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<document id="848990920F74184463D617DE102ACA70" ID-DOI="10.2478/if-2019-0007" ID-ISSN="2533-4069" ID-Zenodo-Dep="5383369" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630296318417" checkinUser="diego" docAuthor="Budil, Petr &amp; Mergl, Michal" docDate="2019" docId="912E6907FFBAFFF8FF688456C4F0F962" docLanguage="en" docName="10_Budil.pdf" docOrigin="Fossil Imprint 75 (1)" docSource="http://dx.doi.org/10.2478/if-2019-0007" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Chotecops hoseri" docType="treatment" docVersion="3" lastPageNumber="81" masterDocId="6D17117FFFB8FFFAFFAF8520C57BFFD6" masterDocTitle="Trilobite Assemblage Of Calceola - Bearing Beds In Acanthopyge Limestone (Choteč Formation, Middle Devonian, Eifelian, Prague Basin, The Czech Republic)" masterLastPageNumber="91" masterPageNumber="79" pageNumber="81" updateTime="1699219012125" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="4E543C3399E65EB062A937B951BE05FC">Trilobite Assemblage Of Calceola - Bearing Beds In Acanthopyge Limestone (Choteč Formation, Middle Devonian, Eifelian, Prague Basin, The Czech Republic)</mods:title>
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<mods:date id="6C8A8C2B49DB7715C25F0A10846CEE95">2019</mods:date>
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<treatment id="912E6907FFBAFFF8FF688456C4F0F962" LSID="urn:lsid:plazi:treatment:912E6907FFBAFFF8FF688456C4F0F962" httpUri="http://treatment.plazi.org/id/912E6907FFBAFFF8FF688456C4F0F962" lastPageNumber="81" pageId="2" pageNumber="81">
<subSubSection id="519D8B9AFFBAFFF8FF688456C773FE78" pageId="2" pageNumber="81" type="nomenclature">
<paragraph id="1938D811FFBAFFF8FF688456C7C3FE58" blockId="2.[199,696,374,398]" box="[199,696,374,398]" pageId="2" pageNumber="81">
<heading id="42706F7DFFBAFFF8FF688456C7C3FE58" box="[199,696,374,398]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="81" reason="2">
<taxonomicName id="DE87A392FFBAFFF8FF688456C7C3FE58" authority="(HAWLE et CORDA, 1847)" baseAuthorityName="HAWLE et CORDA" baseAuthorityYear="1847" box="[199,696,374,398]" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="hoseri">
<emphasis id="2BF30403FFBAFFF8FF688456C7C3FE58" bold="true" box="[199,696,374,398]" pageId="2" pageNumber="81">
<emphasis id="2BF30403FFBAFFF8FF688456C442FE58" bold="true" box="[199,313,374,398]" italics="true" pageId="2" pageNumber="81">Chotecops</emphasis>
cf.
<emphasis id="2BF30403FFBAFFF8FECD8456C4DDFE58" bold="true" box="[354,422,374,398]" italics="true" pageId="2" pageNumber="81">hoseri</emphasis>
(
<bibRefCitation id="7D16A5E0FFBAFFF8FE1A8456C77AFE5B" author="Hawle, I. &amp; Corda, A. J. C." box="[437,513,374,398]" pageId="2" pageNumber="81" pagination="117 - 292" refId="ref7970" refString="Hawle, I., Corda, A. J. C. (1847): Prodrom einer Monogra- phie der bohmischen Trilobiten. - Abhandlungen der ko- niglichen bohmischen Gesellschaft der Wissenschaften, Prag, 5 (5): 117 - 292." type="journal article" year="1847">HAWLE</bibRefCitation>
et CORDA, 1847)
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</paragraph>
<paragraph id="1938D811FFBAFFF8FED884B6C773FE78" blockId="2.[375,520,406,430]" box="[375,520,406,430]" pageId="2" pageNumber="81">
<figureCitation id="81BCC494FFBAFFF8FED884B6C773FE78" box="[375,520,406,430]" captionStart="Text-fig" captionStartId="4.[129,210,1755,1776]" captionText="Text-fig. 3. Trilobite assemblage at Zadní Kobyla small quarry outcrop (Calceola-bearing locality). a–f – Chotecops cf. hoseri (HAWLE et CORDA, 1847); a, c – almost complete cephalon, CGS MM 556, internal mould, a – dorsal view, c – right frontolateral view; b – incomplete cephalon, CGS MM 557, internal mould, dorsal view; d – left free cheek with visual surface of eye, CGS MM 558, internal mould with remains of exoskeleton, lateral view; e – incomplete pygidium, CGS MM 559, dorsal view; f – incomplete pygidium, CGS MM 560, dorsal view. g – Chotecops? sp., incomplete corroded pygidium, CGS MM 561, dorsal view. h –?Lioharpes (Fritchaspis) sp., fragment of fringe, internal mould, lateral view, CGS MM 562. i – Crotalocephalus cf. affinis (HAWLE et CORDA, 1847), incomplete cranidium, CGS MM 563, internal mould, dorsal view. j–k – Acanthopyge (Acanthopyge) cf. haueri (BARRANDE, 1846), incomplete hypostomes, dorsal views, j – CGS MM 564, k – CGS MM 565. l – Acanthopyge? sp., fragment of cranidium, internal mould with remains of exoskeleton, CGS MM 566, dorsal view. m – Otarion cf. lacrimarum PŘIBYL et VANĚK, 1981, incomplete corroded cranidium, internal mould, CGS MM 567, dorsal view. n – Orbitoproetus cf. angelini (HAWLE et CORDA, 1847), juvenile pygidium, internal mould with exoskeleton, CGS MM 568, dorsal view. o – Eremiproetus? sp., negative counterpart of poorly preserved fragment of cranidium, CGS MM 569, dorsal view" pageId="2" pageNumber="81">Text-fig. 3a–f</figureCitation>
</paragraph>
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<subSubSection id="519D8B9AFFBAFFF8FF0984E6C4F0F962" pageId="2" pageNumber="81" type="description">
<paragraph id="1938D811FFBAFFF8FF0984E6C4E5FD88" blockId="2.[128,768,454,607]" pageId="2" pageNumber="81">M a t e r i a l. Eight incomplete cephala, six pygidia, numerous isolated fragments of cephala, pygidia and thoracic segments, two incomplete cheeks with preserved visual surface of the eye. Internal moulds and their counterparts, twenty five total remnants.</paragraph>
<paragraph id="1938D811FFBAFFF8FF098756C4F0F962" blockId="2.[128,768,630,1716]" pageId="2" pageNumber="81">
R e m a r k s. Preservation of even the best-preserved remains of phacopid exoskeletons does not allow positive determination of material to the species level, but it shares most features with
<taxonomicName id="DE87A392FFBAFFF8FECD87F6C5B9FCD9" authority="(HAWLE et CORDA, 1847)" baseAuthorityName="HAWLE et CORDA" baseAuthorityYear="1847" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="hoseri">
<emphasis id="2BF30403FFBAFFF8FECD87F6C75FFD38" box="[354,548,726,750]" italics="true" pageId="2" pageNumber="81">Chotecops hoseri</emphasis>
(
<bibRefCitation id="7D16A5E0FFBAFFF8FD9487F7C5C2FCD9" author="Hawle, I. &amp; Corda, A. J. C." pageId="2" pageNumber="81" pagination="117 - 292" refId="ref7970" refString="Hawle, I., Corda, A. J. C. (1847): Prodrom einer Monogra- phie der bohmischen Trilobiten. - Abhandlungen der ko- niglichen bohmischen Gesellschaft der Wissenschaften, Prag, 5 (5): 117 - 292." type="journal article" year="1847">HAWLE et CORDA, 1847</bibRefCitation>
)
</taxonomicName>
. The angle of divergence in axial furrows is, however, slightly larger in the studied material (67°), but this value is barely larger than the upper limit of the variation range (60–65°) previously known in this species (
<bibRefCitation id="7D16A5E0FFBAFFF8FDF08677C5E5FC59" author="Chlupac, I." pageId="2" pageNumber="81" pagination="1 - 172" refId="ref7228" refString="Chlupac, I. (1977): The phacopid trilobites of the Silurian and Devonian of Czechoslovakia. - Rozpravy Ustred- niho ustavu geologickeho, 23: 1 - 172." type="journal article" year="1977">Chlupáč 1977: 53</bibRefCitation>
); this feature may have also been slightly affected by deformation. In the studied samples, the maximal number of lenses in the dorso-ventral row of the eye reaches 6 (5) ommatidia (in two young holaspid specimens, see
<figureCitation id="81BCC494FFBAFFF8FD0A86F8C5E7FBC6" captionStart="Text-fig" captionStartId="4.[129,210,1755,1776]" captionText="Text-fig. 3. Trilobite assemblage at Zadní Kobyla small quarry outcrop (Calceola-bearing locality). a–f – Chotecops cf. hoseri (HAWLE et CORDA, 1847); a, c – almost complete cephalon, CGS MM 556, internal mould, a – dorsal view, c – right frontolateral view; b – incomplete cephalon, CGS MM 557, internal mould, dorsal view; d – left free cheek with visual surface of eye, CGS MM 558, internal mould with remains of exoskeleton, lateral view; e – incomplete pygidium, CGS MM 559, dorsal view; f – incomplete pygidium, CGS MM 560, dorsal view. g – Chotecops? sp., incomplete corroded pygidium, CGS MM 561, dorsal view. h –?Lioharpes (Fritchaspis) sp., fragment of fringe, internal mould, lateral view, CGS MM 562. i – Crotalocephalus cf. affinis (HAWLE et CORDA, 1847), incomplete cranidium, CGS MM 563, internal mould, dorsal view. j–k – Acanthopyge (Acanthopyge) cf. haueri (BARRANDE, 1846), incomplete hypostomes, dorsal views, j – CGS MM 564, k – CGS MM 565. l – Acanthopyge? sp., fragment of cranidium, internal mould with remains of exoskeleton, CGS MM 566, dorsal view. m – Otarion cf. lacrimarum PŘIBYL et VANĚK, 1981, incomplete corroded cranidium, internal mould, CGS MM 567, dorsal view. n – Orbitoproetus cf. angelini (HAWLE et CORDA, 1847), juvenile pygidium, internal mould with exoskeleton, CGS MM 568, dorsal view. o – Eremiproetus? sp., negative counterpart of poorly preserved fragment of cranidium, CGS MM 569, dorsal view" pageId="2" pageNumber="81">Text-fig. 3d</figureCitation>
and younger unfigured specimen CGS MM 575), which fits well with description of
<taxonomicName id="DE87A392FFBAFFF8FE148138C702FBE6" baseAuthorityName="HAWLE et CORDA" baseAuthorityYear="1847" box="[443,633,1048,1072]" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="hoseri">
<emphasis id="2BF30403FFBAFFF8FE148138C702FBE6" box="[443,633,1048,1072]" italics="true" pageId="2" pageNumber="81">Chotecops hoseri</emphasis>
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by
<bibRefCitation id="7D16A5E0FFBAFFF8FD098138C582FB86" author="Chlupac, I." pageId="2" pageNumber="81" pagination="1 - 172" refId="ref7228" refString="Chlupac, I. (1977): The phacopid trilobites of the Silurian and Devonian of Czechoslovakia. - Rozpravy Ustred- niho ustavu geologickeho, 23: 1 - 172." type="journal article" year="1977">Chlupáč (1977: 53)</bibRefCitation>
. A lesser maximal number of ommatidia in a dorso-ventral row in the younger specimen can be explained by its early ontogenetic stage. In the largest specimen, the number of dorso-ventral rows in an eye is 17, and some are clearly missing (broken off). Following
<bibRefCitation id="7D16A5E0FFBAFFF8FDE38199C780FB07" author="Chlupac, I." box="[588,763,1209,1233]" pageId="2" pageNumber="81" pagination="1 - 172" refId="ref7228" refString="Chlupac, I. (1977): The phacopid trilobites of the Silurian and Devonian of Czechoslovakia. - Rozpravy Ustred- niho ustavu geologickeho, 23: 1 - 172." type="journal article" year="1977">Chlupáč (1977)</bibRefCitation>
, specimens of
<taxonomicName id="DE87A392FFBAFFF8FE8C81F9C49EFB27" baseAuthorityName="HAWLE et CORDA" baseAuthorityYear="1847" box="[291,485,1241,1265]" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="hoseri">
<emphasis id="2BF30403FFBAFFF8FE8C81F9C49EFB27" box="[291,485,1241,1265]" italics="true" pageId="2" pageNumber="81">Chotecops hoseri</emphasis>
</taxonomicName>
should reach 18 dorsoventral rows. Pygidia correspond with
<taxonomicName id="DE87A392FFBAFFF8FD8A81DAC7F2FAC7" baseAuthorityName="HAWLE et CORDA" baseAuthorityYear="1847" box="[549,649,1273,1297]" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="hoseri">
<emphasis id="2BF30403FFBAFFF8FD8A81DAC74CFAC7" box="[549,567,1274,1297]" italics="true" pageId="2" pageNumber="81">C</emphasis>
.
<emphasis id="2BF30403FFBAFFF8FDE981D9C7F2FAC7" box="[582,649,1273,1297]" italics="true" pageId="2" pageNumber="81">hoseri</emphasis>
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in number of ribs (4–5, 6 in large specimens only; see
<bibRefCitation id="7D16A5E0FFBAFFF8FDFD803AC792FAE4" author="Chlupac, I." box="[594,745,1306,1330]" pageId="2" pageNumber="81" pagination="1 - 172" refId="ref7228" refString="Chlupac, I. (1977): The phacopid trilobites of the Silurian and Devonian of Czechoslovakia. - Rozpravy Ustred- niho ustavu geologickeho, 23: 1 - 172." type="journal article" year="1977">Chlupáč 1977</bibRefCitation>
)), by their PL/PW ratios, and by prominent effacement of pleural furrows, except for the first one (two) pairs. The exact number of axial rings in studied specimens remains unknown because of poor preservation, but certainly exceeds four well-expressed rings. The studied remains differ from
<taxonomicName id="DE87A392FFBAFFF8FF2F80FBC489FA25" authority="CHLUPAC, 1971" authorityName="CHLUPAC" authorityYear="1971" box="[128,498,1499,1523]" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="auspex">
<emphasis id="2BF30403FFBAFFF8FF2F80FBC43FFA25" box="[128,324,1499,1523]" italics="true" pageId="2" pageNumber="81">Chotecops auspex</emphasis>
<bibRefCitation id="7D16A5E0FFBAFFF8FEE180FBC489FA25" author="Chlupac, I." box="[334,498,1499,1523]" pageId="2" pageNumber="81" pagination="255 - 262" refId="ref7198" refString="Chlupac, I. (1971): New phacopid trilobites from the Devonian of Czechoslovakia. - Casopis pro mineralogii a geologii, 16: 255 - 262." type="journal article" year="1971">CHLUPÁČ, 1971</bibRefCitation>
</taxonomicName>
, also rarely occurring in the
<taxonomicName id="DE87A392FFBAFFF8FF0980DBC4D7F9C5" authority="Limestone" authorityName="Limestone (Chlupac" authorityYear="1983" box="[166,428,1531,1555]" class="Trilobita" family="Lichidae" genus="Acanthopyge" kingdom="Animalia" order="Lichida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="genus">Acanthopyge Limestone</taxonomicName>
(
<bibRefCitation id="7D16A5E0FFBAFFF8FE1680DBC729F9C5" author="Chlupac, I." box="[441,594,1531,1555]" pageId="2" pageNumber="81" pagination="255 - 262" refId="ref7198" refString="Chlupac, I. (1971): New phacopid trilobites from the Devonian of Czechoslovakia. - Casopis pro mineralogii a geologii, 16: 255 - 262." type="journal article" year="1971">Chlupáč 1971</bibRefCitation>
,
<bibRefCitation id="7D16A5E0FFBAFFF8FDF280DBC7EEF9C5" author="Chlupac, I." box="[605,661,1531,1555]" pageId="2" pageNumber="81" pagination="1 - 172" refId="ref7228" refString="Chlupac, I. (1977): The phacopid trilobites of the Silurian and Devonian of Czechoslovakia. - Rozpravy Ustred- niho ustavu geologickeho, 23: 1 - 172." type="journal article" year="1977">1977</bibRefCitation>
, 1983), in having slightly larger eyes with more dorsoventral files of ommatidia, and by better expressed pygidial segmentation (much effaced in
<taxonomicName id="DE87A392FFBAFFF8FEEE837CC4D7F9A5" authorityName="CHLUPAC" authorityYear="1971" box="[321,428,1628,1651]" class="Trilobita" family="Phacopidae" genus="Chotecops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="81" phylum="Arthropoda" rank="species" species="auspex">
<emphasis id="2BF30403FFBAFFF8FEEE837CC428F9A5" box="[321,339,1628,1651]" italics="true" pageId="2" pageNumber="81">C</emphasis>
.
<emphasis id="2BF30403FFBAFFF8FECD837CC4D7F9A5" box="[354,428,1628,1651]" italics="true" pageId="2" pageNumber="81">auspex</emphasis>
</taxonomicName>
). However, the observed angle of divergence in cephalic axial furrows fits better to the latter-mentioned species.
</paragraph>
</subSubSection>
</treatment>
</document>
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