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<mods:title id="9333EB58493D78C52D4E00A96D9B3FF3">New species and findings of Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from Central Asia, with DNA barcoding of known species of the genus</mods:title>
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<mods:namePart id="C7DCA15052776D68350E7E9E160C0067">Semenchenko, Alexander A.</mods:namePart>
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<mods:affiliation id="75B9456B8A26CB98AE8D5E170D6DE9A4">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia. semenchenko _ alexander @ mail. ru; https: // orcid. org / 0000 - 0001 - 7207 - 9529</mods:affiliation>
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<mods:affiliation id="5D148EF59FBDE6C1559146F256D7A09B">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia. triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
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Key to the known species of
<taxonomicName id="69B471DDFF82FFA574E1FD4BFD53FCD5" ID-CoL="6C9D" ID-ENA="994279" authority="Oliver" authorityName="Oliver" authorityYear="1959" box="[505,696,752,778]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="9CC0D64CFF82FFA574E1FD4BFD8AFCD5" bold="true" box="[505,609,752,778]" italics="true" pageId="2" pageNumber="561">Pagastia</emphasis>
Oliver
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<paragraph id="AE0B0A5EFF82FFA5758FFC83FF30FC8D" blockId="2.[151,1442,824,1847]" box="[151,219,824,850]" pageId="2" pageNumber="561">Males</paragraph>
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1. Eyes hairy. Aedeagal lobes reduced (
<emphasis id="9CC0D64CFF82FFA5775CFCE2FC4CFCB0" bold="true" box="[580,935,856,879]" pageId="2" pageNumber="561">
subgenus
<taxonomicName id="69B471DDFF82FFA577B1FCE3FC4CFCB0" authority="Sublette" authorityName="Sublette" box="[681,935,856,879]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="subGenus" subGenus="Hesperodiamesa">
<emphasis id="9CC0D64CFF82FFA577B1FCE3FCA6FCB1" bold="true" box="[681,845,856,878]" italics="true" pageId="2" pageNumber="561">Hesperodiamesa</emphasis>
Sublette
</taxonomicName>
</emphasis>
). Basal lobe of gonocoxite small...................................................................
<taxonomicName id="69B471DDFF82FFA5767DFCCCFB97FC53" ID-CoL="88ZRY" authority="(Garrett, 1925)" baseAuthorityName="Garrett" baseAuthorityYear="1925" box="[869,1148,886,908]" class="Insecta" family="Chironomidae" genus="Prodiamesa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="sequax" subGenus="Hesperodiamesa">
<emphasis id="9CC0D64CFF82FFA5767DFCCCFC93FC53" box="[869,888,887,908]" italics="true" pageId="2" pageNumber="561">P.</emphasis>
(
<emphasis id="9CC0D64CFF82FFA57666FCCCFC7CFC53" box="[894,919,887,908]" italics="true" pageId="2" pageNumber="561">H.</emphasis>
)
<emphasis id="9CC0D64CFF82FFA576BCFCCCFC0CFC53" box="[932,999,887,908]" italics="true" pageId="2" pageNumber="561">sequax</emphasis>
(Garrett, 1925)
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57191FCCDFAADFC54" author="Makarchenko, E. A." box="[1161,1350,886,908]" pageId="2" pageNumber="561" pagination="115 - 128" refId="ref8752" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
, Fig. 18)
</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFC2FFA77FC76" blockId="2.[151,1442,824,1847]" box="[151,1436,915,938]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFC2FFA77FC76" box="[151,1436,915,938]" pageId="2" pageNumber="561">
- Eyes bare or pubescent. One or two aedeagal lobes present (
<emphasis id="9CC0D64CFF82FFA5763CFC2FFBCEFC76" bold="true" box="[804,1061,915,937]" pageId="2" pageNumber="561">
subgenus
<taxonomicName id="69B471DDFF82FFA57690FC28FBCEFC76" ID-CoL="6C9D" authority="Oliver" authorityName="Oliver" authorityYear="1959" box="[904,1061,915,937]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="9CC0D64CFF82FFA57690FC28FC36FC77" bold="true" box="[904,989,915,936]" italics="true" pageId="2" pageNumber="561">Pagastia</emphasis>
Oliver
</taxonomicName>
</emphasis>
). Basal lobe of gonocoxite large... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FFC0AFA77FC3C" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFC0AFA77FC19" blockId="2.[151,1442,824,1847]" box="[151,1436,945,967]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFC0AFA77FC19" box="[151,1436,945,967]" pageId="2" pageNumber="561">2. Both median and lateral aedeagal lobes present.............................................................. 3</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFC74FA77FC3C" blockId="2.[151,1442,824,1847]" box="[151,1436,974,996]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFC74FA77FC3C" box="[151,1436,974,996]" pageId="2" pageNumber="561">- Only lateral aedeagal lobe present........................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FFC57FA77FB86" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFC57FA77FBC0" blockId="2.[151,1442,824,1847]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFC57FA77FBC0" pageId="2" pageNumber="561">3. Antepronotum with dorsal and lateral setae in widely separated groups. Median aedeagal lobe digitated, widest in distal part or wedge-shaped and widest at base......................................................................... 5</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFB9CFA77FB86" blockId="2.[151,1442,824,1847]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFB9CFA77FB86" pageId="2" pageNumber="561">- Antepronotum completely covered with setae. Median aedeagal lobe widest medially, with apex forming a sharp hook or wedge-shaped, with fingerlike apex....................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FFBDAFA77FB6E" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFBDAFA77FB4B" blockId="2.[151,1442,824,1847]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFBDAFA77FB4B" pageId="2" pageNumber="561">
4. Median aedeagal lobe widest medially, then tapering abruptly and with the apex forming a sharp hook.............................................................
<taxonomicName id="69B471DDFF82FFA577F6FBC4FBBEFB4B" ID-CoL="75KP4" authority="(Roback, 1957)" baseAuthorityName="Roback" baseAuthorityYear="1957" box="[750,1109,1150,1172]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="partica" subGenus="Pagastia">
<emphasis id="9CC0D64CFF82FFA577F6FBC4FD16FB4B" box="[750,765,1151,1172]" italics="true" pageId="2" pageNumber="561">P</emphasis>
. (
<emphasis id="9CC0D64CFF82FFA57609FBC4FC8DFB4B" box="[785,870,1151,1172]" italics="true" pageId="2" pageNumber="561">Pagastia</emphasis>
)
<emphasis id="9CC0D64CFF82FFA5766CFBC4FC52FB4B" box="[884,953,1151,1172]" italics="true" pageId="2" pageNumber="561">partica</emphasis>
(Roback, 1957)
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57179FBC5FAF4FB4C" author="Makarchenko, E. A." box="[1121,1311,1150,1172]" pageId="2" pageNumber="561" pagination="115 - 128" refId="ref8752" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
,
<figureCitation id="368F16DBFF82FFA57031FBC5FA9AFB4B" box="[1321,1393,1150,1172]" captionStart="FIGURE 16" captionStartId="9.[151,250,1389,1413]" captionTargetBox="[152,1404,193,1358]" captionTargetId="figure-18@9.[151,1435,181,1366]" captionTargetPageId="9" captionText="FIGURE 16. Bayesian tree based on mitochondrial COI and COII genes for available members of the genus Pagastia Oliver with Pseudodiamesa stackelbergi (Goetghebuer) as outgroup. Bayesian posterior probabilities (PP) are given above tree nodes and bootstrap support values found in the ML analysis are shown below nodes." figureDoi="http://doi.org/10.5281/zenodo.4668495" httpUri="https://zenodo.org/record/4668495/files/figure.png" pageId="2" pageNumber="561">Figs.16</figureCitation>
17)
</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFB27FA77FB6E" blockId="2.[151,1442,824,1847]" box="[151,1436,1180,1202]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFB27FA77FB6E" box="[151,1436,1180,1202]" pageId="2" pageNumber="561">
- Median aedeagal lobe wedge-shaped, with finger-like apex............................
<taxonomicName id="69B471DDFF82FFA57174FB27FB17FB6E" ID-CoL="6C9D" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2021" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2021" box="[1132,1276,1180,1201]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="hanseni" status="sp. nov." subGenus="Pagastia">
<emphasis id="9CC0D64CFF82FFA57174FB27FB90FB6E" box="[1132,1147,1180,1201]" italics="true" pageId="2" pageNumber="561">P</emphasis>
. (
<emphasis id="9CC0D64CFF82FFA57196FB27FB76FB6E" box="[1166,1181,1180,1201]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA571A9FB27FB17FB6E" box="[1201,1276,1180,1201]" italics="true" pageId="2" pageNumber="561">hanseni</emphasis>
</taxonomicName>
<emphasis id="9CC0D64CFF82FFA5701AFB27FAA5FB6E" bold="true" box="[1282,1358,1180,1201]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="87F36B37FF82FFA5701AFB27FAA5FB6E" box="[1282,1358,1180,1201]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="368F16DBFF82FFA57043FB27FA7EFB6E" box="[1371,1429,1180,1202]" captionStart="FIGURES 17" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="2" pageNumber="561">Fig. 7</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FFB02FA77FB33" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFB02FA77FB10" blockId="2.[151,1442,824,1847]" box="[151,1436,1209,1231]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFB02FA77FB10" box="[151,1436,1209,1231]" pageId="2" pageNumber="561">5. Median aedeagal lobe digitated, widest in distal part......................................................... 6</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFB6CFA77FB33" blockId="2.[151,1442,824,1847]" box="[151,1436,1238,1260]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFB6CFA77FB33" box="[151,1436,1238,1260]" pageId="2" pageNumber="561">
- Median aedeagal lobe wedge-shaped, with toothed apex......................
<emphasis id="9CC0D64CFF82FFA5710CFB6CFBC8FB33" box="[1044,1059,1239,1260]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.
<taxonomicName id="69B471DDFF82FFA57136FB6DFB17FB34" ID-CoL="6C9D" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2021" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2021" box="[1070,1276,1238,1260]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="caelestomontana" status="sp. nov." subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF82FFA5712EFB6CFBAEFB33" box="[1078,1093,1239,1260]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA57141FB6DFB17FB34" box="[1113,1276,1238,1259]" italics="true" pageId="2" pageNumber="561">caelestomontana</emphasis>
</taxonomicName>
<emphasis id="9CC0D64CFF82FFA5701AFB6DFAA5FB34" bold="true" box="[1282,1358,1238,1259]" pageId="2" pageNumber="561">
<taxonomicNameLabel id="87F36B37FF82FFA5701AFB6DFAA5FB34" box="[1282,1358,1238,1259]" pageId="2" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="368F16DBFF82FFA57043FB6DFA7EFB33" box="[1371,1429,1238,1260]" captionStart="FIGURES 17" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="2" pageNumber="561">Fig. 2</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FFB4FFA77FAF9" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFB4FFA77FAD6" blockId="2.[151,1442,824,1847]" box="[151,1436,1267,1290]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFB4FFA77FAD6" box="[151,1436,1267,1290]" pageId="2" pageNumber="561">
6. Gonostylus subapical with “heel”..................
<emphasis id="9CC0D64CFF82FFA57609FB4FFCCBFAD6" box="[785,800,1268,1289]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.
<taxonomicName id="69B471DDFF82FFA57634FB4FFBDEFAD5" ID-CoL="6TKP3" authority="(Tokunaga, 1936)" baseAuthorityName="Tokunaga" baseAuthorityYear="1936" box="[812,1077,1268,1290]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="nivis" subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF82FFA5762CFB4FFCA8FAD6" box="[820,835,1268,1289]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA5764FFB4FFC6FFAD6" box="[855,900,1268,1289]" italics="true" pageId="2" pageNumber="561">nivis</emphasis>
(Tokunaga, 1936)
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57159FB4FFB15FAD5" author="Makarchenko, E. A." box="[1089,1278,1268,1290]" pageId="2" pageNumber="561" pagination="607 - 621" refId="ref8638" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276, 468 - 480, 607 - 621. [in Russian]" type="journal article" year="2006">Makarchenko 2006</bibRefCitation>
, Fig. 180, 510)
</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFAAAFA77FAF9" blockId="2.[151,1442,824,1847]" box="[151,1436,1297,1319]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFAAAFA77FAF9" box="[151,1436,1297,1319]" pageId="2" pageNumber="561">- Gonostylus subapical without “heel”..................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FFA94FA49FAA0" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFA94FA49FA9B" blockId="2.[151,1442,824,1847]" box="[151,1442,1326,1348]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFA94FA49FA9B" box="[151,1442,1326,1348]" pageId="2" pageNumber="561">
7. Apex of the gonostylus is angled...............
<emphasis id="9CC0D64CFF82FFA577CCFA94FD0CFA9B" box="[724,743,1327,1348]" italics="true" pageId="2" pageNumber="561">P.</emphasis>
<taxonomicName id="69B471DDFF82FFA577F4FA95FBABFA9B" ID-CoL="75KP3" authority="(Tshernovskij, 1949)" baseAuthorityName="Tshernovskij" baseAuthorityYear="1949" box="[748,1088,1326,1348]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="orientalis" subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF82FFA577ECFA94FCE8FA9B" box="[756,771,1327,1348]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA5760FFA95FC98FA9C" box="[791,883,1326,1347]" italics="true" pageId="2" pageNumber="561">orientalis</emphasis>
(Tshernovskij, 1949)
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57155FA95FAE1FA9B" author="Makarchenko, E. A." box="[1101,1290,1326,1348]" pageId="2" pageNumber="561" pagination="607 - 621" refId="ref8638" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276, 468 - 480, 607 - 621. [in Russian]" type="journal article" year="2006">Makarchenko 2006</bibRefCitation>
, Fig. 181, 15)
</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFAF7FA49FAA0" blockId="2.[151,1442,824,1847]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFAF7FA49FAA0" pageId="2" pageNumber="561">
- Apex of the gonostylus is broadly rounded..............................................................................................
<emphasis id="9CC0D64CFF82FFA57733FAD1FDD1FAA0" box="[555,570,1386,1407]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.
<taxonomicName id="69B471DDFF82FFA5775EFAD2FBB9FAA1" ID-CoL="75L2C" authority="Makarchenko, Kerkis et Ivanchenko, 1997" authorityName="Makarchenko, Kerkis et Ivanchenko" authorityYear="1997" box="[582,1106,1385,1407]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="altaica" subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF82FFA57756FAD1FDB6FAA0" box="[590,605,1386,1407]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA57768FAD2FD58FAA1" box="[624,691,1385,1406]" italics="true" pageId="2" pageNumber="561">altaica</emphasis>
<bibRefCitation id="CA2577AFFF82FFA577A1FAD2FBB9FAA1" author="Makarchenko, E. A. &amp; Kerkis, I. E. &amp; Ivanchenko, O. V." box="[697,1106,1385,1407]" pageId="2" pageNumber="561" pagination="1 - 8" refId="ref8504" refString="Makarchenko, E. A., Kerkis, I. E. &amp; Ivanchenko, O. V. (1997) Morphokaryological description of Pagastia altaica sp. n. (Diptera, Chironomidae) from Altai Mountains, with the key to Holarctic species of Pagastia Oliver. Far Eastern Entomologist, 43, 1 - 8." type="journal article" year="1997">Makarchenko, Kerkis et Ivanchenko, 1997</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57147FAD2FAB9FAA1" author="Makarchenko, E. A. &amp; Kerkis, I. E. &amp; Ivanchenko, O. V." box="[1119,1362,1385,1407]" pageId="2" pageNumber="561" pagination="1 - 8" refId="ref8504" refString="Makarchenko, E. A., Kerkis, I. E. &amp; Ivanchenko, O. V. (1997) Morphokaryological description of Pagastia altaica sp. n. (Diptera, Chironomidae) from Altai Mountains, with the key to Holarctic species of Pagastia Oliver. Far Eastern Entomologist, 43, 1 - 8." type="journal article" year="1997">
Makarchenko
<emphasis id="9CC0D64CFF82FFA571F2FAD1FAF0FAA1" box="[1258,1307,1385,1407]" italics="true" pageId="2" pageNumber="561">et al.</emphasis>
1997
</bibRefCitation>
,
<figureCitation id="368F16DBFF82FFA57044FAD2FA7DFAA0" box="[1372,1430,1385,1407]" captionStart="FIGURES 17" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="2" pageNumber="561">Fig. 1</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<paragraph id="AE0B0A5EFF82FFA5758FFA3DFDA2FA43" blockId="2.[151,1442,824,1847]" box="[151,585,1414,1436]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFA3DFDA2FA43" box="[151,585,1414,1436]" pageId="2" pageNumber="561">
<keyStep id="154012FBFF82FFA5758FFA3DFDA2FA43" box="[151,585,1414,1436]" pageId="2" pageNumber="561">8. Anal point absent. Alula as well as M</keyStep>
</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA57752FA2DFD8BFA7D" blockId="2.[151,1442,824,1847]" box="[586,608,1430,1442]" pageId="2" pageNumber="561">1+2</paragraph>
<paragraph id="AE0B0A5EFF82FFA57778FA3CFD6AFA43" blockId="2.[151,1442,824,1847]" box="[608,641,1415,1436]" pageId="2" pageNumber="561">, M</paragraph>
<paragraph id="AE0B0A5EFF82FFA57799FA2DFD7DFA7D" blockId="2.[151,1442,824,1847]" box="[641,662,1430,1442]" pageId="2" pageNumber="561">3+4</paragraph>
<paragraph id="AE0B0A5EFF82FFA5778FFA3CFD03FA44" blockId="2.[151,1442,824,1847]" box="[663,744,1414,1436]" pageId="2" pageNumber="561">, and Cu</paragraph>
<paragraph id="AE0B0A5EFF82FFA577F0FA2DFD04FA7D" blockId="2.[151,1442,824,1847]" box="[744,751,1430,1442]" pageId="2" pageNumber="561">1</paragraph>
<paragraph id="AE0B0A5EFF82FFA577EDFA3DFA77FA65" blockId="2.[151,1442,824,1847]" pageId="2" pageNumber="561">
of wing with setae.................................................................................
<emphasis id="9CC0D64CFF82FFA577CAFA1FFD0AFA66" box="[722,737,1444,1465]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.
<taxonomicName id="69B471DDFF82FFA577F5FA1FFBF4FA65" ID-CoL="9HNS6" authority="Makarchenko" authorityName="Makarchenko" authorityYear="2019" box="[749,1055,1444,1466]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="subletteorum" subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF82FFA577EDFA1FFCEFFA66" box="[757,772,1444,1465]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA57600FA1FFC7EFA66" box="[792,917,1444,1465]" italics="true" pageId="2" pageNumber="561">subletteorum</emphasis>
Makarchenko
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57133FA1FFB02FA66" author="Makarchenko, E. A." box="[1067,1257,1444,1466]" pageId="2" pageNumber="561" pagination="115 - 128" refId="ref8752" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
,
<figureCitation id="368F16DBFF82FFA571EBFA1FFAA6FA65" box="[1267,1357,1444,1466]" captionStart="FIGURES 17" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="2" pageNumber="561">Figs. 16</figureCitation>
, 1920)
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFA7AFA77FA09" blockId="2.[151,1442,824,1847]" box="[151,1436,1473,1500]" pageId="2" pageNumber="561">
- Anal point present. Alula and M
<subScript id="3230081BFF82FFA5770FFA6BFDC6FA03" attach="left" box="[535,557,1488,1500]" fontSize="5" pageId="2" pageNumber="561">1+2</subScript>
, M
<subScript id="3230081BFF82FFA57756FA6BFD8FFA03" attach="left" box="[590,612,1488,1500]" fontSize="5" pageId="2" pageNumber="561">3+4</subScript>
, Cu
<subScript id="3230081BFF82FFA57794FA6BFD78FA03" attach="left" box="[652,659,1488,1500]" fontSize="5" pageId="2" pageNumber="561">1</subScript>
without setae.................................................... 9
</paragraph>
<keyStep id="154012FBFF82FFA5758FFA65FA77F9CE" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FFA65FA77FA2C" blockId="2.[151,1442,824,1847]" box="[151,1436,1502,1524]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFA65FA77FA2C" box="[151,1436,1502,1524]" pageId="2" pageNumber="561">9. Anal point almost parallel-sided in dorsal view; lateral aedeagal lobe tapering to thin apex.......................... 10</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FFA47FA77F9CE" blockId="2.[151,1442,824,1847]" box="[151,1436,1532,1554]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FFA47FA77F9CE" box="[151,1436,1532,1554]" pageId="2" pageNumber="561">- Anal point tapering to apex; lateral aedeagal lobe broad in distal part........................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FF9A2FA77F994" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FF9A2FA79F9F0" blockId="2.[151,1442,824,1847]" box="[151,1426,1561,1583]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FF9A2FA79F9F0" box="[151,1426,1561,1583]" pageId="2" pageNumber="561">
10. Anal point with rounded apex and without apical peg ...
<emphasis id="9CC0D64CFF82FFA577E6F9A1FCE6F9F0" box="[766,781,1562,1583]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.
<taxonomicName id="69B471DDFF82FFA57601F9A2FBC6F9F0" ID-CoL="6TKP4" authority="Oliver, 1959" authorityName="Oliver" authorityYear="1959" box="[793,1069,1561,1583]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="orthogonia" subGenus="Pagastia">
(
<emphasis id="9CC0D64CFF82FFA57639F9A1FCDBF9F0" box="[801,816,1562,1583]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA5765CF9A2FC44F9F1" box="[836,943,1561,1582]" italics="true" pageId="2" pageNumber="561">orthogonia</emphasis>
<bibRefCitation id="CA2577AFFF82FFA576ACF9A2FBC6F9F0" author="Oliver, D. R." box="[948,1069,1561,1583]" pageId="2" pageNumber="561" pagination="48 - 64" refId="ref9044" refString="Oliver, D. R. (1959) Some Diamesini (Chironomidae) from the Nearctic and Palaearctic. Entomologisk Tidskrift, 80, 48 - 64." type="journal article" year="1959">Oliver, 1959</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="CA2577AFFF82FFA57121F9A2FB1CF9F1" author="Makarchenko, E. A." box="[1081,1271,1561,1583]" pageId="2" pageNumber="561" pagination="115 - 128" refId="ref8752" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
,
<figureCitation id="368F16DBFF82FFA57019F9A2FA83F9F1" box="[1281,1384,1561,1583]" captionStart-0="FIGURES 17" captionStart-1="FIGURES 811" captionStart-2="FIGURES 1215" captionStartId-0="4.[151,264,1710,1734]" captionStartId-1="5.[151,264,1704,1728]" captionStartId-2="7.[151,264,1495,1519]" captionTargetBox-0="[162,1428,201,1667]" captionTargetBox-1="[151,1435,429,1666]" captionTargetBox-2="[151,1435,194,1457]" captionTargetId-0="figure-24@4.[151,1436,181,1686]" captionTargetId-1="figure-134@5.[151,1435,417,1680]" captionTargetId-2="figure-18@7.[151,1435,181,1471]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." captionText-1="FIGURES 811. Pagastia (P.) caelestomontana sp. nov. (89) and P. (P.) hanseni sp. nov. (1011), males. 8, 10, hypopygium in dorsal view; 9, gonostylus; 11, anal point." captionText-2="FIGURES 1215. Type localities of Pagastia (P.) caelestomontana sp. nov. (12), P. (P.) hanseni sp. nov. (13), and P. (P.) aff. lanceolata (Tokunaga) (1415). 12, Otmek River, Talas district, Kirgizstan (photo by E.S. Chertoprud); 13, Soktosh River, Gorno-Badakhshan Autonomous Region, Roshtqala District, Tajikistan (photo by D.M. Palatov); 14, Tuzumtaikul River, Gorno- Badakhshan Autonomous Region, Shughnon District, Tajikistan (photo by M.V. Vinarski); 15, spring in front of the mouth of the Bartang River, Gorno-Badakhshan Autonomous Region, Rushanskiy District, Tajikistan (photo by M.V. Vinarski)." figureDoi-0="http://doi.org/10.5281/zenodo.4668489" figureDoi-1="http://doi.org/10.5281/zenodo.4668491" figureDoi-2="http://doi.org/10.5281/zenodo.4668493" httpUri-0="https://zenodo.org/record/4668489/files/figure.png" httpUri-1="https://zenodo.org/record/4668491/files/figure.png" httpUri-2="https://zenodo.org/record/4668493/files/figure.png" pageId="2" pageNumber="561">Figs. 713</figureCitation>
, 21)
</keyLead>
</paragraph>
<paragraph id="AE0B0A5EFF82FFA5758FF98CFA77F994" blockId="2.[151,1442,824,1847]" box="[151,1436,1590,1612]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FF98CFA77F994" box="[151,1436,1590,1612]" pageId="2" pageNumber="561">- Anal point with pointed apical peg....................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="154012FBFF82FFA5758FF9EFFA77F91D" pageId="2" pageNumber="561">
<paragraph id="AE0B0A5EFF82FFA5758FF9EFFA76F958" blockId="2.[151,1442,824,1847]" pageId="2" pageNumber="561">
<keyLead id="1545A96BFF82FFA5758FF9EFFA76F958" pageId="2" pageNumber="561">
11. Gonostylus with small basal lobe in outer side; apical part rounded, without “heel”. AR 1.561.75...........................................................................
<emphasis id="9CC0D64CFF82FFA57674F9C9FC94F958" box="[876,895,1650,1671]" italics="true" pageId="2" pageNumber="561">P.</emphasis>
<taxonomicName id="69B471DDFF82FFA5769CF9CAFB53F959" ID-CoL="6TKP2" authority="Endo, 2004" authorityName="Endo" authorityYear="2004" box="[900,1208,1649,1671]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="561" phylum="Arthropoda" rank="species" species="hidakamontana" subGenus="Pagastia">
(
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.)
<emphasis id="9CC0D64CFF82FFA576B6F9CAFBA8F959" box="[942,1091,1649,1670]" italics="true" pageId="2" pageNumber="561">hidakamontana</emphasis>
<bibRefCitation id="CA2577AFFF82FFA57152F9CAFB53F959" author="Endo, K." box="[1098,1208,1649,1671]" pageId="2" pageNumber="561" pagination="277 - 289" refId="ref7984" refString="Endo, K. (2004) Genus Pagastia Oliver (Diptera: Chironomidae) from Japan, with description of a new species. Entomological Science, 7, 277 - 289. https: // doi. org / 10.1111 / j. 1479 - 8298.2004.00074. x" type="journal article" year="2004">Endo, 2004</bibRefCitation>
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(
<bibRefCitation id="CA2577AFFF82FFA571DDF9CAFADAF959" author="Endo, K." box="[1221,1329,1649,1671]" pageId="2" pageNumber="561" pagination="277 - 289" refId="ref7984" refString="Endo, K. (2004) Genus Pagastia Oliver (Diptera: Chironomidae) from Japan, with description of a new species. Entomological Science, 7, 277 - 289. https: // doi. org / 10.1111 / j. 1479 - 8298.2004.00074. x" type="journal article" year="2004">Endo 2004</bibRefCitation>
,
<figureCitation id="368F16DBFF82FFA57023F9CAFA7EF958" box="[1339,1429,1649,1671]" captionStart="FIGURES 17" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="2" pageNumber="561">Figs. 16</figureCitation>
)
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- Gonostylus with rounded basal lobe in outer side and with lobe like “heel” in apical part. AR 1.221.26.........................................
<emphasis id="9CC0D64CFF82FFA57718F917FDF8F91E" box="[512,531,1708,1729]" italics="true" pageId="2" pageNumber="561">P.</emphasis>
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(
<emphasis id="9CC0D64CFF82FFA57738F917FDC4F91E" box="[544,559,1708,1729]" italics="true" pageId="2" pageNumber="561">P</emphasis>
.)
<emphasis id="9CC0D64CFF82FFA5775AF917FD4EF91E" box="[578,677,1708,1729]" italics="true" pageId="2" pageNumber="561">donoliveri</emphasis>
<bibRefCitation id="CA2577AFFF82FFA577B3F917FC3BF91E" author="Makarchenko, E. A. &amp; Hansen, D. C." box="[683,976,1708,1730]" pageId="2" pageNumber="561" pagination="171 - 176" refId="ref8815" refString="Makarchenko, E. A. &amp; Hansen, D. C. (2020) Pagastia (P.) donoliveri sp. nov. - a new Nearctic alpine stream chironomid species (Diptera: Chironomidae: Diamesinae) from the Beartooth Mountains, Wyoming, U. S. A. Zootaxa, 4755 (1), 171 - 176. https: // doi. org / 10.11646 / zootaxa. 4755.1.9" type="journal article" year="2020">Makarchenko et Hansen, 2020</bibRefCitation>
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(
<bibRefCitation id="CA2577AFFF82FFA576C5F917FAEAF91E" author="Makarchenko, E. A. &amp; Hansen, D. C." box="[989,1281,1708,1730]" pageId="2" pageNumber="561" pagination="171 - 176" refId="ref8815" refString="Makarchenko, E. A. &amp; Hansen, D. C. (2020) Pagastia (P.) donoliveri sp. nov. - a new Nearctic alpine stream chironomid species (Diptera: Chironomidae: Diamesinae) from the Beartooth Mountains, Wyoming, U. S. A. Zootaxa, 4755 (1), 171 - 176. https: // doi. org / 10.11646 / zootaxa. 4755.1.9" type="journal article" year="2020">Makarchenko &amp; Hansen 2020</bibRefCitation>
,
<figureCitation id="368F16DBFF82FFA57013F917FA95F91D" box="[1291,1406,1708,1730]" captionStart="FIGURES 17" captionStartId="4.[151,264,1710,1734]" captionTargetBox="[162,1428,201,1667]" captionTargetId="figure-24@4.[151,1436,181,1686]" captionTargetPageId="4" captionText="FIGURES 17. Pagastia (P.) caelestomontana sp. nov. (14) and P. (P.) hanseni sp. nov. (57), males. 1, 6, anteropronotum in dorsal view; 2, 7, aedeagal lobes; 3, 5, hypopygium in dorsal view; 4, anal point. Bl, basal lobe of gonocoxite; LAl, lateral aedeagal lobe; MAl, median aedeagal lobe TSa, transverse sternapodeme. Scale bars for figures 1, 3, 5, 6 200 µm, for figures 2, 4, 7 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4668489" httpUri="https://zenodo.org/record/4668489/files/figure.png" pageId="2" pageNumber="561">Figs. 4, 67</figureCitation>
,
<figureCitation id="368F16DBFF82FFA57090F917FA7EF91D" box="[1416,1429,1708,1730]" captionStart="FIGURES 811" captionStartId="5.[151,264,1704,1728]" captionTargetBox="[151,1435,429,1666]" captionTargetId="figure-134@5.[151,1435,417,1680]" captionTargetPageId="5" captionText="FIGURES 811. Pagastia (P.) caelestomontana sp. nov. (89) and P. (P.) hanseni sp. nov. (1011), males. 8, 10, hypopygium in dorsal view; 9, gonostylus; 11, anal point." figureDoi="http://doi.org/10.5281/zenodo.4668491" httpUri="https://zenodo.org/record/4668491/files/figure.png" pageId="2" pageNumber="561">9</figureCitation>
)
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12. Anal point narrow, tapering to pointed apex, without apical peg; gonostylus in basal part with outer angle-shaped projection. AR 2.182.42.................
<emphasis id="9CC0D64CFF82FFA57721F95CFDA7F923" box="[569,588,1767,1788]" italics="true" pageId="2" pageNumber="561">P.</emphasis>
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.)
<emphasis id="9CC0D64CFF82FFA57763F95CFCF9F923" box="[635,786,1767,1788]" italics="true" pageId="2" pageNumber="561">tianmumontana</emphasis>
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Makarchenko
<emphasis id="9CC0D64CFF82FFA576BAF95CFC58F923" box="[930,947,1767,1788]" italics="true" pageId="2" pageNumber="561">et</emphasis>
Wang, 2017
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(
<bibRefCitation id="CA2577AFFF82FFA57121F95DFAADF924" author="Makarchenko, E. A. &amp; Wang, X." box="[1081,1350,1766,1788]" pageId="2" pageNumber="561" pagination="13 - 15" refId="ref8705" refString="Makarchenko, E. A. &amp; Wang, X. (2017) Pagastia tianmumontana sp. n. - a new species of chironomids (Diptera: Chironomidae: Diamesinae) from South China. Far Eastern Entomologist, 336, 13 - 15." type="journal article" year="2017">Makarchenko &amp;Wang 2017</bibRefCitation>
, Fig. 17)
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- Anal point widest in basal part and thin apically, pointed and sometimes with apical peg; gonostylus without outer basal projec- tion and with rounded apex. AR 1.812.10........
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.
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(
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.)
<emphasis id="9CC0D64CFF82FFA57633F89AFC7BF8E9" box="[811,912,1825,1846]" italics="true" pageId="2" pageNumber="561">lanceolata</emphasis>
(Tokunaga, 1936)
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(
<bibRefCitation id="CA2577AFFF82FFA57155F89AFAE1F8E8" author="Makarchenko, E. A." box="[1101,1290,1825,1847]" pageId="2" pageNumber="561" pagination="607 - 621" refId="ref8638" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276, 468 - 480, 607 - 621. [in Russian]" type="journal article" year="2006">Makarchenko 2006</bibRefCitation>
, Fig. 180, 14)
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</subSubSection>
</treatment>
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