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<document id="883210B2006A0F981267B7E95E03A4F7" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58516FFA45991FB7E500FFD7C" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) kolleri Silva, Louzada &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="488" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="487" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="607BD1EE8A632F4AA88FB19C477B4314">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="67F7245824FAD002277C16B8C8B51030">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="28FB6D702821504D7326A36F481CD13D">Louzada, Júlio</mods:namePart>
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<mods:namePart id="243EE9E2A4B3B6953500B584D02E30DC">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:date id="FAFD889BCFF10068D6C508755FAB322B">2015</mods:date>
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<treatment id="03DE87D58516FFA45991FB7E500FFD7C" ID-DOI="http://doi.org/10.5281/zenodo.6115756" ID-GBIF-Taxon="119598958" ID-Zenodo-Dep="6115756" LSID="urn:lsid:plazi:treatment:03DE87D58516FFA45991FB7E500FFD7C" httpUri="http://treatment.plazi.org/id/03DE87D58516FFA45991FB7E500FFD7C" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487">
<subSubSection id="C36D65488516FFA55991FB7E54FCFB10" pageId="36" pageNumber="487" type="nomenclature">
<paragraph id="8BC836C38516FFA55991FB7E579AFB76" blockId="36.[151,725,1159,1221]" box="[151,725,1159,1186]" pageId="36" pageNumber="487">
<heading id="D08081AF8516FFA55991FB7E579AFB76" bold="true" box="[151,725,1159,1186]" fontSize="11" level="1" pageId="36" pageNumber="487" reason="1">
<emphasis id="B903EAD18516FFA55991FB7E579AFB76" bold="true" box="[151,725,1159,1186]" pageId="36" pageNumber="487">
<taxonomicName id="4C774D408516FFA55991FB7E5772FB75" ID-CoL="6CJ5H" box="[151,573,1159,1186]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="kolleri" status="sp. nov." subGenus="Aganhyboma">
<emphasis id="B903EAD18516FFA55991FB7E5462FB75" bold="true" box="[151,301,1159,1185]" italics="true" pageId="36" pageNumber="487">Deltochilum</emphasis>
(
<emphasis id="B903EAD18516FFA5583BFB7E5491FB75" bold="true" box="[317,478,1159,1185]" italics="true" pageId="36" pageNumber="487">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18516FFA558F6FB7E5772FB75" bold="true" box="[496,573,1159,1185]" italics="true" pageId="36" pageNumber="487">kolleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057AA8516FFA55B43FB71579AFB76" box="[581,725,1160,1186]" pageId="36" pageNumber="487" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38516FFA55991FB5554FCFB10" blockId="36.[151,725,1159,1221]" box="[151,435,1196,1221]" pageId="36" pageNumber="487">
(
<figureCitation id="134C2A468516FFA55999FB5555BAFB10" box="[159,245,1196,1221]" captionStart="FIGURES 12 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="36" pageNumber="487">Figs 19</figureCitation>
, 49, 56, 94, 122)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488516FFA55991FB0D5068FAD9" box="[151,1319,1268,1293]" pageId="36" pageNumber="487" type="etymology">
<paragraph id="8BC836C38516FFA55991FB0D5068FAD9" blockId="36.[151,1437,1268,1977]" box="[151,1319,1268,1293]" pageId="36" pageNumber="487">
<emphasis id="B903EAD18516FFA55991FB0D5451FAD9" bold="true" box="[151,286,1268,1293]" pageId="36" pageNumber="487">Etymology.</emphasis>
Named in honour of Wilson Werner Koller, the collecter of one of the specimens examined.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488516FFA559C1FAEE5631F86D" pageId="36" pageNumber="487" type="description">
<paragraph id="8BC836C38516FFA559C1FAEE5631F86D" blockId="36.[151,1437,1268,1977]" pageId="36" pageNumber="487">
<emphasis id="B903EAD18516FFA559C1FAEE54EBFAE4" bold="true" box="[199,420,1303,1328]" pageId="36" pageNumber="487">Description. Body.</emphasis>
Head, pronotum, elytra and ventral surface dark brown or black, with a greenish metallic sheen. Length&gt;
<quantity id="4C8F9B268516FFA55858FAC554FAFA80" box="[350,437,1340,1364]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="36" pageNumber="487" unit="mm" value="13.0">13 mm</quantity>
.
<emphasis id="B903EAD18516FFA558C2FAC55744FA81" bold="true" box="[452,523,1340,1365]" pageId="36" pageNumber="487">Head.</emphasis>
Foveiform punctures dense, touching on dorsal interocular surface. Punctures around clypeus shallower, smaller and more dispersed than those on the interocular surface. Dorsal interocular distance approximately four times eye width. Lateral margin of gena strongly curved outward, with row of setae around the eyes.
<emphasis id="B903EAD18516FFA55855FA5E54FCFA14" bold="true" box="[339,435,1447,1472]" pageId="36" pageNumber="487">Thorax.</emphasis>
Surface of pronotum with dense and deep foveiform punctures, touching on the sides and anterior angles. Discal punctures smaller and more dispersed than those of surrounding surface. Posteromedial punctures usually separated by less than one diameter. Margin between lateral and posterior angles straight.
<emphasis id="B903EAD18516FFA55C4FFA1650D3F9DC" bold="true" box="[1353,1436,1519,1544]" pageId="36" pageNumber="487">Elytra.</emphasis>
Lateral margins almost pararell (
<figureCitation id="134C2A468516FFA55B12F9ED5737F9F8" box="[532,632,1556,1581]" captionStart="FIGURES 119 123" captionStartId="51.[151,264,1596,1619]" captionTargetBox="[178,1436,193,1570]" captionTargetId="figure@51.[150,1436,193,1576]" captionTargetPageId="51" captionText="FIGURES 119 123. Species of D. (Aganhyboma). 119. D. ritamourae new species. 120. D. icariforme. 121. D. paresi new species. 122. D. kolleri new species. 123. D. subrubrum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244810/files/figure.png" pageId="36" pageNumber="487">Fig. 122</figureCitation>
) and surface slightly irregular (rugose). Elytral striae wider, carinate margins clearly separated (as in Fig. 75). Interstriae clearly convex, with its central portion nearly flat. Interstrial surface with foveiform punctures surrounded by microtubercles weakly defined, conspicuous on lateral of elytra. Interstriae usually more punctate on the sides, close to striae. Foveiform punctures of third interstria denser laterally (punctures can be so scattered at center of interstria to produce smooth midlongitudinal strip) (as in Fig. 86). Foveiform puncture surrounded by approximately six microtubercles. Basal carina of seventh interstria elongate, length about four or more times seventh interestria width. Basal carina of ninth interstria present in threequarters of elytra (not reaching anterior limit of apical carina of seventh interstria). Apical carina of third interstria rounded, better defined as tubercle, more prominent than apical carina of second and fourth interstriae. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria less than or equal to the sum of lengths of apical carina of fifth and sixth interstriae. Pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63).
</paragraph>
</subSubSection>
<subSubSection id="C36D65488517FFA459C1FF6E50DDFEB1" pageId="37" pageNumber="488" type="materials_examined">
<paragraph id="8BC836C38517FFA459C1FF6E50D9FF64" blockId="37.[151,1437,151,680]" box="[199,1430,151,176]" pageId="37" pageNumber="488">
<emphasis id="B903EAD18517FFA459C1FF6E57CCFF64" bold="true" box="[199,643,151,176]" pageId="37" pageNumber="488">
Material studied.
<typeStatus id="54CC88618517FFA4589DFF6E5743FF64" box="[411,524,151,176]" pageId="37" pageNumber="488" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F36076538517FFA45B1EFF6E57CCFF64" box="[536,643,151,176]" name="Brazil" pageId="37" pageNumber="488">BRAZIL</collectingCountry>
</emphasis>
:
<collectionCode id="ED66AE068517FFA45B94FF6157AEFF64" box="[658,737,152,176]" pageId="37" pageNumber="488">MATO</collectionCode>
GROSSO DO SUL: Coxim,
<date id="FFC910038517FFA45D2DFF6E51EDFF64" box="[1067,1186,151,176]" pageId="37" pageNumber="488" value="1986-02-26">26.II.1986</date>
, Koller (
<specimenCount id="9D71FD4A8517FFA45C01FF615060FF7B" box="[1287,1327,152,176]" pageId="37" pageNumber="488" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068517FFA45C31FF6150C5FF64" box="[1335,1418,152,176]" pageId="37" pageNumber="488">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38517FFA459C1FF4557F1FEC8" blockId="37.[151,1437,151,680]" pageId="37" pageNumber="488">
<emphasis id="B903EAD18517FFA459C1FF4554A1FF01" bold="true" box="[199,494,188,213]" pageId="37" pageNumber="488">
<typeStatus id="54CC88618517FFA459C1FF455470FF01" box="[199,319,188,213]" pageId="37" pageNumber="488" type="paratype">Paratypes</typeStatus>
[5]:
<collectingCountry id="F36076538517FFA45885FF4554A1FF01" box="[387,494,188,213]" name="Brazil" pageId="37" pageNumber="488">BRAZIL</collectingCountry>
</emphasis>
:
<collectionCode id="ED66AE068517FFA45B02FF455711FF00" box="[516,606,188,212]" pageId="37" pageNumber="488">MINAS</collectionCode>
GERAIS: São Roque de Minas, Parque Nacional Serra da Canastra, (
<geoCoordinate id="EE4350048517FFA459A6FF26546DFF2C" box="[160,290,223,248]" direction="south" orientation="latitude" pageId="37" pageNumber="488" precision="15" value="-20.254723">20°15'17&quot;S</geoCoordinate>
,
<geoCoordinate id="EE4350048517FFA45835FF2654F0FF2C" box="[307,447,223,248]" direction="west" orientation="longitude" pageId="37" pageNumber="488" precision="15" value="-46.419724">46°25'11&quot;W</geoCoordinate>
),
<date id="FFC910038517FFA458DEFF195703FF2C" box="[472,588,224,248]" pageId="37" pageNumber="488" value="2007-11-09">9.XI.2007</date>
, M. F. Souza (
<specimenCount id="9D71FD4A8517FFA45A00FF19567CFF23" box="[774,819,224,248]" pageId="37" pageNumber="488" type="generic">1 ♂</specimenCount>
2 ♀
<collectionCode id="ED66AE068517FFA45A7DFF195681FF2C" box="[891,974,224,248]" pageId="37" pageNumber="488">CEMT</collectionCode>
);
<collectionCode id="ED66AE068517FFA45AE0FF265151FF2C" box="[998,1054,223,248]" httpUri="http://grbio.org/cool/2pta-s8y1" name="Sammlung Oberli" pageId="37" pageNumber="488">SÃO</collectionCode>
<collectionCode id="ED66AE068517FFA45D2DFF1951C5FF2C" box="[1067,1162,224,248]" pageId="37" pageNumber="488">PAULO</collectionCode>
: Piracicaba, E.E.C.J.V.,
<date id="FFC910038517FFA45991FEFD545DFEC9" box="[151,274,260,285]" pageId="37" pageNumber="488" value="1966-04-30">30.IV.1966</date>
(
<specimenCount id="9D71FD4A8517FFA45823FEFD5402FEC8" box="[293,333,260,284]" pageId="37" pageNumber="488" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068517FFA45852FEFD54E8FEC8" box="[340,423,260,284]" pageId="37" pageNumber="488">CEMT</collectionCode>
); [no data] (
<specimenCount id="9D71FD4A8517FFA45B36FEFD5716FEC8" box="[560,601,260,284]" pageId="37" pageNumber="488" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE068517FFA45B59FEFD57FDFEC8" box="[607,690,260,284]" pageId="37" pageNumber="488">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38517FFA459C1FEDE5799FE94" blockId="37.[151,1437,151,680]" box="[199,726,295,320]" pageId="37" pageNumber="488">
<emphasis id="B903EAD18517FFA459C1FEDE54E3FE94" bold="true" box="[199,428,295,320]" pageId="37" pageNumber="488">Sampling methods.</emphasis>
at light-trap [
<specimenCount id="9D71FD4A8517FFA45B41FED15784FE94" box="[583,715,295,320]" pageId="37" pageNumber="488" type="generic">1 specimen</specimenCount>
].
</paragraph>
<paragraph id="8BC836C38517FFA459C1FEB550DDFEB1" blockId="37.[151,1437,151,680]" box="[199,1426,332,357]" pageId="37" pageNumber="488">
<emphasis id="B903EAD18517FFA459C1FEB55465FEB1" bold="true" box="[199,298,332,357]" pageId="37" pageNumber="488">Habitat.</emphasis>
<collectingCountry id="F36076538517FFA45837FEB5543AFEB1" box="[305,373,332,357]" name="Brazil" pageId="37" pageNumber="488">Brazil</collectingCountry>
(Minas Gerais and Mato Grosso) (
<figureCitation id="134C2A468517FFA45BFAFEB5561FFEB0" box="[764,848,332,357]" captionStart="FIGURES 93 94" captionStartId="47.[151,264,1007,1030]" captionTargetBox="[151,1436,193,984]" captionTargetId="figure@47.[151,1436,193,985]" captionTargetPageId="47" captionText="FIGURES 93 94. Known distribution of D. (Aganhyboma). 93. D. icaroides and D. valgum. 94. icariforme subcomplex." httpUri="https://zenodo.org/record/244806/files/figure.png" pageId="37" pageNumber="488">Fig. 94</figureCitation>
). From available data this species inhabits Cerrado.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488517FFA459C1FE96500FFD7C" pageId="37" pageNumber="488" type="diagnosis">
<paragraph id="8BC836C38517FFA459C1FE9657ABFDB4" blockId="37.[151,1437,151,680]" pageId="37" pageNumber="488">
<emphasis id="B903EAD18517FFA459C1FE9654AEFE5C" bold="true" box="[199,481,367,392]" pageId="37" pageNumber="488">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408517FFA45B75FE8957A6FE5C" box="[627,745,368,392]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="icariforme">
<emphasis id="B903EAD18517FFA45B75FE8957A6FE5C" box="[627,745,368,392]" italics="true" pageId="37" pageNumber="488">icariforme</emphasis>
</taxonomicName>
subcomplex): length&gt;
<quantity id="4C8F9B268517FFA45D06FE895119FE53" box="[1024,1110,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="37" pageNumber="488" unit="mm" value="13.0">13 mm</quantity>
; posteromedial punctures of pronotum usually separated by less than one diameter; length of apical carina of seventh interstria less than or equal to the sum of lengths of apical carinae of fifth and sixth interstriae; lateral margins of elytra almost parallel. The following aedeagus characters are also diagnostic: parameres symmetric and elongate, directed downward, almost at an angle of 60° with phallobase; apex weakly acuminate (
<figureCitation id="134C2A468517FFA45A8EFE0656ACFDCC" box="[904,995,511,536]" captionStart="FIGURES 12 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="37" pageNumber="488">Fig. 19</figureCitation>
). Lamella copulatrix (LC): left lobe subrectangular (as in
<figureCitation id="134C2A468517FFA4588EFDDD5494FDE8" box="[392,475,548,573]" captionStart="FIGURES 37 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="37" pageNumber="488">Fig. 41</figureCitation>
); right lobe “b”-shaped (as in
<figureCitation id="134C2A468517FFA45A28FDDD56CFFDE8" box="[814,896,548,573]" captionStart="FIGURES 37 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="37" pageNumber="488">Fig. 41</figureCitation>
). Superior right peripheral sclerite as in Fig. 49. Fronto-lateral peripheral sclerite (FLP) as in
<figureCitation id="134C2A468517FFA45B8BFDBE5790FDB4" box="[653,735,583,608]" captionStart="FIGURES 55 58" captionStartId="43.[151,264,1783,1806]" captionTargetBox="[205,1383,207,1725]" captionTargetId="figure@43.[174,1383,193,1728]" captionTargetPageId="43" captionText="FIGURES 55 58. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 55. D. valgum. 56. D. kolleri new species. 57. D. icariforme. 58. D. ritamourae new species. Figure 59. Complex of axial and subaxial sclerites (A + AS) in D. viridescens new status. Figures 60 61. Head in D. (Aganhyboma). 60. D. viridescens new status. 61. D. longiceps new status. Figures 62 63. Lateral view of the elytra in D. (Aganhyboma). 62. D. viridescens new status (arrow points the basal carina of ninth interstria). 63. D. finestriatum new species. (arrow points the basal carina of ninth interstria). Figures 64 69. Legs. 64. Metafemur in D. viridescens new status. 65. Metafemur in D. longiceps new status. 66. Protibia in D. trisignatum. 67. Metatibia in D. trisignatum. 68. Metatibia in D. viridescens new status. 69. Profemur in D. trisignatum (arrow indicates the anterior margin discontinuous). Figures scale = 1 mm." httpUri="https://zenodo.org/record/244802/files/figure.png" pageId="37" pageNumber="488">Fig. 56</figureCitation>
.
</paragraph>
<paragraph id="8BC836C38517FFA459C1FD95500FFD7C" blockId="37.[151,1437,151,680]" pageId="37" pageNumber="488">
The punctures at center of elytral interstriae, on anterior portion of elytra, are the most dense of
<taxonomicName id="4C774D408517FFA45C20FD9550D3FD50" box="[1318,1436,620,644]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="488" phylum="Arthropoda" rank="species" species="icariforme">
<emphasis id="B903EAD18517FFA45C20FD9550D3FD50" box="[1318,1436,620,644]" italics="true" pageId="37" pageNumber="488">icariforme</emphasis>
</taxonomicName>
subcomplex. Apical carina of sixth interstria can present the same length of apical carina of fifth interstria.
</paragraph>
</subSubSection>
</treatment>
</document>