234 lines
27 KiB
XML
234 lines
27 KiB
XML
<document id="7C9C8288C9B5FF6EB8A4751C6860CEC0" ID-CLB-Dataset="298186" ID-DOI="10.3897/zookeys.1204.114932" ID-GBIF-Dataset="33ca18a1-99c6-4b56-9a8e-92fe6f6f9209" ID-ZooBank="F5134CC6-A83C-48B6-A1EA-235E6858D5CB" ID-publisher-id="114932" URI-arpha="EAF89A3A-8354-506D-AF9B-9E1C53605541" URI-zoobank="http://zoobank.org/F5134CC6-A83C-48B6-A1EA-235E6858D5CB" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1717680124246" checkinUser="pensoft" docAuthor="Nascimento, Elynton Alves & Bocakova, Milada" docDate="2024" docId="2C643453BC6F545EA4FB74E4DD6E086A" docLanguage="en" docName="ZooKeys 1204: 241-259" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/114932/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Gorhamium bidentatum Nascimento & Bocakova 2024, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="259" masterDocId="EAF89A3A8354506DAF9B9E1C53605541" masterDocTitle="Phylogenetic analysis reveals a new net-winged beetle genus of Eurrhacini (Coleoptera, Lycidae) from the Pacific slopes of Central America and Ecuador" masterLastPageNumber="259" masterPageNumber="241" pageNumber="241" updateTime="1717709099581" updateUser="ExternalLinkService">
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<mods:mods id="7E591E104DCB5EF4B7BB92984262FB70" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="487AAF010216BE02D10F94E516828FBC">
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<mods:title id="49448037E46DB56B39E30743646C1FB2">Phylogenetic analysis reveals a new net-winged beetle genus of Eurrhacini (Coleoptera, Lycidae) from the Pacific slopes of Central America and Ecuador</mods:title>
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</mods:titleInfo>
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<mods:name id="935AFA5B4EF5A1752447AF524F6C48AA" type="personal">
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<mods:roleTerm id="73E44D54927AB2620CE59DBA8D4B2BB7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F89EC004BE17D7E7BAEF3B406300EEFA">Nascimento, Elynton Alves</mods:namePart>
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<mods:nameIdentifier id="368D661C20996FE9BE37333AD1258601" type="ORCID">0000-0002-9071-2823</mods:nameIdentifier>
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<mods:affiliation id="7D440C534ED94775F3B9C1FD238352F1">Departamento de Engenharia Ambiental, Universidade Estadual do Centro-Oeste, Rua Professora Maria Roza Zanon de Almeida, s / n, Engenheiro Gutierrez, Irati – PR, CEP 84505 - 677, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="0089049F821DB2D7F90DE5DF3C76C029" type="personal">
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<mods:roleTerm id="AD9743D42A1E701C1D8932E9F53E079A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BD0E83D88E60904C6E21C9BF143D9E5A">Bocakova, Milada</mods:namePart>
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<mods:nameIdentifier id="EBF82730B6CD4608AA7BD57076AD2AE4" type="ORCID">0000-0002-2507-0887</mods:nameIdentifier>
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<mods:affiliation id="03AC945CE8DC308F1BB694F29F2E84DC">Department of Biology, Faculty of Education, Palacky University, Purkrabska 2, CZ- 77140 Olomouc, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="8B9EAD3F3A930D486F19306C9B5E720C">text</mods:typeOfResource>
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<mods:titleInfo id="358FF8604A84E6A5F43C8D24EE277BF0">
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<mods:title id="2AD92DEC1A9D91F981CC38031AEE8782">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="00795EA7CF247E91E20C0EFC0E79699E">
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<mods:date id="86A4F3DAE500EBB1BF4FC7B36EAD1F3B">2024</mods:date>
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<mods:detail id="978071DBD4877F6F671B7DAD355FE0BF" type="pubDate">
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<mods:number id="F41DCB7B4A8CE6FDFD500F9777F646A5">2024-06-06</mods:number>
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</mods:detail>
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<mods:detail id="E83850A8D6B76B9DE5FF46DC2803ADFA" type="volume">
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<mods:number id="9551CF168BC0E5C6851ACA7839040D84">1204</mods:number>
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<mods:start id="D0ACCB6479C425C842E34101BD6E3770">241</mods:start>
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<mods:end id="67B35AF24A7C4221C631B23F982FF6DF">259</mods:end>
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<mods:classification id="74DECDEDF092FB6E255AE22B238041A2">journal article</mods:classification>
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<mods:identifier id="24A3727FE48EF46D092DD738A5B8B8B0" type="CLB-Dataset">298186</mods:identifier>
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<mods:identifier id="D56B50115AEAB3EDCD331E96F8B0C050" type="DOI">10.3897/zookeys.1204.114932</mods:identifier>
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<mods:identifier id="960D600D1B91BC7321A97328D32611D3" type="GBIF-Dataset">33ca18a1-99c6-4b56-9a8e-92fe6f6f9209</mods:identifier>
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<treatment id="2C643453BC6F545EA4FB74E4DD6E086A" ID-DOI="http://doi.org/10.5281/zenodo.11505169" ID-GBIF-Taxon="232157813" ID-Zenodo-Dep="11505169" ID-arpha="2C643453-BC6F-545E-A4FB-74E4DD6E086A" ID-zoobank="https://zoobank.org/C1E7CBDC-5EBC-457E-BB8D-5B51E6ACE0F6" LSID="urn:lsid:plazi:treatment:2C643453BC6F545EA4FB74E4DD6E086A" LSID-ZBK="urn:lsid:zoobank.org:act:C1E7CBDC-5EBC-457E-BB8D-5B51E6ACE0F6" httpUri="http://treatment.plazi.org/id/2C643453BC6F545EA4FB74E4DD6E086A">
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<subSubSection id="2968F653D01C5D21FA456279914828C6" type="nomenclature">
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||
<paragraph id="503024EDC525B73CA1A47DD577973171">
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||
<taxonomicName id="EC117131948535A405874A3DBE0AC66A" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
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<emphasis id="FA185058EA12D89C229AA824ADB05F4E" italics="true">Gorhamium bidentatum</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="C4B0DC54539185AFEAC3C708FC8F9221" rank="species">sp. nov.</taxonomicNameLabel>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="F8A2471D68ABA18C3B521F460285B22E" type="description">
|
||
<paragraph id="DAE5C08AC3704C477B827FB6F579F795">
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||
<figureCitation id="62314F0E4EC6D6FBCD2BFFD967AD1DE1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">Figs 2 A, B</figureCitation>
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||
,
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<figureCitation id="3F4BB262E11094B86A296A21EC949F81" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pronotum A Gorhamium bidentatum sp. nov., male B Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure3" httpUri="https://binary.pensoft.net/fig/1063777">3 A</figureCitation>
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||
,
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<figureCitation id="13886317C16F0C154EC84B4710B5D050" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 A, B</figureCitation>
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||
,
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<figureCitation id="20FD64040521255CA2FD204A69D93B95" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 A – D</figureCitation>
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||
,
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<figureCitation id="0E61E437B306F143B674CCA2D5CAB7BE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C – E, H</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0ER4AG" type="type material">
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||
<paragraph id="2D7AE5C94010D6811B80A54B18556B90">
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||
<heading id="6E0DCD8E360D4AFD31A9030F45909F72" reason="title">Type material.</heading>
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||
</paragraph>
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||
<paragraph id="3BD632FD828DAB70F8763AAF65849703">
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||
<materialsCitation id="8B2274378744FB07EA1DB90D5F763103" ID-GBIF-Occurrence="4873767301" collectionCode="V" collectorName="B. C. A. Col." country="Panama" elevation="613" location="Champion" specimenCount="1" specimenCount-male="1" stateProvince="V. de Chiriqui" typeStatus="holotype">
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||
<emphasis id="D93DEECE94D88AE49B41DD5C7DCE1636" bold="true" italics="true">
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||
<typeStatus id="EC68D0763E683AC32D8A18E36FDCCE08" type="holotype">Holotype</typeStatus>
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</emphasis>
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•
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<specimenCount id="0CBF782EDBCED1F91F874331AA9BFBE3" count="1" type="male">male</specimenCount>
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, “
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<collectingCountry id="4830DE5E18A5412B557B847C5805BCF3" name="Panama">Panama</collectingCountry>
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,
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||
<collectingRegion id="6318A0FCD6BA09DA262FE13C479A2DE2">V. de Chiriqui</collectingRegion>
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||
,
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||
<quantity id="1B971418581D8EBD99960206D74E0125" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">
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||
<elevation id="95A91965C22DD941B5BC7DBA41B34273" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">25–4000 ft.</elevation>
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||
</quantity>
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||
<location id="04EBA913FA0F112BBF273D5074154F0E" LSID="urn:lsid:plazi:treatment:2C643453BC6F545EA4FB74E4DD6E086A:04EBA913FA0F112BBF273D5074154F0E" country="Panama" name="Champion" stateProvince="V. de Chiriqui">Champion</location>
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||
”, secondary labels -
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<collectorName id="3BD3842372247A1A5FED03F6659BFBCD">B. C. A. Col.</collectorName>
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||
III. (2).
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||
<taxonomicName id="E888481EFC4350B928D139507CD0692A" authorityName="Gorham" authorityYear="1884" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chiriquense">
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||
<emphasis id="A5DA3468A7429CCF733B809A33A21532" italics="true">
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||
<originalDetermination id="1A927D5ECB78C62C7AE1618373514E11">Calocladon chiriquense</originalDetermination>
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||
</emphasis>
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||
</taxonomicName>
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||
</materialsCitation>
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||
,
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||
<typeStatus id="E115881A9EEABC6B2E728410A1196D76" type="syntype">SYNTYPE</typeStatus>
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||
- blue-edged circle (
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<collectionCode id="BAC4C9F237E9C3FD5C7F9032D8A5A436">BMNH</collectionCode>
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||
). [Volcan de
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<collectingRegion id="95537E5AF296FFFD8A845708A9A17035" country="Panama" name="Chiriqui">Chiriqui</collectingRegion>
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is now referred to as Volcán Barú].
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||
</paragraph>
|
||
<paragraph id="D841D5375A4BED0CA913BEC9755DCAED">
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||
<materialsCitation id="802D290CC50582DB86FE7D7B7FB02555" ID-GBIF-Occurrence="4873767304" collectionCode="BMNH" collectorName="B. C. A. Col." country="Panama" elevation="613" location="Panama" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
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||
<emphasis id="98B0A1C20F8D9F61B72BF1454482893D" bold="true" italics="true">
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||
<typeStatus id="A4E5C8AFD36633CF1E97F94FBD133965" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
•
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||
<collectingRegion id="84FA39328A869B9650BC0D24461B481D" country="Panama" name="Panama">Panama</collectingRegion>
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||
, same data as for holotype,
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<specimenCount id="CAE31F6316C25ABE22D4FF0E6478E742" count="1" type="male">1 male</specimenCount>
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||
,
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||
<specimenCount id="0CEB85D106D35618828A01AE5D69C123" count="3" type="female">3 females</specimenCount>
|
||
(
|
||
<collectionCode id="A73BBF27B984A8B8C6DA47C6CF648F79">BMNH</collectionCode>
|
||
)
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||
</materialsCitation>
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||
; •
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||
<materialsCitation id="F1ADCDA0E2A022551C6FE96FC9F38B57" ID-GBIF-Occurrence="4873767302" collectionCode="V" collectorName="B. C. A. Col." country="Panama" elevation="458" location="Champion" specimenCount="1" stateProvince="V. de Chiriqui" typeStatus="paratype">
|
||
“
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||
<collectingCountry id="03BE56561E2E4DE91A32F6C2D439135C" name="Panama">PANAMA</collectingCountry>
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||
,
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||
<collectingRegion id="4D969376B1F702CCB81D7EFC7B28532D">V. de Chiriqui</collectingRegion>
|
||
,
|
||
<quantity id="AF8914E5603AF975097F75B705558232" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">
|
||
<elevation id="764089CF46325410DF84BC3C1366FA2E" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">2-3000 ft.</elevation>
|
||
</quantity>
|
||
<location id="360EF646142FFA1B587EEDD31373CB29" LSID="urn:lsid:plazi:treatment:2C643453BC6F545EA4FB74E4DD6E086A:360EF646142FFA1B587EEDD31373CB29" country="Panama" name="Champion" stateProvince="V. de Chiriqui">Champion</location>
|
||
”, secondary labels - same data as for holotype
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="7EA2401C03431C72DAD05A4AD5C60CAC" ID-GBIF-Occurrence="4873767305" collectionCode="BMNH" country="Panama" specimenCount="1" specimenCount-female="1" stateProvince="Chiriqui" typeStatus="syntype">
|
||
<typeStatus id="3EA815CDAF1E8E0518E4A021A800BEE9" type="syntype">SYNTYPE</typeStatus>
|
||
- blue-edged circle,
|
||
<specimenCount id="6D6E6F6763FB50C2DFD5E869E84FB3B2" count="1" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="686D14070553B1C90595F533FBCB3DB0">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; • “
|
||
<materialsCitation id="028004CECEFEF7ACF17E64E3DA21AC49" ID-GBIF-Occurrence="4873767303" collectionCode="V" collectorName="B. C. A. Col." country="Panama" elevation="1524" location="Champion" specimenCount="1" stateProvince="V. de Chiriqui" typeStatus="syntype">
|
||
<collectingCountry id="AE13E8F354CD6EC3B8147EC1621CDCC3">Panama</collectingCountry>
|
||
,
|
||
<collectingRegion id="F341ACF1C3D89B7D55AD1A75DCE0EEE2">V. de Chiriqui</collectingRegion>
|
||
,
|
||
<quantity id="094328BBBA5E87082853AA4D6929A0DF" metricMagnitude="3" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" unit="ft" value="5000.0" valueMax="6000.0" valueMin="4000.0">
|
||
<elevation id="3062455D6633C9E2ECBB438FAE69545F" metricMagnitude="3" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" unit="ft" value="5000.0" valueMax="6000.0" valueMin="4000.0">4000–6000 ft.</elevation>
|
||
</quantity>
|
||
<location id="150808D9BD4F4ED1B1ED0F315F0FF2B5" LSID="urn:lsid:plazi:treatment:2C643453BC6F545EA4FB74E4DD6E086A:150808D9BD4F4ED1B1ED0F315F0FF2B5" country="Panama" name="Champion" stateProvince="V. de Chiriqui">Champion</location>
|
||
, secondary labels - same data as for holotype
|
||
</materialsCitation>
|
||
,
|
||
<typeStatus id="A2146CFB526E84DDEB17CFB06C211E57" type="syntype">SYNTYPE</typeStatus>
|
||
- blue-edged circle,
|
||
<specimenCount id="3ED200FBBEF973E59C3D301F7F0C7D5C" count="3" type="male">3 males</specimenCount>
|
||
(
|
||
<collectionCode id="4F1DA60D46C5C0C6ED562E020B167D4A">BMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EN5AG" type="diagnosis">
|
||
<paragraph id="44E7643011B6C3FD82E691F8AD9823B0">
|
||
<heading id="B4EF4758EDE48AE7737EAF34808C7809" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="70A35429C95AAC9646AD81C1B6AB0D95">
|
||
Pronotum and elytra largely yellow, only median longitudinal stripe on pronotum, basal half of elytral suture, and elytral apex black. Phallus rod-like apically, ventromedial opening oval, widest medially. Parameres shorter than half of phallus, their ventrobasal projects separated (d
|
||
<subScript id="5AF42E0204377337644AD8CD35A42B05">1</subScript>
|
||
, Fig.
|
||
<figureCitation id="2A3DFC4E1B21BA660123A01EBB5317C3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C</figureCitation>
|
||
), apex of parameres bidentate, internal sac largely membranous, micro spurs barely visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZ5AG" type="description">
|
||
<paragraph id="3BDB62B16C009B95827F012D15D064F7">
|
||
<heading id="1B7E629456C555EB2EA73E337997F9B0" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="A6F9E76B304C9E6B7B2BA6FF857BB264">
|
||
Body length:
|
||
<quantity id="42BD7CDE903C92D5534B84EB0D968CFE" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.0" metricValueMin="5.1" unit="mm" value="5.55" valueMax="6.0" valueMin="5.1">5.1–6 mm</quantity>
|
||
, width across the humeri:
|
||
<quantity id="2E7D2BF851AE89C614E8B2B2AE19414D" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
|
||
. Body dark brown, only anterior pronotal margin, broad sides of pronotum, trochanters, bases of femora, scutellum, and most of elytra yellow (Fig.
|
||
<figureCitation id="9F13A801D9C2F2F9B93FB2DBB61FFA59" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 A, B</figureCitation>
|
||
). Sutural stripe in basal half of elytra and apical 1 / 30 of elytra black. Head largely covered by pronotum. Elytra 4–4.8 × longer than humeral width (Fig.
|
||
<figureCitation id="2E6DC6610CC70CFE058FC9184132C4F7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 A</figureCitation>
|
||
). Primary costae 2 and 4 and basal quarter of primary costa 3 more elevated. Reticulate cells irregular, secondary costae 3 and 4 present only basally.
|
||
</paragraph>
|
||
<paragraph id="0DFB3FCBEC81F1193DAE5C4D0F64424D">
|
||
<emphasis id="C6D2C7A3B5A8807B3D1E4D9A2D17C48E" bold="true">Male.</emphasis>
|
||
Eyes large, hemispherically prominent, eye diameter 1.5–1.7 × longer than interocular distance. Antennae with antennomere 3 (= A 3) 1.15 × shorter than A 4, A 4 – A 10 subequal in length; antennal branches flattened, lamella of A 3 1.7 × longer than antennomere length, remaining lamellae considerably longer, ~ 2.4 × longer than antennomere length (Fig.
|
||
<figureCitation id="4FB7513ADB662EBB3A327441EA95D575" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 B</figureCitation>
|
||
). Abdominal sternum VIII with a broad, shallow emargination distally (up to 1 / 5 of sternum length), its proximal margin minutely emarginated up to 1 / 10 of sternum length (Fig.
|
||
<figureCitation id="0F7887223AC0981BA84E7DD735A0A31E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 C</figureCitation>
|
||
). Tergum X small, only 1.3 × longer than preceding sternum IX on the sides (Fig.
|
||
<figureCitation id="BC1DA8A0E2156644D0E79D8B77678E96" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 B</figureCitation>
|
||
). Phallus rod-like in distal 1 / 3, slightly widened apically, with a dorsal hook in median portion (c, Fig.
|
||
<figureCitation id="6770BAB5D9D0726BE791099D05547D9E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 D</figureCitation>
|
||
) and large ventral opening widest medially (b, Fig.
|
||
<figureCitation id="4617D878FBB8D7D554D70E2E5BC27849" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C</figureCitation>
|
||
). Parameres moderately shorter than half of phallus, each with two coarse teeth apically, ventrobasal parameral protrusions slender, medially separated by 1 / 3 of phallic width (Fig.
|
||
<figureCitation id="9329F6A7F2F3D024B93B0B99210DE65E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0077B820917483B96A48AF179DC20B1F">
|
||
<emphasis id="6AA1696E000AD583B130A0B9DA7811EC" bold="true">Female.</emphasis>
|
||
Eyes small, eye diameter 1.3 × shorter than interocular distance. Antennae serrate (Fig.
|
||
<figureCitation id="26780DE4A1EDFF4CBE584DED49F5F9D0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 B</figureCitation>
|
||
). Terminal sternum with spiculum gastrale rudimentary, triangular to slightly pointed (Fig.
|
||
<figureCitation id="9B65B63639CB020266516CF6207AD28D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 D</figureCitation>
|
||
). Ovipositor elongate (Fig.
|
||
<figureCitation id="DB474D96BF1EEC9C91BDC50531EED513" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 H</figureCitation>
|
||
), valvifers rod-like, 1.4 × longer than coxites, basally coalescent. Coxites medially distant, their base and apex closer, styli as long as half of coxites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ETABG" type="etymology">
|
||
<paragraph id="C5570112AC366D7EA45A37A19812D36F">
|
||
<heading id="7FC2715B883D7C3579A71532FE44FBD2" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="480DB9C178D8CEF5DEF68A9C9573F5BC">Named after the shape of apical portion of parameres.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EYABG" type="distribution">
|
||
<paragraph id="89B018AFA34D6EBB8CD68D72C0E72C2B">
|
||
<heading id="C525E6117ADD3E0D9B029F428044913B" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="9FBF99A11B15A275B5CACDE5040D5FD2">
|
||
<collectingCountry id="F14D002B3361E29C2F92D0F7B137EB84" name="Panama">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |