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<document id="A9B3C9DA53AC2A54FC948BA318E35861" ID-CLB-Dataset="20369" ID-DOI="10.11646/zootaxa.5104.4.9" ID-GBIF-Dataset="b507936a-58a7-4ae6-a21d-80ac84c985df" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332233" ID-ZooBank="D8D1FF6D-8B84-4D88-8EC7-8103DBFAF7F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646595789018" checkinUser="plazi" docAuthor="Kazandzhieva, Svetlozara &amp; Bechev, Dimitar" docDate="2022" docId="533B090A7718FFB1FF4ECBBEA1D8C383" docLanguage="en" docName="zootaxa.5104.4.9.pdf" docOrigin="Zootaxa 5104 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chetoneura Colless 1962" docType="key" docVersion="9" lastPageNumber="598" masterDocId="AF027172771CFFB4FFD9CD1FA444C36B" masterDocTitle="A new species of Chetoneura Colless from Borneo (Diptera: Keroplatidae)" masterLastPageNumber="599" masterPageNumber="593" pageNumber="597" updateTime="1699316992557" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DA518AD85E8125D6228956496E497517">A new species of Chetoneura Colless from Borneo (Diptera: Keroplatidae)</mods:title>
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<mods:namePart id="08727DA4B2504281B6B51B9C8FC8F7D1">Kazandzhieva, Svetlozara</mods:namePart>
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<mods:affiliation id="B99CB872B183413F0BFD816DCCD2E614">Department of Zoology, University of Plovdiv, 24 Tzar Assen Str., BG 4000 Plovdiv, Bulgaria &amp; Regional Museum of Natural History - Plovdiv, 34 Hristo G. Danov Str., BG 4000 Plovdiv, Bulgaria &amp; svetlozara. k @ gmail. com; https: // orcid. org / 0000 - 0001 - 5018 - 8951</mods:affiliation>
<mods:nameIdentifier id="A4093350415D160B835E06F9DDD54CF4" type="email">svetlozara.k@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FCADF016ACD993C4437C52C6E227FF62">Bechev, Dimitar</mods:namePart>
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<mods:affiliation id="FD458ECFCC3C7EC39CE857FC52EDCD94">Department of Zoology, University of Plovdiv, 24 Tzar Assen Str., BG 4000 Plovdiv, Bulgaria &amp; dbechev @ abv. bg; https: // orcid. org / 0000 - 0001 - 5697 - 1197</mods:affiliation>
<mods:nameIdentifier id="E619E8CED0487449B546B2D5713B520A" type="email">dbechev@abv.bg</mods:nameIdentifier>
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<mods:title id="BE99F6BD5703B0D93955B49374461C7C">Zootaxa</mods:title>
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<treatment id="533B090A7718FFB1FF4ECBBEA1D8C383" ID-DOI="http://doi.org/10.5281/zenodo.6333344" ID-GBIF-Taxon="193878520" ID-Zenodo-Dep="6333344" LSID="urn:lsid:plazi:treatment:533B090A7718FFB1FF4ECBBEA1D8C383" httpUri="http://treatment.plazi.org/id/533B090A7718FFB1FF4ECBBEA1D8C383" lastPageId="5" lastPageNumber="598" pageId="4" pageNumber="597">
<subSubSection id="9388EB977718FFB0FF4ECBBEA668C5D7" box="[151,556,1697,1724]" pageId="4" pageNumber="597" type="nomenclature">
<paragraph id="DB2DB81C7718FFB0FF4ECBBEA668C5D7" blockId="4.[151,556,1697,1724]" box="[151,556,1697,1724]" pageId="4" pageNumber="597">
<heading id="80650F707718FFB0FF4ECBBEA668C5D7" bold="true" box="[151,556,1697,1724]" fontSize="11" level="1" pageId="4" pageNumber="597" reason="1">
<emphasis id="E9E6640E7718FFB0FF4ECBBEA668C5D7" bold="true" box="[151,556,1697,1724]" pageId="4" pageNumber="597">
Key to the species of
<taxonomicName id="1C92C39F7718FFB0FE44CBBEA668C5D7" ID-CoL="62LTP" ID-ENA="481811" authorityName="Colless" authorityYear="1962" box="[413,556,1697,1724]" class="Insecta" family="Keroplatidae" genus="Chetoneura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="E9E6640E7718FFB0FE44CBBEA668C5D7" bold="true" box="[413,556,1697,1724]" italics="true" pageId="4" pageNumber="597">Chetoneura</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9388EB977718FFB1FF4ECBF9A1D8C383" lastPageId="5" lastPageNumber="598" pageId="4" pageNumber="597" type="key">
<keyStep id="6066A0B97718FFB0FF4ECBF9A1D8C472" pageId="4" pageNumber="597">
<paragraph id="DB2DB81C7718FFB0FF4ECBF9A1D8C590" blockId="4.[151,1437,1766,2023]" box="[151,1436,1766,1788]" pageId="4" pageNumber="597">
<keyLead id="60631B297718FFB0FF4ECBF9A1D8C590" box="[151,1436,1766,1788]" pageId="4" pageNumber="597">
1. R-M fusion very short (
<figureCitation id="43A9A4997718FFB0FE1CCBF9A644C597" box="[453,512,1766,1788]" captionStart="FIGURE 2" captionStartId="2.[151,250,795,819]" captionTargetId="figure-29@2.[296,1292,181,771]" captionTargetPageId="2" captionText="FIGURE 2. Chetoneura lagangеnsis sp. n., wing, view from below." figureDoi="http://doi.org/10.5281/zenodo.6335586" httpUri="https://zenodo.org/record/6335586/files/figure.png" pageId="4" pageNumber="597">Fig. 2</figureCitation>
). Tergite IX slightly longer than broad or up to about 1.7 as long as wide, not subtriangular... 2
</keyLead>
</paragraph>
<paragraph id="DB2DB81C7718FFB0FF4ECA1BA1D8C472" blockId="4.[151,1437,1766,2023]" box="[151,1436,1795,1817]" pageId="4" pageNumber="597">
<keyLead id="60631B297718FFB0FF4ECA1BA1D8C472" box="[151,1436,1795,1817]" pageId="4" pageNumber="597">- R-M fusion as long as r-m or more. Tergite IX shorter than broad, subtriangular.................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="6066A0B97718FFB0FF4ECA3EA1D8C4E5" pageId="4" pageNumber="597">
<paragraph id="DB2DB81C7718FFB0FF4ECA3EA1D8C43F" blockId="4.[151,1437,1766,2023]" pageId="4" pageNumber="597">
<keyLead id="60631B297718FFB0FF4ECA3EA1D8C43F" pageId="4" pageNumber="597">
2. Scutum yellowish, with lateral margins and two submedian longitudinal stripes dark. Tergite IX slightly longer than broad, length/width ratio about 1/1. Form of tergite IX—
<figureCitation id="43A9A4997718FFB0FD1DCA21A6BDC43F" box="[708,761,1854,1876]" captionStart="FIGURES 79" captionStartId="4.[151,264,595,619]" captionTargetBox="[307,1287,181,571]" captionTargetId="figure-29@4.[300,1288,181,572]" captionTargetPageId="4" captionText="FIGURES 79. Form of tergite IX (not in scale): 7—Ch. davidi (by Ševčík et al. 2021: Fig. 13C); 8—Ch. cavernae (by Colless 1962: Fig. 1c); 9—Ch. lagangеnsis sp. n." figureDoi="http://doi.org/10.5281/zenodo.6332237" httpUri="https://zenodo.org/record/6332237/files/figure.png" pageId="4" pageNumber="597">Fig. 7</figureCitation>
........................
<taxonomicName id="1C92C39F7718FFB0FBFFCA21A1D8C43F" ID-CoL="9HYP4" authority="Sevcik, Hippa &amp; Burdikova" authorityName="Sevcik, Hippa &amp; Burdikova" box="[1062,1436,1854,1878]" class="Insecta" family="Keroplatidae" genus="Chetoneura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="E9E6640E7718FFB0FBFFCA21A0CEC438" box="[1062,1162,1854,1875]" italics="true" pageId="4" pageNumber="597">Ch. davidi</emphasis>
Ševčík, Hippa &amp; Burdiková
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DB2DB81C7718FFB0FF4ECA43A1D8C4E5" blockId="4.[151,1437,1766,2023]" pageId="4" pageNumber="597">
<keyLead id="60631B297718FFB0FF4ECA43A1D8C4E5" pageId="4" pageNumber="597">- Scutum light yellowish brown, without bands or brown, with thin darker brown oblique bands. Tergite IX apparently longer than broad, length/width ratio about 4/3 or more............................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="6066A0B97718FFB1FF4ECA89A1D8C3C6" lastPageId="5" lastPageNumber="598" pageId="4" pageNumber="597">
<paragraph id="DB2DB81C7718FFB0FF4ECA89A1D9C4A2" blockId="4.[151,1437,1766,2023]" pageId="4" pageNumber="597">
<keyLead id="60631B297718FFB0FF4ECA89A1D9C4A2" pageId="4" pageNumber="597">
3. Tergite IX length/width ratio about 4/3. Form of tergite IX—
<figureCitation id="43A9A4997718FFB0FCEFCA89A72BC4C0" box="[822,879,1942,1964]" captionStart="FIGURES 79" captionStartId="4.[151,264,595,619]" captionTargetBox="[307,1287,181,571]" captionTargetId="figure-29@4.[300,1288,181,572]" captionTargetPageId="4" captionText="FIGURES 79. Form of tergite IX (not in scale): 7—Ch. davidi (by Ševčík et al. 2021: Fig. 13C); 8—Ch. cavernae (by Colless 1962: Fig. 1c); 9—Ch. lagangеnsis sp. n." figureDoi="http://doi.org/10.5281/zenodo.6332237" httpUri="https://zenodo.org/record/6332237/files/figure.png" pageId="4" pageNumber="597">Fig. 8</figureCitation>
. Gonostylus stout, with a thorn-like apical spine (
<bibRefCitation id="BF03C5ED7718FFB0FA8DCA89A552C4A2" author="Colless, D. H." pageId="4" pageNumber="597" pagination="437 - 439" refId="ref3136" refString="Colless, D. H. (1962) Chetoneura cavernae n. gen., n. sp. from Batu Caves, Malaya (Diptera: Mycetophilidae). Pacific Insects, Honolulu, 4 (2), 437 - 439." type="journal article" year="1962">Colless 1962</bibRefCitation>
:
<figureCitation id="43A9A4997718FFB0FEC6CAACA525C4A1" box="[287,353,1971,1994]" captionStart="FIGURE 1" captionStartId="1.[151,250,1947,1971]" captionTargetId="figure-281@1.[478,1110,991,1921]" captionTargetPageId="1" captionText="FIGURE 1. Chetoneura lagangеnsis sp. n., habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6335584" httpUri="https://zenodo.org/record/6335584/files/figure.png" pageId="4" pageNumber="597">Fig. 1c</figureCitation>
)........................................................................
<taxonomicName id="1C92C39F7718FFB0FB09CAACA1D9C4A2" ID-CoL="TYJK" authority="Colless" authorityName="Colless" authorityYear="1962" box="[1232,1437,1971,1993]" class="Insecta" family="Keroplatidae" genus="Chetoneura" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="597" phylum="Arthropoda" rank="species" species="cavernae">
<emphasis id="E9E6640E7718FFB0FB09CAACA114C4A2" box="[1232,1360,1971,1993]" italics="true" pageId="4" pageNumber="597">Ch. cavernae</emphasis>
Colless
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DB2DB81C7718FFB1FF4ECACEA1D8C3C6" blockId="4.[151,1437,1766,2023]" lastBlockId="5.[151,1437,151,232]" lastPageId="5" lastPageNumber="598" pageId="4" pageNumber="597">
<keyLead id="60631B297718FFB1FF4ECACEA1D8C3C6" lastPageId="5" lastPageNumber="598" pageId="4" pageNumber="597">
- Tergite IX length/width ratio about 5/3. Form of tergite IX—
<figureCitation id="43A9A4997718FFB0FCF3CACEA726C48D" box="[810,866,2001,2023]" captionStart="FIGURES 79" captionStartId="4.[151,264,595,619]" captionTargetBox="[307,1287,181,571]" captionTargetId="figure-29@4.[300,1288,181,572]" captionTargetPageId="4" captionText="FIGURES 79. Form of tergite IX (not in scale): 7—Ch. davidi (by Ševčík et al. 2021: Fig. 13C); 8—Ch. cavernae (by Colless 1962: Fig. 1c); 9—Ch. lagangеnsis sp. n." figureDoi="http://doi.org/10.5281/zenodo.6332237" httpUri="https://zenodo.org/record/6332237/files/figure.png" pageId="4" pageNumber="597">Fig. 9</figureCitation>
. Gonostylus not stout, elongated in ventral view, with elongated sharp apical spine (
<figureCitation id="43A9A4997719FFB1FE0DCD88A66EC3C6" box="[468,554,151,173]" captionStart="FIGURES 36" captionStartId="3.[151,264,1731,1755]" captionTargetBox="[201,1424,197,1695]" captionTargetId="figure-17@3.[162,1425,181,1707]" captionTargetPageId="3" captionText="FIGURES 36. Chetoneura lagangеnsis sp. n., male terminalia: 3—dorsal view, 4—lateral view; 5—ventral view; 6—posterior view. Abbreviation: aps = apical spine of gonostylus; cer = cerci; gcx = gonocoxite; gst = gonostylus; hyp = hypoproct; T9 = tergite IX." figureDoi="http://doi.org/10.5281/zenodo.6332235" httpUri="https://zenodo.org/record/6332235/files/figure.png" pageId="5" pageNumber="598">Figs 56</figureCitation>
).................................
<taxonomicName id="1C92C39F7719FFB1FC1ECD88A119C3C6" ID-CoL="9J73F" authority="Kazandzhieva &amp; Bechev" authorityName=", Kazandzhieva &amp; Bechev" authorityYear="2022" box="[967,1373,151,173]" class="Insecta" family="Keroplatidae" genus="Chetoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="lagangensis" status="sp. nov.">
<emphasis id="E9E6640E7719FFB1FC1ECD88A026C3C7" box="[967,1122,151,172]" italics="true" pageId="5" pageNumber="598">Ch. lagangensis</emphasis>
, Kazandzhieva &amp; Bechev
</taxonomicName>
,
<emphasis id="E9E6640E7719FFB1FABFCD87A1D8C3C6" bold="true" box="[1382,1436,152,173]" pageId="5" pageNumber="598">
<taxonomicNameLabel id="F2D5D9757719FFB1FABFCD87A1D8C3C6" box="[1382,1436,152,173]" pageId="5" pageNumber="598" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="6066A0B97719FFB1FF4ECDAAA1D8C383" pageId="5" pageNumber="598">
<paragraph id="DB2DB81C7719FFB1FF4ECDAAA1D9C3A0" blockId="5.[151,1437,151,232]" box="[151,1437,181,203]" pageId="5" pageNumber="598">
<keyLead id="60631B297719FFB1FF4ECDAAA1D9C3A0" box="[151,1437,181,203]" pageId="5" pageNumber="598">
4. Fore tibia without apical spur.........................................................
<taxonomicName id="1C92C39F7719FFB1FB6FCDAAA1D9C3A0" ID-CoL="8BFXN" authority="(Papp)" baseAuthorityName="Papp" baseAuthorityYear="2006" box="[1206,1437,181,203]" class="Insecta" family="Keroplatidae" genus="Chetoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="oligoradiata">
<emphasis id="E9E6640E7719FFB1FB6FCDAAA112C3A1" box="[1206,1366,181,202]" italics="true" pageId="5" pageNumber="598">Ch. oligoradiata</emphasis>
(Papp)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="DB2DB81C7719FFB1FF4ECDCCA1D8C383" blockId="5.[151,1437,151,232]" box="[151,1436,210,232]" pageId="5" pageNumber="598">
<keyLead id="60631B297719FFB1FF4ECDCCA1D8C383" box="[151,1436,210,232]" pageId="5" pageNumber="598">
- Fore tibia with very short apical spur.......................................
<taxonomicName id="1C92C39F7719FFB1FBC7CDCDA1D8C383" ID-CoL="TYJL" authority="Amorim &amp; Niu" authorityName="Amorim &amp; Niu" authorityYear="2008" box="[1054,1436,210,232]" class="Insecta" family="Keroplatidae" genus="Chetoneura" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="shennonggongensis">
<emphasis id="E9E6640E7719FFB1FBC7CDCDA146C38C" box="[1054,1282,210,231]" italics="true" pageId="5" pageNumber="598">Ch. shennonggongensis</emphasis>
Amorim &amp; Niu
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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