:part id="AF541499B917515695DBB8E37B65E943">
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<mods:date id="30A09A90C3C213DEABAEE15C15A05DD3">2010</mods:date>
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<mods:number id="B4E9F131F6C8611A4497A359C4E5DB70">2500</mods:number>
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<mods:identifier id="12D14DDDC5828DBE5B9AF0FE3247F2AE" type="DOI">10.5281/zenodo.195826</mods:identifier>
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<mods:identifier id="DC7EB09106E553AED6C3A133A648FBBC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="00D79109C9191B12F68C11CEF6925564" type="Zenodo-Dep">195826</mods:identifier>
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<treatment id="03F235737E28FFA3FF27C4F72758F84F" ID-DOI="http://doi.org/10.5281/zenodo.6200842" ID-GBIF-Taxon="119387844" ID-Zenodo-Dep="6200842" LSID="urn:lsid:plazi:treatment:03F235737E28FFA3FF27C4F72758F84F" httpUri="http://treatment.plazi.org/id/03F235737E28FFA3FF27C4F72758F84F" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="64">
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<subSubSection id="C341D7EE7E28FFA2FF27C4F7217DF912" box="[151,576,1685,1711]" pageId="6" pageNumber="64" type="nomenclature">
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<paragraph id="8BE484657E28FFA2FF27C4F7217DF912" blockId="6.[151,576,1685,1711]" box="[151,576,1685,1711]" pageId="6" pageNumber="64">
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<heading id="D0AC33097E28FFA2FF27C4F7217DF912" bold="true" box="[151,576,1685,1711]" fontSize="11" level="1" pageId="6" pageNumber="64" reason="1">
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<taxonomicName id="4C5BFFE67E28FFA2FF27C4F7217DF912" authority="Boulenger, 1905" authorityName="Boulenger" authorityYear="1905" box="[151,576,1685,1711]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="liolepis">
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<emphasis id="B92F58777E28FFA2FF27C4F7217DF912" bold="true" box="[151,576,1685,1711]" pageId="6" pageNumber="64">
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<emphasis id="B92F58777E28FFA2FF27C4F72263F912" bold="true" box="[151,350,1685,1711]" italics="true" pageId="6" pageNumber="64">Podarcis liolepis</emphasis>
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(
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<bibRefCitation id="EFCAF9947E28FFA2FEDDC4F72105F912" author="Boulenger" box="[365,568,1685,1711]" pageId="6" pageNumber="66" refString="Boulenger, G. A. (1905) A contribution to our knowledge of the varieties of the Wall-Lizard (Lacerta muralis) in Western Europe and North Africa. Transactions of the Zoological Society of London, 17, 351 - 437." type="journal article" year="1905">Boulenger, 1905</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C341D7EE7E28FFA3FF27C4BF22E4FF72" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="64" type="reference_group">
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<paragraph id="8BE484657E28FFA2FF27C4BF20C7F8C9" blockId="6.[151,1436,1757,1908]" pageId="6" pageNumber="64">
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<taxonomicName id="4C5BFFE67E28FFA2FF27C4BF227CF949" box="[151,321,1757,1780]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="muralis">
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<emphasis id="B92F58777E28FFA2FF27C4BF227CF949" box="[151,321,1757,1780]" italics="true" pageId="6" pageNumber="64">Lacerta muralis</emphasis>
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</taxonomicName>
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<taxonomicName id="4C5BFFE67E28FFA2FEF9C4BF21B8F949" authority="Boulenger, 1905" authorityName="Boulenger" authorityYear="1905" box="[329,645,1757,1780]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="variety" species="bocagei" variety="liolepis">
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var.
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<emphasis id="B92F58777E28FFA2FEC6C4BF22F9F949" box="[374,452,1757,1780]" italics="true" pageId="6" pageNumber="64">liolepis</emphasis>
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,
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<bibRefCitation id="EFCAF9947E28FFA2FE62C4BF21B8F949" author="Boulenger" box="[466,645,1757,1780]" pageId="6" pageNumber="66" refString="Boulenger, G. A. (1905) A contribution to our knowledge of the varieties of the Wall-Lizard (Lacerta muralis) in Western Europe and North Africa. Transactions of the Zoological Society of London, 17, 351 - 437." type="journal article" year="1905">Boulenger, 1905</bibRefCitation>
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</taxonomicName>
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, Transactions of the zoological Society of London 17: 363. Name-bearing
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<typeStatus id="54E03AC77E28FFA2FF77C49F23C4F8A9" box="[199,249,1789,1812]" pageId="6" pageNumber="64">type</typeStatus>
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: Natural History Museum (London) BMNH 86.12.29.3, an adult male from Valencia, coll. or leg by Prof. E. Boscá,
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<typeStatus id="54E03AC77E28FFA2FEA6C57F2245F889" box="[278,376,1821,1844]" pageId="6" pageNumber="64" type="lectotype">lectotype</typeStatus>
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by present designation (see
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<figureCitation id="136098E07E28FFA2FD19C57F21D4F889" box="[681,745,1821,1844]" captionStart="FIGURE 2" captionStartId="5.[151,255,1073,1097]" captionTargetBox="[163,1424,208,1038]" captionTargetId="figure@5.[151,1436,197,1049]" captionTargetPageId="5" captionText="FIGURE 2. Latero-dorsal views of Podarcis liolepis (Boulenger, 1905). A. Specimen BMNH 86.12.29.3, lectotype of Lacerta muralis var. liolepis Boulenger, 1905, Valencia, Spain. Photo C. McCarthy. B. Specimen BEV 9853 with “ Valencia ” mitochondrial lineage (mtDNA), male, collected in Rafelbuñol, north of Valencia, Spain. Photo PGe. C. Specimen BEV 9815 with “ Type 3 ” mtDNA, male, collected 3 km NW of Miami-Platja along the national road 340, province Tarragone, Spain. Photo PAC. Note the great morphological resemblance of both Podarcis liolepis (BEV 9853 and BEV 9815) showing however a deep mitochondrial divergence" httpUri="https://zenodo.org/record/195828/files/figure.png" pageId="6" pageNumber="64">Fig. 2</figureCitation>
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A). The list of the known original
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<typeStatus id="54E03AC77E28FFA2FBEAC57F278AF889" box="[1114,1207,1821,1844]" pageId="6" pageNumber="64" type="syntype">syntypes</typeStatus>
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is given above.
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<typeStatus id="54E03AC77E28FFA2FAD7C57F26A1F889" box="[1383,1436,1821,1844]" pageId="6" pageNumber="64">Type</typeStatus>
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locality: Valencia,
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<collectingCountry id="F34CC4F57E28FFA2FE3FC55F22F2F8E9" box="[399,463,1853,1876]" name="Spain" pageId="6" pageNumber="64">Spain</collectingCountry>
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, by present
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<typeStatus id="54E03AC77E28FFA2FDE1C55F218EF8E9" box="[593,691,1853,1876]" pageId="6" pageNumber="64" type="lectotype">lectotype</typeStatus>
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designation. Originally “Valencia”, “Foyos” (= Foios) near Valencia, “Sevilla”, and along the road from Valencia to Dehesa de la Albufera,
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<collectingCountry id="F34CC4F57E28FFA2FC06C53F20CBF8C9" box="[950,1014,1885,1908]" name="Spain" pageId="6" pageNumber="64">Spain</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BE484657E28FFA2FF27C5FF227FF809" blockId="6.[151,1437,1949,2036]" box="[151,322,1949,1972]" pageId="6" pageNumber="64">Junior synonym</paragraph>
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<paragraph id="8BE484657E28FFA3FF27C5DF22E4FF72" blockId="6.[151,1437,1949,2036]" lastBlockId="7.[199,1434,152,207]" lastPageId="7" lastPageNumber="65" pageId="6" pageNumber="64">
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<taxonomicName id="4C5BFFE67E28FFA2FF27C5DF227FF869" box="[151,322,1981,2004]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="64" phylum="Chordata" rank="species" species="muralis">
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<emphasis id="B92F58777E28FFA2FF27C5DF227FF869" box="[151,322,1981,2004]" italics="true" pageId="6" pageNumber="64">Lacerta muralis</emphasis>
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</taxonomicName>
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<taxonomicName id="4C5BFFE67E28FFA2FEF9C5DF21ABF869" authority="Schreiber, 1912" authorityName="Schreiber" authorityYear="1912" box="[329,662,1981,2004]" pageId="6" pageNumber="64" rank="variety" variety="hesperica">
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var.
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<emphasis id="B92F58777E28FFA2FEC6C5DF22E3F869" box="[374,478,1981,2004]" italics="true" pageId="6" pageNumber="64">hesperica</emphasis>
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,
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<bibRefCitation id="EFCAF9947E28FFA2FE5CC5DF21ABF869" author="Schreiber" box="[492,662,1981,2004]" pageId="6" pageNumber="68" refString="Schreiber, E. (1912) Herpetologia europaea. Eine systematische Bearbeitung der Amphibien und Reptilien welche bisher in Europa aufgefunden sind. Verlag von Gustav Fischer, Jena." type="book" year="1912">Schreiber, 1912</bibRefCitation>
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</taxonomicName>
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, Herpetologia europaea. Eine systematische Bearbeitung der Amphibien und Reptilien welche bisher in Europa aufgefunden sind: 943. Name-bearing
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<typeStatus id="54E03AC77E28FFA2FB9CC5BF2755F849" box="[1068,1128,2013,2036]" pageId="6" pageNumber="64">types</typeStatus>
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: Five
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<typeStatus id="54E03AC77E28FFA2FB01C5BF262CF849" box="[1201,1297,2013,2036]" pageId="6" pageNumber="64" type="syntype">syntypes</typeStatus>
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, still extant: Naturhistorisches Museum Wien NMW 16020 and NMW 16021: 1-4,
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<typeStatus id="54E03AC77E29FFA3FC6FC2FA2701FF12" box="[991,1084,152,175]" pageId="7" pageNumber="65" type="syntype">syntypes</typeStatus>
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(
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<bibRefCitation id="EFCAF9947E29FFA3FBFFC2FA266CFF12" author="Tiedemann" box="[1103,1361,152,175]" pageId="7" pageNumber="68" refString="Tiedemann, F., Haupl, M. & Grillitsch, H. (1994) Katalog der Typen der herpetologischen Sammlung nach dem Stand vom Janner 1994, Teil II: Reptilia. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, Wien. 10 (Vertebrata 4): Selbstverlag Naturhistorisches Museum, Wien, 110 pp." type="book" year="1994">
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Tiedemann
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<emphasis id="B92F58777E29FFA3FB63C2FA2630FF12" box="[1235,1293,152,175]" italics="true" pageId="7" pageNumber="65">et al.</emphasis>
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1994
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</bibRefCitation>
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).
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<typeStatus id="54E03AC77E29FFA3FAD7C2FA26A7FF12" box="[1383,1434,152,175]" pageId="7" pageNumber="65">Type</typeStatus>
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locality: Zaragoza,
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<collectingCountry id="F34CC4F57E29FFA3FE25C2DA22E8FF72" box="[405,469,184,207]" name="Spain" pageId="7" pageNumber="65">Spain</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C341D7EE7E29FFA3FF27C29F2774FC2F" pageId="7" pageNumber="65" type="discussion">
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<paragraph id="8BE484657E29FFA3FF27C29F2736FE4A" blockId="7.[151,1437,253,2034]" pageId="7" pageNumber="65">
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<emphasis id="B92F58777E29FFA3FF27C29F222BFEAA" bold="true" box="[151,278,253,279]" pageId="7" pageNumber="65">Remarks:</emphasis>
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1) This nomen was treated as an infrasubspecific nomen by
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<bibRefCitation id="EFCAF9947E29FFA3FC4FC29F2655FEAA" author="Mertens" box="[1023,1384,253,279]" pageId="7" pageNumber="67" refString="Mertens, R. & Wermuth, H. (1960) Die Amphibien und Reptilien Europas. Verlag Waldemar Kramer, Frankfurt am Main, 264 pp." type="book" year="1960">Mertens and Wermuth (1960)</bibRefCitation>
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and
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<bibRefCitation id="EFCAF9947E29FFA3FF27C3402266FE81" author="Salvador" box="[151,347,290,316]" pageId="7" pageNumber="68" refString="Salvador, A. (1986) Podarcis hispanica (Steindachner, 1870) - Iberische Mauereidechse. In Bohme, W. (Eds), Handbuch der Reptilien und Amphibien Europas, Band 2 (2), Echsen 3. Aula, Wiesbaden, pp. 71 - 82." type="journal article" year="1986">
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<collectingCountry id="F34CC4F57E29FFA3FF27C340223CFE81" box="[151,257,290,316]" name="El Salvador" pageId="7" pageNumber="65">Salvador</collectingCountry>
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(1986)
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</bibRefCitation>
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, and therefore as not available (“nomen illegitimum”). This is clearly a mistake:
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<bibRefCitation id="EFCAF9947E29FFA3FA9AC34023D9FEDF" author="Schreiber" pageId="7" pageNumber="68" refString="Schreiber, E. (1912) Herpetologia europaea. Eine systematische Bearbeitung der Amphibien und Reptilien welche bisher in Europa aufgefunden sind. Verlag von Gustav Fischer, Jena." type="book" year="1912">Schreiber (1912)</bibRefCitation>
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described
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<emphasis id="B92F58777E29FFA3FED4C32A22EBFEDC" box="[356,470,328,353]" italics="true" pageId="7" pageNumber="65">hesperica</emphasis>
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as a variety of
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<taxonomicName id="4C5BFFE67E29FFA3FD39C32A2078FEDC" box="[649,837,328,353]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="muralis">
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<emphasis id="B92F58777E29FFA3FD39C32A2078FEDC" box="[649,837,328,353]" italics="true" pageId="7" pageNumber="65">Lacerta muralis</emphasis>
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</taxonomicName>
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(see heading of page 943), treated it as a variety of
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<taxonomicName id="4C5BFFE67E29FFA3FF27C30F23CEFE3B" box="[151,243,365,390]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="muralis">
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<emphasis id="B92F58777E29FFA3FF27C30F23CEFE3B" box="[151,243,365,390]" italics="true" pageId="7" pageNumber="65">muralis</emphasis>
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</taxonomicName>
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(see page 410). This is confirmed by the original labels of the
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<emphasis id="B92F58777E29FFA3FC44C30F2754FE3B" box="[1012,1129,365,390]" italics="true" pageId="7" pageNumber="65">hesperica</emphasis>
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<typeStatus id="54E03AC77E29FFA3FBC1C30F27E7FE3A" box="[1137,1242,365,391]" pageId="7" pageNumber="65" type="syntype">syntypes</typeStatus>
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in Wien, which writes as: “NMW 16020,
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<taxonomicName id="4C5BFFE67E29FFA3FE7EC3F0200CFE11" authority="Blgr" authorityName="Blgr" box="[462,817,402,428]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="subSpecies" species="bocagei" subSpecies="liolepis">
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<emphasis id="B92F58777E29FFA3FE7EC3F021CEFE16" box="[462,755,402,427]" italics="true" pageId="7" pageNumber="65">Lacerta bocagei liolepis</emphasis>
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Blgr
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</taxonomicName>
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(
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<typeStatus id="54E03AC77E29FFA3FCF1C3F020F5FE11" box="[833,968,402,428]" pageId="7" pageNumber="65" type="lectotype">Lectotypus</typeStatus>
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of
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|
<taxonomicName id="4C5BFFE67E29FFA3FC42C3F02750FE16" box="[1010,1133,402,427]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="muralis">
|
|
<emphasis id="B92F58777E29FFA3FC42C3F02750FE16" box="[1010,1133,402,427]" italics="true" pageId="7" pageNumber="65">L. muralis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicName id="4C5BFFE67E29FFA3FBC6C3F026B2FE11" authority="Schreib." authorityName="Schreib." box="[1142,1423,402,428]" pageId="7" pageNumber="65" rank="variety" variety="hesperica">
|
|
var.
|
|
<emphasis id="B92F58777E29FFA3FB1BC3F0261DFE16" box="[1195,1312,402,427]" italics="true" pageId="7" pageNumber="65">hesperica</emphasis>
|
|
Schreib.
|
|
</taxonomicName>
|
|
), Saragossa, Southoff leg,
|
|
<date id="FFE5A2A57E29FFA3FE7EC3DA2171FE6F" box="[462,588,440,466]" pageId="7" pageNumber="65" value="1912-08">VIII.1912</date>
|
|
, Coll. Schreiber / NMW 16021:1-4,
|
|
<taxonomicName id="4C5BFFE67E29FFA3FBA6C3DA27E0FE6C" box="[1046,1245,440,465]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="muralis">
|
|
<emphasis id="B92F58777E29FFA3FBA6C3DA27E0FE6C" box="[1046,1245,440,465]" italics="true" pageId="7" pageNumber="65">Lacerta muralis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicName id="4C5BFFE67E29FFA3FB56C3DA2231FE4A" authority="Saragossa" authorityName="Saragossa" pageId="7" pageNumber="65" rank="variety" variety="hesperica">
|
|
var.
|
|
<emphasis id="B92F58777E29FFA3FAACC3DA26A9FE6C" box="[1308,1428,440,465]" italics="true" pageId="7" pageNumber="65">hesperica</emphasis>
|
|
, Saragossa
|
|
</taxonomicName>
|
|
,
|
|
<collectingCountry id="F34CC4F57E29FFA3FEABC3BF2241FE4A" box="[283,380,477,503]" name="Spain" pageId="7" pageNumber="65">Spanien</collectingCountry>
|
|
, 1915.
|
|
<date id="FFE5A2A57E29FFA3FE79C3BF213FFE4A" box="[457,514,477,503]" pageId="7" pageNumber="65" value="1922-02">II.22</date>
|
|
E, Coll. Schreiber, Nr. 84,89, leg. Southoff”.
|
|
</paragraph>
|
|
<paragraph id="8BE484657E29FFA3FF76C060219CFDDA" blockId="7.[151,1437,253,2034]" pageId="7" pageNumber="65">
|
|
2) Despite the mention on the labels that one of the specimen is a
|
|
<typeStatus id="54E03AC77E29FFA3FC47C060275BFDA1" box="[1015,1126,514,540]" pageId="7" pageNumber="65" type="lectotype">lectotype</typeStatus>
|
|
, neither
|
|
<bibRefCitation id="EFCAF9947E29FFA3FB61C06023D9FDFF" pageId="7" pageNumber="68" refString="Tiedemann, F., Haupl, M. & Grillitsch, H. (1994) Katalog der Typen der herpetologischen Sammlung nach dem Stand vom Janner 1994, Teil II: Reptilia. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, Wien. 10 (Vertebrata 4): Selbstverlag Naturhistorisches Museum, Wien, 110 pp." type="book">
|
|
Tiedemann
|
|
<emphasis id="B92F58777E29FFA3FAEEC06026A1FDA6" box="[1374,1436,514,539]" italics="true" pageId="7" pageNumber="65">et al.</emphasis>
|
|
(1994)
|
|
</bibRefCitation>
|
|
nor ourselves were able to find a published
|
|
<typeStatus id="54E03AC77E29FFA3FD42C04A2060FDFF" box="[754,861,552,578]" pageId="7" pageNumber="65" type="lectotype">lectotype</typeStatus>
|
|
designation for
|
|
<emphasis id="B92F58777E29FFA3FBAEC04A27ADFDFC" box="[1054,1168,552,577]" italics="true" pageId="7" pageNumber="65">hesperica</emphasis>
|
|
. The specimens of the original
|
|
<typeStatus id="54E03AC77E29FFA3FF49C02F2211FDDA" box="[249,300,589,615]" pageId="7" pageNumber="65">type</typeStatus>
|
|
series thus all remain
|
|
<typeStatus id="54E03AC77E29FFA3FD82C02F21A1FDDA" box="[562,668,589,615]" pageId="7" pageNumber="65" type="syntype">syntypes</typeStatus>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BE484657E29FFA3FF76C0102771FD6A" blockId="7.[151,1437,253,2034]" pageId="7" pageNumber="65">
|
|
Proposed status: subjective junior synonym of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FD45C0102080FD36" box="[757,957,626,651]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FD45C0102080FD36" box="[757,957,626,651]" italics="true" pageId="7" pageNumber="65">Podarcis liolepis</emphasis>
|
|
</taxonomicName>
|
|
. This nomen would be available for the northern TYP3 populations (carrying the “
|
|
<typeStatus id="54E03AC77E29FFA3FD3DC0FA21F6FD0F" box="[653,715,664,690]" pageId="7" pageNumber="65">Type</typeStatus>
|
|
3” mtDNA lineage) if they were recognised as distinct from the southern TYP3 ones (carrying the south-eastern “Valencia” mtDNA lineage).
|
|
</paragraph>
|
|
<paragraph id="8BE484657E29FFA3FF76C0802774FC2F" blockId="7.[151,1437,253,2034]" pageId="7" pageNumber="65">
|
|
A series of other nomina apply to various populations of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FC36C080276DFD46" box="[902,1104,738,763]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FC36C080276DFD46" box="[902,1104,738,763]" italics="true" pageId="7" pageNumber="65">Podarcis liolepis</emphasis>
|
|
</taxonomicName>
|
|
. They are either subjective junior synonyms of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FE22C16A22D3FC9C" box="[402,494,776,801]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FE22C16A22D3FC9C" box="[402,494,776,801]" italics="true" pageId="7" pageNumber="65">liolepis</emphasis>
|
|
</taxonomicName>
|
|
(if
|
|
<taxonomicName id="4C5BFFE67E29FFA3FDAEC16A21CCFC9C" box="[542,753,776,801]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FDAEC16A21CCFC9C" box="[542,753,776,801]" italics="true" pageId="7" pageNumber="65">Podarcis liolepis</emphasis>
|
|
</taxonomicName>
|
|
is treated as a monotypic species) or valid nomina for subspecies of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FEF1C14F2130FCFB" box="[321,525,813,838]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FEF1C14F2130FCFB" box="[321,525,813,838]" italics="true" pageId="7" pageNumber="65">Podarcis liolepis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<collectingCountry id="F34CC4F57E29FFA3FDADC14F2103FCFA" box="[541,574,813,839]" name="American Samoa" pageId="7" pageNumber="65">As</collectingCountry>
|
|
a review of the morphological variation and subspecific systematics of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FF27C130225BFCD6" box="[151,358,850,875]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FF27C130225BFCD6" box="[151,358,850,875]" italics="true" pageId="7" pageNumber="65">Podarcis liolepis</emphasis>
|
|
</taxonomicName>
|
|
is beyond the scope of this paper, we do not address their status here but will do it in a forthcoming work dedicated to the infraspecific systematics of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FCCBC11A277CFC2C" box="[891,1089,888,913]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FCCBC11A277CFC2C" box="[891,1089,888,913]" italics="true" pageId="7" pageNumber="65">Podarcis liolepis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C341D7EE7E29FFA3FF76C1FF2016F871" pageId="7" pageNumber="65" type="diagnosis">
|
|
<paragraph id="8BE484657E29FFA3FF76C1FF2142F83F" blockId="7.[151,1437,253,2034]" pageId="7" pageNumber="65">
|
|
<emphasis id="B92F58777E29FFA3FF76C1FF2275FC0A" bold="true" box="[198,328,925,951]" pageId="7" pageNumber="65">Diagnosis:</emphasis>
|
|
a member of the
|
|
<taxonomicName id="4C5BFFE67E29FFA3FD87C1FF2017FC0B" box="[567,810,925,950]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="hispanicus">
|
|
<emphasis id="B92F58777E29FFA3FD87C1FF2017FC0B" box="[567,810,925,950]" italics="true" pageId="7" pageNumber="65">Podarcis hispanicus</emphasis>
|
|
</taxonomicName>
|
|
complex characterized by the combination of the following characters: masseteric shield and dark vertebral stripe frequently present but most often discontinuous, dark vertebral stripe never splits near its anterior part, generally lack of yellow coloration on the belly and under tail, pale dorsolateral stripes wider than the dark supradorsolateral stripes except in uniformly coloured individuals. Dorsum generally brown or greyish but often green in some populations from the southernmost parts of the range. Belly and undertail white, salmon-pink or brick red, never yellowish (but females can have yellow under the throat). When present, combination of a dark vertebral stripe and a distinct masseteric shield on one or two sides excludes all other taxa in this complex except
|
|
<taxonomicName id="4C5BFFE67E29FFA3FBDCC6C126A1FB01" box="[1132,1436,1187,1212]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="subSpecies" species="carbonelli" subSpecies="berlengensis">
|
|
<emphasis id="B92F58777E29FFA3FBDCC6C126A1FB01" box="[1132,1436,1187,1212]" italics="true" pageId="7" pageNumber="65">P. carbonelli berlengensis</emphasis>
|
|
</taxonomicName>
|
|
and some TYP
|
|
<specimenCount id="9D5D4FEC7E29FFA3FEE1C6AA22DCFB5F" box="[337,481,1224,1250]" pageId="7" pageNumber="65" type="generic">1 specimens</specimenCount>
|
|
(in which the dark vertebral stripe would be limited to the anterior part of the back). In addition, TYP1 (both 1A and 1B lineages, see
|
|
<bibRefCitation id="EFCAF9947E29FFA3FCD6C68F277EFABA" author="Pinho" box="[870,1091,1261,1287]" pageId="7" pageNumber="67" refString="Pinho, C., Ferrand, N. & Harris, D. J. (2006) Reexamination of the Iberian and North African Podarcis (Squamata: Lacertidae) phylogeny based on increased mitochondrial DNA sequencing. Molecular Phylogenetics and Evolution, 38, 266 - 273." type="journal article" year="2006">
|
|
Pinho
|
|
<emphasis id="B92F58777E29FFA3FC08C68F20C4FABB" box="[952,1017,1261,1286]" italics="true" pageId="7" pageNumber="65">et al.</emphasis>
|
|
2006
|
|
</bibRefCitation>
|
|
) differs from
|
|
<taxonomicName id="4C5BFFE67E29FFA3FB46C68F2649FABB" box="[1270,1396,1261,1286]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FB46C68F2649FABB" box="[1270,1396,1261,1286]" italics="true" pageId="7" pageNumber="65">P. liolepis</emphasis>
|
|
</taxonomicName>
|
|
by contrastingly white or pale dorsolateral stripes (usually less contrasting in
|
|
<emphasis id="B92F58777E29FFA3FBB0C771274DFA91" box="[1024,1136,1299,1324]" italics="true" pageId="7" pageNumber="65">P. l i o l e p i s</emphasis>
|
|
, rarely white, sometimes absent) which are regularly and strongly dashed in males and some females (less regularly fragmented in
|
|
<taxonomicName id="4C5BFFE67E29FFA3FA36C75A23CCFACB" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FA36C75A23CCFACB" italics="true" pageId="7" pageNumber="65">P. liolepis</emphasis>
|
|
</taxonomicName>
|
|
) or continuous in the other females, and narrower than the black supradorsolateral stripes in both sexes (as wide as or wider than the black supradorsolaterals in
|
|
<taxonomicName id="4C5BFFE67E29FFA3FC3CC7E1273FFA21" box="[908,1026,1411,1436]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FC3CC7E1273FFA21" box="[908,1026,1411,1436]" italics="true" pageId="7" pageNumber="65">P. liolepis</emphasis>
|
|
</taxonomicName>
|
|
). Those
|
|
<taxonomicName id="4C5BFFE67E29FFA3FBDDC7E127DEFA21" box="[1133,1251,1411,1436]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FBDDC7E127DEFA21" box="[1133,1251,1411,1436]" italics="true" pageId="7" pageNumber="65">P. liolepis</emphasis>
|
|
</taxonomicName>
|
|
specimens that lack a dark vertebral stripe need to be separated from the other taxa by additional characters. TYP2 (which never shows a dark vertebral stripe) also differs from
|
|
<taxonomicName id="4C5BFFE67E29FFA3FCBCC7AF205FFA5B" box="[780,866,1485,1510]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FCBCC7AF205FFA5B" box="[780,866,1485,1510]" italics="true" pageId="7" pageNumber="65">liolepis</emphasis>
|
|
</taxonomicName>
|
|
by the presence of a yellowish coloration on the rear belly and under tail and by wider dark supradorsolateral stripes (which are usually made of scattered marbling in males, more continuous in females) and the typically green or greenish coloration of the dorsum of males in spring. Specimens of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FD8BC45F21E3F9EB" box="[571,734,1597,1622]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" sensu="stricto" species="hispanicus">
|
|
<emphasis id="B92F58777E29FFA3FD8BC45F21E3F9EB" box="[571,734,1597,1622]" italics="true" pageId="7" pageNumber="65">P. hispanicus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A21CE50C7E29FFA3FD58C45F20B9F9EA" box="[744,900,1597,1623]" pageId="7" pageNumber="65" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
|
lacking a split dark vertebral stripe can be distinguished from small, flattened
|
|
<taxonomicName id="4C5BFFE67E29FFA3FDF8C40121FCF9C1" box="[584,705,1635,1660]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="liolepis">
|
|
<emphasis id="B92F58777E29FFA3FDF8C40121FCF9C1" box="[584,705,1635,1660]" italics="true" pageId="7" pageNumber="65">P. liolepis</emphasis>
|
|
</taxonomicName>
|
|
lacking temporal shields by the yellowish coloration on the undertail present in most
|
|
<taxonomicName id="4C5BFFE67E29FFA3FE78C4EA2158F91C" box="[456,613,1672,1697]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" sensu="stricto" species="hispanicus">
|
|
<emphasis id="B92F58777E29FFA3FE78C4EA2158F91C" box="[456,613,1672,1697]" italics="true" pageId="7" pageNumber="65">P. hispanicus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A21CE50C7E29FFA3FDDEC4EA2038F91F" box="[622,773,1672,1698]" pageId="7" pageNumber="65" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
|
, by their different head shape with more flattened head and more pointed snout and their paler iris. Iberian populations of
|
|
<taxonomicName id="4C5BFFE67E29FFA3FC72C4CF2776F97B" box="[962,1099,1709,1734]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="vaucheri">
|
|
<emphasis id="B92F58777E29FFA3FC72C4CF2776F97B" box="[962,1099,1709,1734]" italics="true" pageId="7" pageNumber="65">P. vaucheri</emphasis>
|
|
</taxonomicName>
|
|
(which rarely shows a dark vertebral stripe) often have a yellow or yellowish belly and dark supradorsolateral stripes that are usually either absent or wider than the pale dorsolateral stripes.
|
|
<taxonomicName id="4C5BFFE67E29FFA3FCF4C49A20FEF8AC" box="[836,963,1784,1809]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="bocagei">
|
|
<emphasis id="B92F58777E29FFA3FCF4C49A20FEF8AC" box="[836,963,1784,1809]" italics="true" pageId="7" pageNumber="65">P. bocagei</emphasis>
|
|
</taxonomicName>
|
|
has a blunt, rounder snout and a green dorsum contrasting with brown flanks (males in spring).
|
|
<taxonomicName id="4C5BFFE67E29FFA3FCF8C57F20DCF88B" box="[840,993,1821,1846]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="carbonelli">
|
|
<emphasis id="B92F58777E29FFA3FCF8C57F20DCF88B" box="[840,993,1821,1846]" italics="true" pageId="7" pageNumber="65">P. carbonelli</emphasis>
|
|
</taxonomicName>
|
|
has a deeper head and more pointed snout and green flanks (often contrasting with brown dorsum) in males in spring. These last two taxa also lack a dark vertebral stripe in most specimens.
|
|
</paragraph>
|
|
<paragraph id="8BE484657E29FFA3FF74C5EF2016F871" blockId="7.[151,1437,253,2034]" pageId="7" pageNumber="65">
|
|
Generally, we wish to insist on the difficulty to identify many isolated specimens of the
|
|
<taxonomicName id="4C5BFFE67E29FFA3FA9CC5EF2229F871" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="65" phylum="Chordata" rank="species" species="hispanicus">
|
|
<emphasis id="B92F58777E29FFA3FA9CC5EF2229F871" italics="true" pageId="7" pageNumber="65">Podarcis hispanicus</emphasis>
|
|
</taxonomicName>
|
|
complex on their phenotypic characters only.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C341D7EE7E29FFA3FF76C5B52758F84F" box="[198,1125,2007,2034]" pageId="7" pageNumber="65" type="distribution">
|
|
<paragraph id="8BE484657E29FFA3FF76C5B52758F84F" blockId="7.[151,1437,253,2034]" box="[198,1125,2007,2034]" pageId="7" pageNumber="65">
|
|
<emphasis id="B92F58777E29FFA3FF76C5B52254F84C" bold="true" box="[198,361,2007,2033]" pageId="7" pageNumber="65">Distribution:</emphasis>
|
|
eastern and northern
|
|
<collectingCountry id="F34CC4F57E29FFA3FDD7C5BA2197F84F" box="[615,682,2008,2034]" name="Spain" pageId="7" pageNumber="65">Spain</collectingCountry>
|
|
and parts of southern
|
|
<collectingCountry id="F34CC4F57E29FFA3FC1EC5BA20C3F84F" box="[942,1022,2008,2034]" name="France" pageId="7" pageNumber="65">France</collectingCountry>
|
|
(
|
|
<figureCitation id="136098E07E29FFA3FBBEC5BA276AF84F" box="[1038,1111,2008,2034]" captionStart="FIGURE 4" captionStartId="8.[151,255,1093,1117]" captionText="FIGURE 4. Distribution of Podarcis liolepis." pageId="7" pageNumber="65">Fig. 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |