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<mods:title id="23F30272E46DEAB61123C6BB2CBEC9C4">Revision of the Seashore-dwelling Subgenera Emplenota Casey and Triochara Bernhauer (Coleoptera: Staphylinidae: genus Aleochara) from Japan</mods:title>
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<treatment id="03DE87D80828FFF5FF28FF612C5038D0" ID-DOI="http://doi.org/10.5281/zenodo.6175364" ID-GBIF-Taxon="119528583" ID-Zenodo-Dep="6175364" LSID="urn:lsid:plazi:treatment:03DE87D80828FFF5FF28FF612C5038D0" httpUri="http://treatment.plazi.org/id/03DE87D80828FFF5FF28FF612C5038D0" lastPageNumber="5" pageId="4" pageNumber="5">
<subSubSection id="C36D65450828FFF5FF28FF612A593BA0" box="[151,604,151,177]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BC836CE0828FFF5FF28FF612A593BA0" blockId="4.[151,604,151,177]" box="[151,604,151,177]" pageId="4" pageNumber="5">
<heading id="D08081A20828FFF5FF28FF612A593BA0" bold="true" box="[151,604,151,177]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B903EADC0828FFF5FF28FF612A593BA0" bold="true" box="[151,604,151,177]" pageId="4" pageNumber="5">
Genus
<taxonomicName id="4C774D4D0828FFF5FF51FF612A593BA0" ID-CoL="8VRG3" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" box="[238,604,151,177]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0828FFF5FF51FF61296E3BA0" bold="true" box="[238,363,151,177]" italics="true" pageId="4" pageNumber="5">Aleochara</emphasis>
<bibRefCitation id="EFE64B3F0828FFF5FECCFF612A593BA0" author="Gravenhorst" box="[371,604,151,177]" pageId="4" pageNumber="5" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. Brunsuigae, Carolus Reichard, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36D65450828FFF5FF28FF292D993A28" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8BC836CE0828FFF5FF28FF292CBD3BE4" blockId="4.[151,1208,222,245]" box="[151,1208,222,245]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB6711E00828FFF5FF28FF292CBD3BE4" box="[151,1208,222,245]" pageId="4" pageNumber="5">
<taxonomicName id="4C774D4D0828FFF5FF28FF2929EC3BE5" ID-CoL="8VRG3" authority="Gravenhorst, 1802: 67" authorityName="Gravenhorst" authorityPageNumber="67" authorityYear="1802" box="[151,489,222,245]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0828FFF5FF28FF2929053BE4" box="[151,256,223,245]" italics="true" pageId="4" pageNumber="5">Aleochara</emphasis>
<treatmentCitation id="0AD610DF0828FFF5FEB9FF2829EC3BE5" author="Gravenhorst" box="[262,489,222,244]" page="67" pageId="4" pageNumber="5" year="1802">
<bibRefCitation id="EFE64B3F0828FFF5FEB9FF2829C33BE5" author="Gravenhorst" box="[262,454,222,244]" pageId="4" pageNumber="5" refString="Gravenhorst, J. L. C. (1802) Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera familias et species distribuit. Brunsuigae, Carolus Reichard, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
: 67
</treatmentCitation>
</taxonomicName>
(original description;
<typeStatus id="54CC886C0828FFF5FD74FF292AF23BE5" box="[715,759,223,244]" pageId="4" pageNumber="5">type</typeStatus>
species:
<taxonomicName id="4C774D4D0828FFF5FCEBFF292CA83BE4" authority="Goeze, 1777" authorityName="Goeze" authorityYear="1777" box="[852,1197,223,245]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B903EADC0828FFF5FCEBFF292C273BE4" box="[852,1058,223,245]" italics="true" pageId="4" pageNumber="5">Staphylinus curtulus</emphasis>
Goeze, 1777
</taxonomicName>
).
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</paragraph>
<paragraph id="8BC836CE0828FFF5FF28FED72D993A28" blockId="4.[151,1437,289,961]" box="[151,1436,289,313]" pageId="4" pageNumber="5">
See,
<bibRefCitation id="EFE64B3F0828FFF5FF72FED729BA3A28" author="Klimaszewski" box="[205,447,289,313]" pageId="4" pageNumber="5" refString="Klimaszewski, J. (1984) A revision of the genus Aleochara Gravenhorst of America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada, 129, 1 - 211." type="journal article" year="1984">Klimaszewski (1984)</bibRefCitation>
,
<bibRefCitation id="EFE64B3F0828FFF5FE74FED72A783A28" author="Smetana" box="[459,637,289,313]" pageId="4" pageNumber="5" refString="Smetana, A. (2004) Aleocharinae. In: Lobl, I &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 2. Apollo Books, Stenstrup, Denmark, pp. 353 - 494." type="book" year="2004">Smetana (2004)</bibRefCitation>
and
<bibRefCitation id="EFE64B3F0828FFF5FD0CFED72C0B3A28" author="Gouix" box="[691,1038,289,313]" pageId="4" pageNumber="5" refString="Gouix, N. &amp; Klimaszewski, J. (2007) Catalogue of Aleocharine Rove Beetles of Canada and Alaska (Coleoptera, Staphylinidae, Aleocharinae). Pensoft Publishers, Sofia-Moscow, 165 pp." type="book" year="2007">Gouix &amp; Klimaszewski (2007)</bibRefCitation>
for further synonymic information.
</paragraph>
</subSubSection>
<subSubSection id="C36D65450828FFF5FF78FEB32A6639D4" pageId="4" pageNumber="5" type="description">
<paragraph id="8BC836CE0828FFF5FF78FEB32A6639D4" blockId="4.[151,1437,289,961]" pageId="4" pageNumber="5">
<emphasis id="B903EADC0828FFF5FF78FEB329BF3A4C" bold="true" box="[199,442,325,350]" pageId="4" pageNumber="5">Redescription. Body</emphasis>
: small to large, medium to large in many cases; overall shape diverse, normally more or less broad: compact and robust, more or less fusiform or cylindrical and rarely slender and narrow (Ashe &amp; Maus, 1998).
<emphasis id="B903EADC0828FFF5FF55FE7B293B3AB7" bold="true" box="[234,318,397,422]" pageId="4" pageNumber="5">Colour</emphasis>
: uniformly reddish brown to black in most cases, sometimes elytra lighter or with maculations.
<emphasis id="B903EADC0828FFF5FF28FE4728D23AD8" bold="true" box="[151,215,433,457]" pageId="4" pageNumber="5">Head</emphasis>
: head capsule somewhat circular to more or less transverse.
<emphasis id="B903EADC0828FFF5FC1CFE462C133AD8" bold="true" box="[931,1046,432,457]" pageId="4" pageNumber="5">Antennae</emphasis>
: 11-segmented, scape somewhat clavated, normally longer or as long as pedicel (
<bibRefCitation id="EFE64B3F0828FFF5FD79FE232BB63AFC" author="Klimaszewski" box="[710,947,469,493]" pageId="4" pageNumber="5" refString="Klimaszewski, J. (1984) A revision of the genus Aleochara Gravenhorst of America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada, 129, 1 - 211." type="journal article" year="1984">Klimaszewski, 1984</bibRefCitation>
).
<emphasis id="B903EADC0828FFF5FC76FE232C5D3AFC" bold="true" box="[969,1112,469,493]" pageId="4" pageNumber="5">Mouthparts</emphasis>
: lacinia wide, multispinose; galea wide, as long as lacinia (
<bibRefCitation id="EFE64B3F0828FFF5FDAFFE0F2B043900" author="Klimaszewski" box="[528,769,505,529]" pageId="4" pageNumber="5" refString="Klimaszewski, J. (1984) A revision of the genus Aleochara Gravenhorst of America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada, 129, 1 - 211." type="journal article" year="1984">Klimaszewski, 1984</bibRefCitation>
).
<emphasis id="B903EADC0828FFF5FCA5FE0F2B773900" bold="true" box="[794,882,505,529]" pageId="4" pageNumber="5">Thorax</emphasis>
: mesoventrite variable, without carina to with complete carina.
<emphasis id="B903EADC0828FFF5FED0FDEB29A23924" bold="true" box="[367,423,541,565]" pageId="4" pageNumber="5">Legs</emphasis>
: normally medium length; mesocoxae narrowly separated, with long process of mesoventrite.
<emphasis id="B903EADC0828FFF5FE89FDB629AE3948" bold="true" box="[310,427,576,601]" pageId="4" pageNumber="5">Abdomen</emphasis>
: simple and normally fusiform; tergite of segments IIIV impressed transversely at base.
<emphasis id="B903EADC0828FFF5FF28FD932900396F" bold="true" box="[151,261,613,638]" pageId="4" pageNumber="5">Genitalia</emphasis>
: median lobe of aedeagus in male, usually with flagellum inside. Spermatheca of female, which are differently modified in different subgenera (
<bibRefCitation id="EFE64B3F0828FFF5FD24FD7F2C0439B0" author="Klimaszewski" box="[667,1025,649,673]" pageId="4" pageNumber="5" refString="Klimaszewski, J. &amp; Jansen, R. E. (1993) Systematics, biology and distribution of Aleochara Gravenhorst from southern Africa. Part 1: subgenus Xenochara Mulsant and Rey (Coleoptera: Staphylinidae). Annals of the Transvaal Museum, 36, 53 - 107." type="journal article" year="1993">Klimaszewski &amp; Jansen, 1993</bibRefCitation>
), but with some exceptions. Detail descriptions are in
<bibRefCitation id="EFE64B3F0828FFF5FED4FD5B2A5B39D4" author="Klimaszewski" box="[363,606,685,709]" pageId="4" pageNumber="5" refString="Klimaszewski, J. (1984) A revision of the genus Aleochara Gravenhorst of America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada, 129, 1 - 211." type="journal article" year="1984">Klimaszewski (1984)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D65450828FFF5FF78FD272C5038D0" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BC836CE0828FFF5FF78FD272B443868" blockId="4.[151,1437,289,961]" pageId="4" pageNumber="5">
<emphasis id="B903EADC0828FFF5FF78FD27294A39FB" bold="true" box="[199,335,721,746]" pageId="4" pageNumber="5">Comments.</emphasis>
Adults predate upon eggs, larvae, and puparia of flies; most known larvae are internal parasitoids of cyclorrhaphous flies (
<bibRefCitation id="EFE64B3F0828FFF5FE18FD032AC2381C" author="Peschke" box="[423,711,757,781]" pageId="4" pageNumber="5" refString="Peschke, K. &amp; Fuldner, D. (1977) Ubersicht und neue Untersuchungen zur Lebensweise der parasitoiden Aleocharinae (Coleoptera, Staphylinidae). Zoologische Jahrbucher, 104, 242 - 262." type="journal article" year="1977">Peschke &amp; Fuldner, 1977</bibRefCitation>
;
<bibRefCitation id="EFE64B3F0828FFF5FD6DFD032B77381C" author="Seevers" box="[722,882,757,781]" pageId="4" pageNumber="5" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology, 71, 1 - 289." type="journal article" year="1978">Seevers, 1978</bibRefCitation>
;
<bibRefCitation id="EFE64B3F0828FFF5FCC1FD032C63381C" author="Klimaszewski" box="[894,1126,757,781]" pageId="4" pageNumber="5" refString="Klimaszewski, J. (1984) A revision of the genus Aleochara Gravenhorst of America north of Mexico (Coleoptera, Staphylinidae, Aleocharinae). Memoirs of the Entomological Society of Canada, 129, 1 - 211." type="journal article" year="1984">Klimaszewski, 1984</bibRefCitation>
; Maus
<emphasis id="B903EADC0828FFF5FB09FD002CE2381C" box="[1206,1255,757,781]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
., 1998ab, 2001;
<bibRefCitation id="EFE64B3F0828FFF5FF28FCEF294B3820" author="Yamazaki" box="[151,334,793,817]" pageId="4" pageNumber="5" refString="Yamazaki, K. (2008) Aleochara fucicola and A. trisulcata (Coleoptera, Staphylinidae) as parasitoids of a kelp fly, Fucellia apicalis (Diptera, Anthomyiidae). Elytra, 36 (1), 151 - 152." type="journal article" year="2008">Yamazaki, 2008</bibRefCitation>
,
<bibRefCitation id="EFE64B3F0828FFF5FEE6FCEF29933820" author="Yamazaki" box="[345,406,793,817]" pageId="4" pageNumber="5" refString="Yamazaki, K. (2012) Seasonal changes in seaweed deposition, seaweed fly abundance, and parasitism at the pupal stage along sandy beaches in central Japan. Entomological Science, 15, 28 - 32." type="journal article" year="2012">2012</bibRefCitation>
). Thus,
<taxonomicName id="4C774D4D0828FFF5FE51FCEF2A663820" box="[494,611,793,817]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0828FFF5FE51FCEF2A663820" box="[494,611,793,817]" italics="true" pageId="4" pageNumber="5">Aleochara</emphasis>
</taxonomicName>
species are considered economically important as host agents. In contrast to the accumulation of ecological knowledge for some particular species, the taxonomy at the subgeneric level and the evolution and phylogeny of this group are poorly known.
</paragraph>
<paragraph id="8BC836CE0828FFF5FF78FC732C5038D0" blockId="4.[151,1437,289,961]" pageId="4" pageNumber="5">
Studies of Japanese species of
<taxonomicName id="4C774D4D0828FFF5FD93FC732AA4388C" box="[556,673,901,925]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B903EADC0828FFF5FD93FC732AA4388C" box="[556,673,901,925]" italics="true" pageId="4" pageNumber="5">Aleochara</emphasis>
</taxonomicName>
are also still incomplete and only 22 species in 6 subgenera have been recorded. Revisions of each subgenus for the Japanese species are also required.
</paragraph>
</subSubSection>
</treatment>
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