185 lines
21 KiB
XML
185 lines
21 KiB
XML
<document id="0E3DAA4B01D1269E80DCA2954B3B3AA3" ID-CLB-Dataset="26372" ID-DOI="10.11646/zootaxa.4623.3.6" ID-GBIF-Dataset="89686326-030c-4d79-8ccb-8c2db484c121" ID-ISSN="1175-5326" ID-Zenodo-Dep="3258640" ID-ZooBank="F85DD0E5-0452-4EAB-843D-BC32F12F0A87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561649240053" checkinUser="plazi" docAuthor="Murcia, Andres David, Cadena-Castañeda, Oscar J., Noriega, Jorge Ari & García, Alexander García" docDate="2019" docId="03D787A40D0BFFD8FF7F680EFE31FF43" docLanguage="en" docName="zootaxa.4623.3.6.pdf" docOrigin="Zootaxa 4623 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyphloea Redtenbacher 1906" docType="treatment" docVersion="8" lastPageNumber="549" masterDocId="FFEEFFDC0D0AFFDCFFE86E53FFE2FFB8" masterDocTitle="New species of Pachyphloea Redtenbacher, 1906 (Phasmida: Pseudophasmatidae Xerosomatinae) with comments on Grylloclonia Zompro, 2004 n. syn." masterLastPageNumber="554" masterPageNumber="545" pageNumber="546" updateTime="1698737462397" updateUser="ExternalLinkService">
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<mods:title id="458F259D25A7909BD36D0F76FBE57E90">New species of Pachyphloea Redtenbacher, 1906 (Phasmida: Pseudophasmatidae Xerosomatinae) with comments on Grylloclonia Zompro, 2004 n. syn.</mods:title>
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<mods:namePart id="F1C6A435D306F7FE8E19A7F150EBA3F8">Murcia, Andres David</mods:namePart>
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<mods:namePart id="5015A31FB66DFACC0C137015244FF048">Cadena-Castañeda, Oscar J.</mods:namePart>
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<mods:namePart id="C832FD33916823F54B176851E8997FFE">Noriega, Jorge Ari</mods:namePart>
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<mods:namePart id="1C438DB4058AF3C9EFF77839E9702436">García, Alexander García</mods:namePart>
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<mods:title id="D5C5CBF66464096E5824E246E1357140">Zootaxa</mods:title>
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<mods:date id="BD2A70402515FA4BC03505F0F6C5EE40">2019</mods:date>
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<mods:number id="6F76E0010487ED493837C09963A07E2F">2019-06-26</mods:number>
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<mods:number id="40B72DB3B6364928D6A079CBB8C12878">4623</mods:number>
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<treatment id="03D787A40D0BFFD8FF7F680EFE31FF43" ID-DOI="http://doi.org/10.5281/zenodo.5921610" ID-GBIF-Taxon="158407713" ID-Zenodo-Dep="5921610" LSID="urn:lsid:plazi:treatment:03D787A40D0BFFD8FF7F680EFE31FF43" httpUri="http://treatment.plazi.org/id/03D787A40D0BFFD8FF7F680EFE31FF43" lastPageId="4" lastPageNumber="549" pageId="1" pageNumber="546">
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<subSubSection id="C36465390D0BFFDDFF7F680EFD6BF9CF" box="[151,649,1629,1656]" pageId="1" pageNumber="546" type="nomenclature">
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<paragraph id="8BC136B20D0BFFDDFF7F680EFD6BF9CF" blockId="1.[151,649,1629,1656]" box="[151,649,1629,1656]" pageId="1" pageNumber="546">
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<heading id="D08981DE0D0BFFDDFF7F680EFD6BF9CF" bold="true" box="[151,649,1629,1656]" fontSize="11" level="1" pageId="1" pageNumber="546" reason="1">
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<emphasis id="B90AEAA00D0BFFDDFF7F680EFD6BF9CF" bold="true" box="[151,649,1629,1656]" pageId="1" pageNumber="546">
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Genus
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<taxonomicName id="4C7E4D310D0BFFDDFF06680EFD6BF9CF" ID-CoL="8NWKN" authority="Redtenbacher, 1906" authorityName="Redtenbacher" authorityYear="1906" box="[238,649,1629,1656]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="1" pageNumber="546" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA00D0BFFDDFF06680EFE68F9C0" bold="true" box="[238,394,1629,1656]" italics="true" pageId="1" pageNumber="546">Pachyphloea</emphasis>
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<bibRefCitation id="EFEF4B430D0BFFDDFE7A680EFD6BF9CF" author="Redtenbacher, J." box="[402,649,1629,1655]" pageId="1" pageNumber="546" pagination="1 - 180" refId="ref4772" refString="Redtenbacher, J. (1906) Phasmidae Areolatae. Die Insektenfamilie der Phasmiden, 1, 1 - 180." type="journal article" year="1906">Redtenbacher, 1906</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36465390D0BFFDDFF7F68F7FE73F867" pageId="1" pageNumber="546" type="diagnosis">
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<paragraph id="8BC136B20D0BFFDDFF7F68F7FE73F867" blockId="1.[151,1437,1700,2015]" pageId="1" pageNumber="546">
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<emphasis id="B90AEAA00D0BFFDDFF7F68F7FEF6F907" bold="true" box="[151,276,1700,1727]" pageId="1" pageNumber="546">Diagnosis:</emphasis>
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Body in brown tones, robust and granular, with moderate foliaceous extensions in the legs and some species in the abdomen, mainly in tergites V to VII, more conspicuous in females than in males. Distal half of the antennae white or light brown, posterior portion of the vertex with spinose prolongations, becoming tubercles or foliate in shape, scapus and pedicellus unarmed. In dorsal view with a median line running from the anterior margin of the pronotum to the last abdominal segment, pronotum and metanotum wider than long; mesonotum longer than pronotum; metanotum longer than wide; the thorax may be armed with mid-sized spines, foliose extensions, or without any of these. Cerci in both sexes conical and very small, partially covered by tergum X in dorsal view. Subgenital plate with posterior margin rounded or triangular in females; males present variable poculum that covers the vomer completely.
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</paragraph>
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</subSubSection>
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<caption id="DF01663A0D08FFDEFF7F6887FAF4F8A9" httpUri="https://zenodo.org/record/3258642/files/figure.png" pageId="2" pageNumber="547" startId="2.[151,250,1748,1773]" targetBox="[151,1436,226,1717]" targetPageId="2">
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<paragraph id="8BC136B20D08FFDEFF7F6887FAF4F8A9" blockId="2.[151,1436,1748,1809]" pageId="2" pageNumber="547">
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<emphasis id="B90AEAA00D08FFDEFF7F6887FEF1F955" bold="true" box="[151,275,1748,1773]" pageId="2" pageNumber="547">FIGURE 1.</emphasis>
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<taxonomicName id="4C7E4D310D08FFDEFEF16887FDEDF954" authority="Murcia & Cadena-Castañeda & Noriega & García, 2019" authorityName="Murcia & Cadena-Castañeda & Noriega & García" authorityYear="2019" box="[281,527,1748,1772]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="547" phylum="Arthropoda" rank="species" species="magnoliae" status="sp. nov.">
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<emphasis id="B90AEAA00D08FFDEFEF16887FDEDF954" box="[281,527,1748,1772]" italics="true" pageId="2" pageNumber="547">Pachyphloea magnoliae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23957DB0D08FFDEFDFD6886FDACF955" box="[533,590,1749,1773]" pageId="2" pageNumber="547" rank="species">
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<emphasis id="B90AEAA00D08FFDEFDFD6886FDACF955" bold="true" box="[533,590,1749,1773]" pageId="2" pageNumber="547">n. sp.</emphasis>
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</taxonomicNameLabel>
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(Male)
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<emphasis id="B90AEAA00D08FFDEFD486887FD34F955" bold="true" box="[672,726,1748,1773]" pageId="2" pageNumber="547">A–B.</emphasis>
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Habitus in lateral and dorsal views, respectively.
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<emphasis id="B90AEAA00D08FFDEFB256887FAE7F955" bold="true" box="[1229,1285,1748,1773]" pageId="2" pageNumber="547">C–D.</emphasis>
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Head and pronotum in lateral and dorsal views, respectively.
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<emphasis id="B90AEAA00D08FFDEFD9268ABFD50F8A9" bold="true" box="[634,690,1784,1809]" pageId="2" pageNumber="547">E–G.</emphasis>
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Terminalia in lateral, dorsal and ventral views, respectively.
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</paragraph>
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</caption>
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<subSubSection id="C36465390D08FFDEFF2F696CFBA8F8E1" box="[199,1098,1854,1881]" pageId="2" pageNumber="547" type="materials_examined">
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<paragraph id="8BC136B20D08FFDEFF2F696CFBA8F8E1" blockId="2.[151,1437,1854,2025]" box="[199,1098,1854,1881]" pageId="2" pageNumber="547">
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<emphasis id="B90AEAA00D08FFDEFF2F696CFE86F8E1" bold="true" box="[199,356,1854,1881]" pageId="2" pageNumber="547">
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<typeStatus id="54C588100D08FFDEFF2F696CFEE0F8E1" box="[199,258,1855,1881]" pageId="2" pageNumber="547">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C7E4D310D08FFDEFE84696CFCB2F8E1" authority="Redtenbacher, 1906" authorityName="Redtenbacher" authorityYear="1906" box="[364,848,1855,1881]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="547" phylum="Arthropoda" rank="species" species="aberrans">
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<emphasis id="B90AEAA00D08FFDEFE84696CFD88F8E1" box="[364,618,1855,1881]" italics="true" pageId="2" pageNumber="547">Pachyphloea aberrans</emphasis>
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<bibRefCitation id="EFEF4B430D08FFDEFD99696CFCB2F8E1" author="Redtenbacher, J." box="[625,848,1855,1881]" pageId="2" pageNumber="547" pagination="1 - 180" refId="ref4772" refString="Redtenbacher, J. (1906) Phasmidae Areolatae. Die Insektenfamilie der Phasmiden, 1, 1 - 180." type="journal article" year="1906">Redtenbacher, 1906</bibRefCitation>
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</taxonomicName>
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by original monotypy
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</paragraph>
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</subSubSection>
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<subSubSection id="C36465390D08FFD8FF2F6931FE31FF43" lastPageId="4" lastPageNumber="549" pageId="2" pageNumber="547" type="discussion">
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<paragraph id="8BC136B20D08FFDFFF2F6931FBB6FE33" blockId="2.[151,1437,1854,2025]" lastBlockId="3.[151,1437,153,395]" lastPageId="3" lastPageNumber="548" pageId="2" pageNumber="547">
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<emphasis id="B90AEAA00D08FFDEFF2F6931FEB0F8C5" bold="true" box="[199,338,1890,1917]" pageId="2" pageNumber="547">Comments:</emphasis>
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In his taxonomic key,
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<bibRefCitation id="EFEF4B430D08FFDEFDBD6930FCE0F8C5" author="Zompro, O." box="[597,770,1891,1917]" pageId="2" pageNumber="547" refId="ref4855" refString="Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea), Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Hamburg, 327 pp." type="book" year="2004">Zompro (2004)</bibRefCitation>
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differentiated
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<taxonomicName id="4C7E4D310D08FFDEFC406930FBD8F8C5" authorityName="Redtenbacher" authorityYear="1906" box="[936,1082,1891,1917]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="547" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA00D08FFDEFC406930FBD8F8C5" box="[936,1082,1891,1917]" italics="true" pageId="2" pageNumber="547">Pachyphloea</emphasis>
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</taxonomicName>
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and the
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<emphasis id="B90AEAA00D08FFDEFB756930FAD7F8C5" box="[1181,1333,1891,1917]" italics="true" pageId="2" pageNumber="547">
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<taxonomicName id="4C7E4D310D08FFDEFB756930FACEF8C5" authorityName="Zompro" authorityYear="2004" box="[1181,1324,1891,1917]" class="Insecta" family="Pseudophasmatidae" genus="Grylloclonia" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="547" phylum="Arthropoda" rank="genus">Grylloclonia</taxonomicName>
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-
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</emphasis>
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<taxonomicName id="4C7E4D310D08FFDEFADE6937FE60F819" authority="Zompro, 2004" authorityName="Zompro" authorityYear="2004" class="Insecta" family="Pseudophasmatidae" genus="Dicranoclonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="547" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA00D08FFDEFADE6937FF3FF819" italics="true" pageId="2" pageNumber="547">Dicranoclonia</emphasis>
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<bibRefCitation id="EFEF4B430D08FFDEFF0B69D4FE60F819" author="Zompro, O." box="[227,386,1927,1953]" pageId="2" pageNumber="547" refId="ref4855" refString="Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea), Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Hamburg, 327 pp." type="book" year="2004">Zompro, 2004</bibRefCitation>
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</taxonomicName>
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complex (synonymized under
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<taxonomicName id="4C7E4D310D08FFDEFD3169D4FC11F819" authority="Stal, 1875" authorityName="Stal" authorityYear="1875" box="[729,1011,1926,1953]" class="Insecta" family="Pseudophasmatidae" genus="Acanthoclonia" kingdom="Animalia" order="Phasmida" pageId="2" pageNumber="547" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA00D08FFDEFD3169D4FC9EF819" box="[729,892,1927,1953]" italics="true" pageId="2" pageNumber="547">Acanthoclonia</emphasis>
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Stål, 1875
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</taxonomicName>
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by
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<bibRefCitation id="EFEF4B430D08FFDEFBF369D4FB03F819" author="Conle, O. & Hennemann, F. & Gutierrez, Y." box="[1051,1249,1927,1953]" pageId="2" pageNumber="547" refId="ref4577" refString="Conle, O., Hennemann, F. & Gutierrez, Y. (2011) The Stick Insects of Colombia. Books on Demand GmbH, Norderstedt, 406 pp." type="book" year="2011">
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Conle
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<emphasis id="B90AEAA00D08FFDEFB8C69DBFB76F819" box="[1124,1172,1927,1953]" italics="true" pageId="2" pageNumber="547">et al</emphasis>
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., 2011
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</bibRefCitation>
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) based on the arrangement of lateral margins of the mesonotum and the presence/absence of a sensorial area on the prosternum. The arrangement of the lateral margins of the mesonotum however is a variable character even within species, normally with females presenting relatively broader margins of mesonotum than males. The larger size and increased robustness of these in females leads to a gradual widening of the mesothorax towards the posterior in which the posterior margin is wider than the anterior. This occurs for species of both genera, without any other notable difference between them for this character. Likewise, the sensorial area, or transverse ridge on the anterior half of the prosternun, is visible in both
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<taxonomicName id="4C7E4D310D09FFDFFEB36F7AFE0FFEFB" authorityName="Redtenbacher" authorityYear="1906" box="[347,493,297,323]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="548" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA00D09FFDFFEB36F7AFE0FFEFB" box="[347,493,297,323]" italics="true" pageId="3" pageNumber="548">Pachyphloea</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7E4D310D09FFDFFDCD6F7AFD56FEFB" authorityName="Zompro" authorityYear="2004" box="[549,692,297,323]" class="Insecta" family="Pseudophasmatidae" genus="Grylloclonia" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="548" phylum="Arthropoda" rank="genus">
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<emphasis id="B90AEAA00D09FFDFFDCD6F7AFD56FEFB" box="[549,692,297,323]" italics="true" pageId="3" pageNumber="548">Grylloclonia</emphasis>
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</taxonomicName>
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. Additional traits that were used by
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<bibRefCitation id="EFEF4B430D09FFDFFBA76F7AFB1EFEFB" author="Zompro, O." box="[1103,1276,297,323]" pageId="3" pageNumber="548" refId="ref4855" refString="Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea), Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Hamburg, 327 pp." type="book" year="2004">Zompro (2004)</bibRefCitation>
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to distinguish between these genera, such as the degree of development of spines and foliaceous extensions or terminalia structure, have been assessed to still serve for species recognition, albeit not to separate genera.
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</paragraph>
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<caption id="DF01663A0D09FFDFFF7F69FDFAD5F853" httpUri="https://zenodo.org/record/3258644/files/figure.png" pageId="3" pageNumber="548" startId="3.[151,250,1966,1991]" targetBox="[217,1369,465,1947]" targetPageId="3">
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<paragraph id="8BC136B20D09FFDFFF7F69FDFAD5F853" blockId="3.[151,1437,1966,2027]" pageId="3" pageNumber="548">
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<emphasis id="B90AEAA00D09FFDFFF7F69FDFEF6F87F" bold="true" box="[151,276,1966,1991]" pageId="3" pageNumber="548">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7E4D310D09FFDFFEF469FCFDF6F87F" authority="Murcia & Cadena-Castañeda & Noriega & García, 2019" authorityName="Murcia & Cadena-Castañeda & Noriega & García" authorityYear="2019" box="[284,532,1967,1991]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="magnoliae" status="sp. nov.">
|
||
<emphasis id="B90AEAA00D09FFDFFEF469FCFDF6F87F" box="[284,532,1967,1991]" italics="true" pageId="3" pageNumber="548">Pachyphloea magnoliae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957DB0D09FFDFFDF369FCFDB4F87F" box="[539,598,1967,1991]" pageId="3" pageNumber="548" rank="species">
|
||
<emphasis id="B90AEAA00D09FFDFFDF369FCFDB4F87F" bold="true" box="[539,598,1967,1991]" pageId="3" pageNumber="548">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Female)
|
||
<emphasis id="B90AEAA00D09FFDFFD2969FDFD1AF87F" bold="true" box="[705,760,1966,1991]" pageId="3" pageNumber="548">A–B.</emphasis>
|
||
Habitus in lateral and dorsal views, respectively.
|
||
<emphasis id="B90AEAA00D09FFDFFB1369FDFAD1F87F" bold="true" box="[1275,1331,1966,1991]" pageId="3" pageNumber="548">C–D.</emphasis>
|
||
Head and pronotum in lateral and dorsal views, respectively.
|
||
<emphasis id="B90AEAA00D09FFDFFD746981FD36F853" bold="true" box="[668,724,2002,2027]" pageId="3" pageNumber="548">E–G.</emphasis>
|
||
Terminalia in lateral, dorsal and ventral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136B20D0EFFD8FF2F6ECAFE31FF43" blockId="4.[151,1437,153,251]" pageId="4" pageNumber="549">
|
||
In view of the aforementioned characteristics, and taking intrageneric variability into account, we propose
|
||
<taxonomicName id="4C7E4D310D0EFFD8FF7F6EEEFE2FFF6F" authority="Zompro, 2004" authorityName="Zompro" authorityYear="2004" box="[151,461,189,215]" class="Insecta" family="Pseudophasmatidae" genus="Grylloclonia" kingdom="Animalia" order="Phasmida" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA00D0EFFD8FF7F6EEEFEC4FF6F" box="[151,294,189,215]" italics="true" pageId="4" pageNumber="549">Grylloclonia</emphasis>
|
||
<bibRefCitation id="EFEF4B430D0EFFD8FEC56EEEFE2FFF6F" author="Zompro, O." box="[301,461,189,215]" pageId="4" pageNumber="549" refId="ref4855" refString="Zompro, O. (2004) Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea), Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, Hamburg, 327 pp." type="book" year="2004">Zompro, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B90AEAA00D0EFFD8FE3C6EEEFDC2FF6F" bold="true" box="[468,544,189,215]" pageId="4" pageNumber="549">n. syn.</emphasis>
|
||
as synonym of
|
||
<taxonomicName id="4C7E4D310D0EFFD8FD386EEEFC80FF6F" authorityName="Redtenbacher" authorityYear="1906" box="[720,866,189,215]" class="Insecta" family="Pseudophasmatidae" genus="Pachyphloea" kingdom="Animalia" order="Phasmida" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90AEAA00D0EFFD8FD386EEEFC80FF6F" box="[720,866,189,215]" italics="true" pageId="4" pageNumber="549">Pachyphloea</emphasis>
|
||
</taxonomicName>
|
||
since there are not enough diagnostic characters to differentiate the two genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |