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<document id="377CCF15497B2A2BF9DAFC23003AC8D0" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFD2FFB9FF70FD17FEB2C53E" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Strobilophila) kwaiensis Skelley, Xu &amp; Tang, new species" docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="53" updateTime="1698439510239" updateUser="ExternalLinkService">
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<mods:title id="B945D85377B393338F3F3CCC23D556BB">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<mods:namePart id="6A1392BCC0ACD59D686243CA569CB2CA">Tang, William</mods:namePart>
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<mods:namePart id="FE9AFFEC59ABFABD726A119D9EB975E1">Lindström, Anders J.</mods:namePart>
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<mods:namePart id="F798B8E05CB57FD9D39BDB98E18B62A1">Khuraijam, Jibankumar Singh</mods:namePart>
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<treatment id="03A38781FFD2FFB9FF70FD17FEB2C53E" ID-DOI="http://doi.org/10.5281/zenodo.5999233" ID-GBIF-Taxon="130809500" ID-Zenodo-Dep="5999233" LSID="urn:lsid:plazi:treatment:03A38781FFD2FFB9FF70FD17FEB2C53E" httpUri="http://treatment.plazi.org/id/03A38781FFD2FFB9FF70FD17FEB2C53E" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
<subSubSection id="C310651CFFD2FFBBFF70FD17FEF1C50F" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="8BB53697FFD2FFBBFF70FD17FC2FC52F" blockId="52.[151,1018,764,823]" box="[151,1018,764,790]" pageId="52" pageNumber="53">
<heading id="D0FD81FBFFD2FFBBFF70FD17FC2FC52F" bold="true" box="[151,1018,764,790]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<emphasis id="B97EEA85FFD2FFBBFF70FD17FC2FC52F" bold="true" box="[151,1018,764,790]" pageId="52" pageNumber="53">
<taxonomicName id="4C0A4D14FFD2FFBBFF70FD17FC88C52F" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[151,861,764,790]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kwaiensis" status="sp. nov." subGenus="Strobilophila">
<emphasis id="B97EEA85FFD2FFBBFF70FD17FEFBC52F" bold="true" box="[151,302,764,790]" italics="true" pageId="52" pageNumber="53">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFD2FFBBFED8FD17FE09C52F" bold="true" box="[319,476,764,790]" italics="true" pageId="52" pageNumber="53">Strobilophila</emphasis>
)
<emphasis id="B97EEA85FFD2FFBBFE0AFD17FDB7C52F" bold="true" box="[493,610,764,790]" italics="true" pageId="52" pageNumber="53">kwaiensis</emphasis>
Skelley, Xu &amp; Tang
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,
<taxonomicNameLabel id="A24D57FEFFD2FFBBFC8DFD17FC2FC52F" box="[874,1018,764,790]" pageId="52" pageNumber="53" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BB53697FFD2FFBBFF70FCF5FEF1C50F" blockId="52.[151,1018,764,823]" box="[151,292,798,823]" pageId="52" pageNumber="53">
<figureCitation id="13312A12FFD2FFBBFF70FCF5FF3AC50E" box="[151,239,798,823]" captionStart="FIGURE 18" captionStartId="53.[151,250,1626,1648]" captionTargetBox="[151,1435,474,1604]" captionTargetId="figure@53.[151,1435,474,1604]" captionTargetPageId="53" captionText="FIGURE 18. Photographs of Cycadophila (Strobilophila) kwaiensis, A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia." httpUri="https://zenodo.org/record/575659/files/figure.png" pageId="52" pageNumber="53">Figs. 18</figureCitation>
AE
</paragraph>
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<subSubSection id="C310651CFFD2FFBBFF70FC8DFD2DC2FB" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="8BB53697FFD2FFBBFF70FC8DFEF6C237" blockId="52.[151,1437,870,2007]" pageId="52" pageNumber="53">
<emphasis id="B97EEA85FFD2FFBBFF70FC8DFE8FC546" bold="true" box="[151,346,870,895]" pageId="52" pageNumber="53">Adult diagnosis.</emphasis>
A member of the
<taxonomicName id="4C0A4D14FFD2FFBBFDD7FC8DFCB9C546" authority="Strobilophila" authorityName="Strobilophila" box="[560,876,870,895]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFD2FFBBFDD7FC8DFD6AC546" box="[560,703,870,895]" italics="true" pageId="52" pageNumber="53">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFD2FFBBFD36FC8DFCB6C546" box="[721,867,870,895]" italics="true" pageId="52" pageNumber="53">Strobilophila</emphasis>
)
</taxonomicName>
<taxonomicName id="4C0A4D14FFD2FFBBFC92FC8DFC08C546" box="[885,989,870,895]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tansachai">tansachai</taxonomicName>
complex distinguished by the pronotal posterior lateral carina lacking an abrupt inward curve at posterior angle, lacking a distinct projection behind the posterior angle pore, a nearly straight basal pronotal margin, and with no dimorphism on the female abdomen. They are also distinguishable by DNA analysis of the 16S rRNA mitochondrial gene (GenBank Accession number
<accessionNumber id="9459AB74FFD2FFBBFF70FC1DFEC0C236" box="[151,277,1014,1039]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY365221" pageId="52" pageNumber="53">KY365221</accessionNumber>
).
</paragraph>
<paragraph id="8BB53697FFD2FFBBFF20FBF1FD2DC2FB" blockId="52.[151,1437,870,2007]" pageId="52" pageNumber="53">
<emphasis id="B97EEA85FFD2FFBBFF20FBF1FE4FC20A" bold="true" box="[199,410,1050,1075]" pageId="52" pageNumber="53">Adult description</emphasis>
. Length
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; width
<quantity id="4CF29B72FFD2FFBBFCE7FBF1FC7DC20B" box="[768,936,1050,1075]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.7" metricValueMin="1.58" pageId="52" pageNumber="53" unit="mm" value="1.64" valueMax="1.7" valueMin="1.58">1.581.70 mm</quantity>
. Body in dorsal view elongate, sides nearly straight, greatest width at middle of elytra; in lateral view convex dorsally. General body and appendage color solid reddish-brown; dorsal surface distinctly punctate, surface appearing slightly dulled, dorsally appearing glabrous, with short procumbent hairs associated with punctation behind eyes and along lateral sides and declivity of elytra, ventrally mostly covered with short procumbent setae.
</paragraph>
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<subSubSection id="C310651CFFD2FFB9FF20FB25FAA3C6CE" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="description">
<paragraph id="8BB53697FFD2FFBBFF20FB25FB8BC0E6" blockId="52.[151,1437,870,2007]" pageId="52" pageNumber="53">
<emphasis id="B97EEA85FFD2FFBBFF20FB25FED1C2DE" bold="true" box="[199,260,1230,1255]" italics="true" pageId="52" pageNumber="53">Head</emphasis>
in dorsal view conical, gradually narrowed apically, surface flat to slightly convex, distinctly punctured; width 0.91.0 mm; dorsal interocular distance
<quantity id="4CF29B72FFD2FFBBFD7FFB19FCEFC333" box="[664,826,1266,1291]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="6.6" metricValueMin="6.0" pageId="52" pageNumber="53" unit="mm" value="0.63" valueMax="0.66" valueMin="0.6">0.600.66 mm</quantity>
, head width/dorsal interocular distance ratio 1.491.52, ventral interocular distance
<quantity id="4CF29B72FFD2FFBBFE34FAFCFDACC317" box="[467,633,1303,1327]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="52" pageNumber="53" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.380.43 mm</quantity>
, head width/ventral interocular distance ratio 2.342.47. Eye globular; bordered dorsally with supraocular stria; projected laterally, with large black facets and short, inconspicuous interfacetal setae. Frons with short, dark, subcuticular suture on each side, approximately 1/4 width of frons in length, extending obliquely to lateral margin above and anterior to antennal insertion. Antennal length about 3/4 pronotal width, and slightly longer than head width, antennomere relative lengths from base to end approximately 21:12:11:10:10:10:10:10:15:15:19; antennomere I (scape) fairly large, slightly elongate, antennomere IIIII slightly longer than IVVIII; club fairly large, IXXI similar in length, XI globular. Clypeus truncate anteriorly, anterior margin with long setae; densely punctate; somewhat emarginate, with narrow margin. Maxillary palpomeres II, III slightly longer than wide; IV elongate oval, apex densely papillate; relative lengths of IIIV approximately 3:3:5. Labium with mentum pentagonal with carina forming triangular plate projecting medially, lacking distinct lateral pockets; labial palpomere III elongate-oval.
<collectingRegion id="49CEF875FFD2FFBBFD6FF994FD30C0AF" box="[648,741,1663,1686]" country="Myanmar" name="Chin" pageId="52" pageNumber="53">Mentum</collectingRegion>
and submentum with moderate punctation and setation, setae short. Gular area smooth, without setae or punctation, except near suture with submentum where a shallow transversely elongate fovea is present bearing dense setose punctures, fovea dark brown.
</paragraph>
<paragraph id="8BB53697FFD2FFBBFF20F901FA48C1EF" blockId="52.[151,1437,870,2007]" pageId="52" pageNumber="53">
<emphasis id="B97EEA85FFD2FFBBFF20F901FEC8C13A" bold="true" box="[199,285,1770,1795]" italics="true" pageId="52" pageNumber="53">Thorax</emphasis>
with pronotum transverse in dorsal view, with marginal beads on all sides; length/width ratio 0.69 0.75, convex, nearly parallel-sided; anterior angles rounded; lateral carina with thickened bead, bead bearing scattered setose punctures; posterior angle exteriorly angulate, not indented at pore; posterior margin projecting medially; with narrow, short longitudinal furrow posterolaterally, beginning at pore on posterior margin, extending forward and ending abruptly, length of furrows relative to pronotal length 0.190.23. Prosternum with anterior margin slightly emarginate, finely denticulate with fringe of long, anteriorly directed setae; posterior prosternal process round and convex. Hypomeron with coarse punctures laterally, medially with longitudinal striations.
</paragraph>
<paragraph id="8BB53697FFD3FFBAFF70FF7CFCA7C79E" blockId="53.[151,1436,151,423]" pageId="53" pageNumber="54">Scutellar shield transverse, posterior margin projected, pentagonal. Elytra elongate, nearly parallel sided, convex; length/width 3.503.67, greatest width near midlength; with marginal bead basally, scutellary striole with 1013 punctures; intervals of striae with distinct punctures. All elytral punctures bearing a single short seta; seta barely emerging out of puncture. Punctation on meso- and metaventrite distinct. Metaventrite long, convex laterally, slightly impressed medially, metathoracic discrimen extending slightly over half metaventrite length. Legs stout, similar in length; femora compressed laterally; tibiae shorter than femora, dilated to wide apex. Protibia triangularly dilated at apex, apex and apical 1/3 of lateral margin with stout spinules, marginal spinules separated by less than length. Meso- and metatibia less triangularly dilated.</paragraph>
<caption id="DF75661FFFD3FFBAFF70F9B1FD74C0AB" httpUri="https://zenodo.org/record/575659/files/figure.png" pageId="53" pageNumber="54" targetBox="[151,1435,474,1604]" targetPageId="53">
<paragraph id="8BB53697FFD3FFBAFF70F9B1FD74C0AB" blockId="53.[151,1436,1626,1682]" pageId="53" pageNumber="54">
<emphasis id="B97EEA85FFD3FFBAFF70F9B1FEF5C049" bold="true" box="[151,288,1626,1648]" pageId="53" pageNumber="54">FIGURE 18.</emphasis>
Photographs of
<taxonomicName id="4C0A4D14FFD3FFBAFE2DF9B1FC87C049" box="[458,850,1626,1648]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="kwaiensis" subGenus="Strobilophila">
<emphasis id="B97EEA85FFD3FFBAFE2DF9B1FD99C049" box="[458,588,1626,1648]" italics="true" pageId="53" pageNumber="54">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFD3FFBAFDBBF9B1FD35C049" box="[604,736,1626,1648]" italics="true" pageId="53" pageNumber="54">Strobilophila</emphasis>
)
<emphasis id="B97EEA85FFD3FFBAFD08F9B1FC87C049" box="[751,850,1626,1648]" italics="true" pageId="53" pageNumber="54">kwaiensis</emphasis>
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, A) dorsal habitus; B) ventral habitus; C) lateral habitus; D) head and prosternum, ventral; E) male genitalia.
</paragraph>
</caption>
<paragraph id="8BB53697FFD3FFBAFF20F92AFD7DC1B0" blockId="53.[151,1436,1729,2036]" pageId="53" pageNumber="54">
<emphasis id="B97EEA85FFD3FFBAFF20F92AFEE0C0E3" bold="true" box="[199,309,1729,1754]" italics="true" pageId="53" pageNumber="54">Abdomen</emphasis>
with 5 ventrites bearing fine punctures; anterior margin with intercoxal process narrow, with triangular point anteromedially, lateral edges slightly projected, lateral and posterior margins arcuate, converging posteriorly; anterior and posterior margins of ventrites more or less straight; ventrite I lacking subcoxal line, slightly longer medially than II; IIV each with pair short hairs located adjacent to midline (often abraded); IIIV subequal in length; V slightly longer than IV with lateral margins converging posteriorly to a rounded apex bearing row of short appressed, densely arranged setae.
</paragraph>
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<emphasis id="B97EEA85FFD3FFBAFF20F87FFEBBC194" bold="true" box="[199,366,1940,1965]" italics="true" pageId="53" pageNumber="54">Male genitalia</emphasis>
(tegmen and median lobe of aedeagus) twisted approximately 45° toward the left side of animal and resting on side. Tegmen sclerotized, triangular, gently twisted; with anterior region ring-like, posterior region sheath-like, posterior margin slightly arcuate; lateral margins gradually converging posteriorly; dorsally with 2 large, elongate, somewhat coniform parameres. Parameres with long setae apically; length/width ratio 2.20. Aedeagus with median lobe short, slender, highly sclerotized, apex long and acuminate. Ratio of median lobe length to penile strut length 1:5.
<emphasis id="B97EEA85FFD0FFB9FDE3FF35FD8EC6CE" bold="true" box="[516,603,222,247]" italics="true" pageId="54" pageNumber="55">Female</emphasis>
similar to male, except with pygidium truncate, not as well developed.
</paragraph>
</subSubSection>
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<paragraph id="8BB53697FFD0FFB9FF20FEEAFC7BC723" blockId="54.[151,1437,151,775]" box="[199,942,257,282]" pageId="54" pageNumber="55">
<materialsCitation id="3B623CCAFFD0FFB9FF20FEEAFC7BC723" ID-GBIF-Occurrence="1502685006" box="[199,942,257,282]" country="Thailand" latitude="14.033334" location="Kanchanaburi Province" longLatPrecision="1289" longitude="99.48333" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Kanchanaburi Province" typeStatus="holotype">
<emphasis id="B97EEA85FFD0FFB9FF20FEEAFEB0C723" bold="true" box="[199,357,257,282]" pageId="54" pageNumber="55">
<typeStatus id="54B18835FFD0FFB9FF20FEEAFED7C723" box="[199,258,257,282]" pageId="54" pageNumber="55">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D7607FFD0FFB9FE8BFEEAFE01C723" box="[364,468,257,282]" name="Thailand" pageId="54" pageNumber="55">Thailand</collectingCountry>
:
<collectingRegion id="49CEF875FFD0FFB9FE39FEEAFD3CC723" box="[478,745,257,282]" country="Thailand" name="Kanchanaburi" pageId="54" pageNumber="55">Kanchanaburi Province</collectingRegion>
,
<geoCoordinate id="EE3E5050FFD0FFB9FD13FEEAFC96C723" box="[756,835,257,282]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="14.033334">14°2N</geoCoordinate>
,
<geoCoordinate id="EE3E5050FFD0FFB9FCA8FEEAFC7FC723" box="[847,938,257,282]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="99.48333">99°29E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFD0FFB9FF20FECFFD96C704" blockId="54.[151,1437,151,775]" box="[199,579,292,317]" pageId="54" pageNumber="55">
<materialsCitation id="3B623CCAFFD0FFB9FF20FECFFD96C704" ID-GBIF-Occurrence="1502685026" box="[199,579,292,317]" country="Thailand" location="Kanchanaburi" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Kanchanaburi">
<emphasis id="B97EEA85FFD0FFB9FF20FECFFEC6C704" bold="true" box="[199,275,292,317]" pageId="54" pageNumber="55">Range</emphasis>
.
<collectingCountry id="F31D7607FFD0FFB9FEC6FECFFE50C704" box="[289,389,292,317]" name="Thailand" pageId="54" pageNumber="55">Thailand</collectingCountry>
(
<collectingRegion id="49CEF875FFD0FFB9FE74FECFFDE2C704" box="[403,567,292,317]" country="Thailand" name="Kanchanaburi" pageId="54" pageNumber="55">Kanchanaburi</collectingRegion>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB53697FFD0FFB9FF20FEACFDEDC7D4" blockId="54.[151,1437,151,775]" pageId="54" pageNumber="55">
<materialsCitation id="3B623CCAFFD0FFB9FF20FEACFBB6C7F3" ID-GBIF-Occurrence="1502685010" collectingDate="2002-07-29" collectionCode="FSCA" collectorName="W. Tang &amp; Deposited" country="Thailand" latitude="14.033334" location="Material" longLatPrecision="1289" longitude="99.48333" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<emphasis id="B97EEA85FFD0FFB9FF20FEACFDF3C759" bold="true" box="[199,550,327,352]" pageId="54" pageNumber="55">
<location id="8ED5604CFFD0FFB9FF20FEACFEFAC759" LSID="urn:lsid:plazi:treatment:03A38781FFD2FFB9FF70FD17FEB2C53E:8ED5604CFFD0FFB9FF20FEACFEFAC759" box="[199,303,327,352]" country="Thailand" latitude="14.033334" longLatPrecision="1289" longitude="99.48333" name="Material" pageId="54" pageNumber="55" stateProvince="Kanchanaburi">Material</location>
examined.
<typeStatus id="54B18835FFD0FFB9FE5EFEACFDF3C759" box="[441,550,327,352]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
(by designation, with damaged left elytron) male with the following labels: 1) [rectangular; white; printed in black ink] “
<collectingCountry id="F31D7607FFD0FFB9FD8AFE80FCD6C7BA" box="[621,771,363,387]" name="Thailand" pageId="54" pageNumber="55">THAILAND</collectingCountry>
,
<collectingRegion id="49CEF875FFD0FFB9FCE9FE80FC7BC7BD" box="[782,942,363,388]" country="Thailand" name="Kanchanaburi" pageId="54" pageNumber="55">Kanchanaburi</collectingRegion>
,
<geoCoordinate id="EE3E5050FFD0FFB9FC5EFE80FBDDC7BD" box="[953,1032,363,388]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="14.033334">14°2N</geoCoordinate>
,
<geoCoordinate id="EE3E5050FFD0FFB9FBF3FE80FBBBC7BD" box="[1044,1134,363,388]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="99.48333">99°29E</geoCoordinate>
, karst tower, ex crown hair of
<taxonomicName id="4C0A4D14FFD0FFB9FF5FFE64FF29C79F" box="[184,252,399,422]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFD0FFB9FF5FFE64FF29C79F" box="[184,252,399,422]" italics="true" pageId="54" pageNumber="55">Cycas</emphasis>
</taxonomicName>
sp.
<taxonomicName id="4C0A4D14FFD0FFB9FED7FE65FE1DC79F" box="[304,456,398,423]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="pectinata">
aff.
<emphasis id="B97EEA85FFD0FFB9FE87FE64FE1DC79F" box="[352,456,399,422]" italics="true" pageId="54" pageNumber="55">pectinata</emphasis>
</taxonomicName>
,
<date id="FFB41057FFD0FFB9FE3EFE64FDBCC79E" box="[473,617,399,423]" pageId="54" pageNumber="55" value="2002-07-29">
<collectingDate id="EFF0E9BFFFD0FFB9FE3EFE64FDBCC79E" box="[473,617,399,423]" pageId="54" pageNumber="55" value="2002-07-29">29-VII-2002</collectingDate>
</date>
,
<collectorName id="26FF5341FFD0FFB9FD9FFE64FD0FC79E" box="[632,730,399,423]" pageId="54" pageNumber="55">W. Tang</collectorName>
”; 2) [rectangular; red; printed in black ink] “
<typeStatus id="54B18835FFD0FFB9FB00FE64FAA8C79E" box="[1255,1405,399,423]" pageId="54" pageNumber="55" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D0CFD1EFFD0FFB9FA60FE64FA49C79F" box="[1415,1436,399,422]" pageId="54" pageNumber="55" type="male"></specimenCount>
<taxonomicName id="4C0A4D14FFD0FFB9FF70FE5AFCE8C7F3" authority="W. Tang, G. Xu &amp; P. Skelley 2015" authorityName="W. Tang, G. Xu &amp; P. Skelley" authorityYear="2015" box="[151,829,433,458]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kwaiensis">
<emphasis id="B97EEA85FFD0FFB9FF70FE5AFE49C7F3" box="[151,412,433,458]" italics="true" pageId="54" pageNumber="55">Cycadophila kwaiensis</emphasis>
, W.
<bibRefCitation id="EF9B4B66FFD0FFB9FE33FE59FCE8C7F3" author="Tang" box="[468,829,433,458]" pageId="54" pageNumber="55" refString="Xu, G., Tang, W., Skelley, P., Liu, N., &amp; Rich, S. (2015) Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa, 3986 (3), 251 - 278." type="journal article" year="2015">Tang, G. Xu &amp; P. Skelley 2015</bibRefCitation>
</taxonomicName>
”.
<collectorName id="26FF5341FFD0FFB9FCB2FE5AFC13C7F3" box="[853,966,433,458]" pageId="54" pageNumber="55">Deposited</collectorName>
in the
<collectionCode id="ED1BAE52FFD0FFB9FBFEFE59FBB6C7F3" box="[1049,1123,434,458]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="54" pageNumber="55">FSCA</collectionCode>
</materialsCitation>
.
<emphasis id="B97EEA85FFD0FFB9FB88FE5AFB01C7F3" bold="true" box="[1135,1236,433,458]" pageId="54" pageNumber="55">Allotype</emphasis>
and 4
<emphasis id="B97EEA85FFD0FFB9FAC1FE5AFA49C7F3" bold="true" box="[1318,1436,433,458]" pageId="54" pageNumber="55">paratypes</emphasis>
(same label data) deposited at FSCA.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFD0FFB9FF20FE1CFEA1C40D" pageId="54" pageNumber="55" type="etymology">
<paragraph id="8BB53697FFD0FFB9FF20FE1CFEA1C40D" blockId="54.[151,1437,151,775]" pageId="54" pageNumber="55">
<emphasis id="B97EEA85FFD0FFB9FF20FE1CFE9BC429" bold="true" box="[199,334,503,528]" pageId="54" pageNumber="55">Etymology.</emphasis>
Named for the locality of collection in the vicinity of the Kwai River Bridge in
<collectingRegion id="49CEF875FFD0FFB9FB18FE1CFF2BC40D" country="Thailand" name="Kanchanaburi" pageId="54" pageNumber="55">Kanchanaburi Province</collectingRegion>
,
<collectingCountry id="F31D7607FFD0FFB9FEEFFDF0FEBAC40D" box="[264,367,539,564]" name="Thailand" pageId="54" pageNumber="55">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFD0FFB9FF20FDD5FEB2C53E" pageId="54" pageNumber="55" type="discussion">
<paragraph id="8BB53697FFD0FFB9FF20FDD5FEB2C53E" blockId="54.[151,1437,151,775]" pageId="54" pageNumber="55">
<materialsCitation id="3B623CCAFFD0FFB9FF20FDD5FBD0C440" ID-GBIF-Occurrence="1502685016" elevation="100" location="Kanchanaburi Province" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Kanchanaburi Province">
<emphasis id="B97EEA85FFD0FFB9FF20FDD5FEE1C46E" bold="true" box="[199,308,574,599]" pageId="54" pageNumber="55">Remarks</emphasis>
. This species of
<taxonomicName id="4C0A4D14FFD0FFB9FE09FDD5FDA8C46E" box="[494,637,574,599]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFD0FFB9FE09FDD5FDA8C46E" box="[494,637,574,599]" italics="true" pageId="54" pageNumber="55">Cycadophila</emphasis>
</taxonomicName>
appears to be restricted to a currently undescribed species of
<taxonomicName id="4C0A4D14FFD0FFB9FAC0FDD4FAB9C46F" box="[1319,1388,575,598]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFD0FFB9FAC0FDD4FAB9C46F" box="[1319,1388,575,598]" italics="true" pageId="54" pageNumber="55">Cycas</emphasis>
</taxonomicName>
that inhabits karst towers in
<collectingRegion id="49CEF875FFD0FFB9FE53FD8AFD15C443" box="[436,704,609,634]" country="Thailand" name="Kanchanaburi" pageId="54" pageNumber="55">Kanchanaburi Province</collectingRegion>
at elevations below
<quantity id="4CF29B72FFD0FFB9FC50FD89FBD0C440" box="[951,1029,610,634]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="100.0">
<specimenCount id="9D0CFD1EFFD0FFB9FC50FD89FBD0C440" box="[951,1029,610,634]" pageId="54" pageNumber="55" type="generic">
<elevation id="0027D1A4FFD0FFB9FC50FD89FBD0C440" box="[951,1029,610,634]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="100.0">100 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
.
<materialsCitation id="3B623CCAFFD0FFB9FBC7FD8AFEB2C53E" ID-GBIF-Occurrence="1502685029" collectionCode="DNA" country="Vietnam" elevation="600" location="This" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Kanchanaburi Province">
Although
<taxonomicName id="4C0A4D14FFD0FFB9FB73FD8AFA49C443" box="[1172,1436,609,634]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="tansachai">
<emphasis id="B97EEA85FFD0FFB9FB73FD8AFA49C443" box="[1172,1436,609,634]" italics="true" pageId="54" pageNumber="55">Cycadophila tansachai</emphasis>
</taxonomicName>
occurs
<collectedFrom id="3816756AFFD0FFB9FF00FD6EFE6AC4A4" box="[231,447,645,669]" pageId="54" pageNumber="55">
on
<taxonomicName id="4C0A4D14FFD0FFB9FEECFD6EFE6AC4A4" box="[267,447,645,669]" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis id="B97EEA85FFD0FFB9FEECFD6EFE6AC4A4" box="[267,447,645,669]" italics="true" pageId="54" pageNumber="55">Cycas pectinata</emphasis>
</taxonomicName>
</collectedFrom>
some
<quantity id="4CF29B72FFD0FFB9FDEEFD6EFD8AC4A4" box="[521,607,644,669]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="km" value="100.0">100 km</quantity>
north at elevations above
<quantity id="4CF29B72FFD0FFB9FC6FFD6FFC1AC4A5" box="[904,975,644,669]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="54" pageNumber="55" unit="m" value="600.0">
<elevation id="0027D1A4FFD0FFB9FC6FFD6FFC1AC4A5" box="[904,975,644,669]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="54" pageNumber="55" unit="m" value="600.0">600 m</elevation>
</quantity>
in mountains in
<collectingRegion id="49CEF875FFD0FFB9FB6AFD6FFA4CC4A4" box="[1165,1433,644,669]" country="Thailand" name="Kanchanaburi" pageId="54" pageNumber="55">Kanchanaburi Province</collectingRegion>
,
<collectionCode id="ED1BAE52FFD0FFB9FF70FD43FF01C4F9" box="[151,212,680,704]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="54" pageNumber="55">DNA</collectionCode>
analysis indicates that
<taxonomicName id="4C0A4D14FFD0FFB9FE3AFD4CFD37C4F9" box="[477,738,679,704]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kwaiensis">
<emphasis id="B97EEA85FFD0FFB9FE3AFD4CFD37C4F9" box="[477,738,679,704]" italics="true" pageId="54" pageNumber="55">Cycadophila kwaiensis</emphasis>
</taxonomicName>
is also closely related to
<taxonomicName id="4C0A4D14FFD0FFB9FBE2FD4CFB00C4F9" box="[1029,1237,679,704]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="hiepi">
<emphasis id="B97EEA85FFD0FFB9FBE2FD4CFB00C4F9" box="[1029,1237,679,704]" italics="true" pageId="54" pageNumber="55">Cycadophila hiepi</emphasis>
</taxonomicName>
in
<collectingCountry id="F31D7607FFD0FFB9FB1BFD4CFA8BC4F9" box="[1276,1374,679,704]" name="Vietnam" pageId="54" pageNumber="55">Vietnam</collectingCountry>
, over
<quantity id="4CF29B72FFD0FFB9FF70FD20FF2FC4DD" box="[151,250,715,740]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="km" value="1000.0">1000 km</quantity>
to the east.
<location id="8ED5604CFFD0FFB9FE64FD20FE60C4DD" LSID="urn:lsid:plazi:treatment:03A38781FFD2FFB9FF70FD17FEB2C53E:8ED5604CFFD0FFB9FE64FD20FE60C4DD" box="[387,437,715,740]" country="Vietnam" name="This" pageId="54" pageNumber="55" stateProvince="Kanchanaburi Province">This</location>
suggests that
<taxonomicName id="4C0A4D14FFD0FFB9FDB3FD27FD37C4DD" box="[596,738,715,740]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="kwaiensis">
<emphasis id="B97EEA85FFD0FFB9FDB3FD27FD37C4DD" box="[596,738,715,740]" italics="true" pageId="54" pageNumber="55">C. kwaiensis</emphasis>
</taxonomicName>
and its host cycad are relicts with formerly wider distribution in lowland forests.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>