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<document id="510F612595AC13163E3F90B20843940D" ID-DOI="10.11646/zootaxa.3694.5.3" ID-GBIF-Dataset="e6118b09-adf4-4ba0-a985-93914739952d" ID-ISSN="1175-5326" ID-Zenodo-Dep="223454" ID-ZooBank="15D560C8-3DAA-4216-85CE-473D06E4351D" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460659505958" checkinUser="plazi" docAuthor="Mauad, Melina, Siri, Augusto &amp; Donato, Mariano" docDate="2013" docId="03DE879CFFA7FFADFF058F34FEF1FB07" docLanguage="en" docName="zt03694p460.pdf" docOrigin="Zootaxa 3694 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Allocladius quadrus Andersen, Saether &amp; Mendes" docType="treatment" docVersion="8" lastPageNumber="457" masterDocId="FFE7FFE4FFACFFA1FF928B20FC31FFE5" masterDocTitle="New species of Pseudosmittia Edwards, 1932 and new records of Allocladius Kieffer, 1913 (Diptera: Chironomidae, Orthocladiinae) from South America" masterLastPageNumber="460" masterPageNumber="445" pageNumber="456" updateTime="1698347377064" updateUser="plazi">
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<mods:title id="BD5F6EF52838BD8C60EC19C5F4A35410">New species of Pseudosmittia Edwards, 1932 and new records of Allocladius Kieffer, 1913 (Diptera: Chironomidae, Orthocladiinae) from South America</mods:title>
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<mods:namePart id="94E608D6919F6D7164069BFFA2F39A51">Mauad, Melina</mods:namePart>
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<mods:namePart id="BD8E38DBB2E7C6D95751B6663ED77DD9">Siri, Augusto</mods:namePart>
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<mods:namePart id="3865C2741569B20E8E318B822BE9BC2E">Donato, Mariano</mods:namePart>
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<mods:title id="18C375490530D973857E4FB6A2708563">Zootaxa</mods:title>
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<mods:date id="BA17DDA0C21AC8CDCAE8FF4431F25350">2013</mods:date>
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<treatment id="03DE879CFFA7FFADFF058F34FEF1FB07" ID-DOI="http://doi.org/10.5281/zenodo.6155056" ID-GBIF-Taxon="125135855" ID-Zenodo-Dep="6155056" LSID="urn:lsid:plazi:treatment:03DE879CFFA7FFADFF058F34FEF1FB07" httpUri="http://treatment.plazi.org/id/03DE879CFFA7FFADFF058F34FEF1FB07" lastPageId="12" lastPageNumber="457" pageId="11" pageNumber="456">
<subSubSection id="C36D6501FFA7FFAAFF058F34F86BFB94" pageId="11" pageNumber="456" type="nomenclature">
<paragraph id="8BC8368AFFA7FFAAFF058F34FECBFBCB" blockId="11.[151,762,1044,1070]" box="[151,762,1044,1070]" pageId="11" pageNumber="456">
<heading id="D08081E6FFA7FFAAFF058F34FECBFBCB" bold="true" box="[151,762,1044,1070]" fontSize="11" level="1" pageId="11" pageNumber="456" reason="1">
<taxonomicName id="4C774D09FFA7FFAAFF058F34FECBFBCB" ID-CoL="89X4P" authority="Andersen, Saether &amp; Mendes" authorityName="Andersen, Saether &amp; Mendes" box="[151,762,1044,1070]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="456" phylum="Arthropoda" rank="species" species="quadrus">
<emphasis id="B903EA98FFA7FFAAFF058F34FECBFBCB" bold="true" box="[151,762,1044,1070]" pageId="11" pageNumber="456">
<emphasis id="B903EA98FFA7FFAAFF058F34FDBBFBCB" bold="true" box="[151,394,1044,1070]" italics="true" pageId="11" pageNumber="456">Allocladius quadrus</emphasis>
Andersen, Saether &amp; Mendes
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC8368AFFA7FFAAFF058F7BF86BFB94" blockId="11.[151,1114,1115,1137]" box="[151,1114,1115,1137]" pageId="11" pageNumber="456">
<taxonomicName id="4C774D09FFA7FFAAFF058F7BFF36FB94" authority="Andersen, Saether and Mendes, 2010: 17" authorityName="Andersen, Saether and Mendes" authorityPageNumber="17" authorityYear="2010" box="[151,775,1115,1137]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="456" phylum="Arthropoda" rank="species" species="quadrus">
<emphasis id="B903EA98FFA7FFAAFF058F7BFD52FB94" box="[151,355,1115,1137]" italics="true" pageId="11" pageNumber="456">Allocladius quadrus</emphasis>
Andersen, Saether and Mendes, 2010: 17
</taxonomicName>
; Ferrington &amp; Saether (2011: 96).
</paragraph>
</subSubSection>
<subSubSection id="C36D6501FFA7FFAAFF058FBEF894FB1A" pageId="11" pageNumber="456" type="materials_examined">
<paragraph id="8BC8368AFFA7FFAAFF058FBEF894FB1A" blockId="11.[151,1437,1182,1963]" pageId="11" pageNumber="456">
<emphasis id="B903EA98FFA7FFAAFF058FBEFD4BFB52" bold="true" box="[151,378,1182,1207]" pageId="11" pageNumber="456">Material examined</emphasis>
.
<collectingCountry id="F360761AFFA7FFAAFE1A8FBFFE01FB52" box="[392,560,1183,1207]" name="Argentina" pageId="11" pageNumber="456">ARGENTINA</collectingCountry>
:
<specimenCount id="9D71FD03FFA7FFAAFDA98FBFFEA4FB52" box="[571,661,1182,1207]" pageId="11" pageNumber="456" type="male">2 males</specimenCount>
, Salta, Quebrada del Agua,
<geoCoordinate id="EE43504DFFA7FFAAFC438FBFF84BFB52" box="[977,1146,1183,1207]" direction="south" orientation="latitude" pageId="11" pageNumber="456" precision="1" value="-24.509167">24º 30' 33.0'' S</geoCoordinate>
,
<geoCoordinate id="EE43504DFFA7FFAAFB1A8FBEF90BFB53" box="[1160,1338,1182,1207]" direction="west" orientation="longitude" pageId="11" pageNumber="456" precision="1" value="-68.181274">68º 10' 52.6'' W</geoCoordinate>
,
<quantity id="4C8F9B6FFFA7FFAAFAD48FBEF9ADFB53" box="[1350,1436,1182,1207]" metricMagnitude="3" metricUnit="m" metricValue="3.678" pageId="11" pageNumber="456" unit="m" value="3678.0">3678 m</quantity>
a.s.l.,
<date id="FFC9104AFFA7FFAAFF4B8FE2FD75FB3E" box="[217,324,1218,1243]" pageId="11" pageNumber="456" value="2005-01-11">11.i.2005</date>
, sweep net, M. Donato;
<specimenCount id="9D71FD03FFA7FFAAFDC78FE2FE92FB3E" box="[597,675,1218,1243]" pageId="11" pageNumber="456" type="male">1 male</specimenCount>
, Río Negro, P.N. Nahuel Huapi, laguna Los Juncos,
<geoCoordinate id="EE43504DFFA7FFAAFB628FE2F9A6FB3F" box="[1264,1431,1218,1242]" direction="south" orientation="latitude" pageId="11" pageNumber="456" precision="1" value="-41.060528">41º 03' 37.9'' S</geoCoordinate>
,
<geoCoordinate id="EE43504DFFA7FFAAFF058FC7FD79FB1B" box="[151,328,1255,1279]" direction="west" orientation="longitude" pageId="11" pageNumber="456" precision="1" value="-71.00953">71º 00' 34.3'' W</geoCoordinate>
,
<quantity id="4C8F9B6FFFA7FFAAFEC18FC6FDABFB1B" box="[339,410,1254,1279]" metricMagnitude="2" metricUnit="m" metricValue="9.059999999999999" pageId="11" pageNumber="456" unit="m" value="906.0">906 m</quantity>
a.s.l.,
<date id="FFC9104AFFA7FFAAFE708FC6FE55FB1A" box="[482,612,1254,1279]" pageId="11" pageNumber="456" value="2007-12-12">12.xii.2007</date>
, sweep net, A. Garre and F. Montes de Oca (MLP).
</paragraph>
</subSubSection>
<subSubSection id="C36D6501FFA7FFAAFF558E2AFE44F8FF" pageId="11" pageNumber="456" type="description">
<paragraph id="8BC8368AFFA7FFAAFF558E2AFE06FA8F" blockId="11.[151,1437,1182,1963]" pageId="11" pageNumber="456">
<emphasis id="B903EA98FFA7FFAAFF558E2AFD35FAC6" bold="true" box="[199,260,1290,1315]" pageId="11" pageNumber="456">Male</emphasis>
(
<emphasis id="B903EA98FFA7FFAAFE8A8E2BFD17FAC7" box="[280,294,1291,1314]" italics="true" pageId="11" pageNumber="456">n</emphasis>
= 3, except when otherwise stated). The following measurements are different from the original description. Total length
<quantity id="4C8F9B6FFFA7FFAAFE228E0FFE0BFAA3" box="[432,570,1327,1351]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="11" pageNumber="456" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.32.4 mm</quantity>
. Wing length
<quantity id="4C8F9B6FFFA7FFAAFD458E0EFF4CFAA3" box="[727,893,1326,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.55" metricValueMin="1.35" pageId="11" pageNumber="456" unit="mm" value="1.45" valueMax="1.55" valueMin="1.35">1.351.55 mm</quantity>
. Total length/wing length extended to 1.7. Wing length/length of profemur 2.703.04.
</paragraph>
<paragraph id="8BC8368AFFA7FFAAFF558E56FE52FA32" blockId="11.[151,1437,1182,1963]" pageId="11" pageNumber="456">Head. AR 1.21.2. Minimum length of ultimate flagellomere 390. Tentorium 115124 long, 2234 wide. Stipes 115120 long, 710 wide. Palpomere lengths (25) 3946, 7183, 88103, 120127. Third palpomere with 2 lanceolate sensilla clavata, 1921 long.</paragraph>
<paragraph id="8BC8368AFFA7FFAAFF558EC2FFDEFA1F" blockId="11.[151,1437,1182,1963]" box="[199,1007,1506,1531]" pageId="11" pageNumber="456">Thorax. Dorsocentrals 710, acrostichals 35, scutellum with 67 setae.</paragraph>
<paragraph id="8BC8368AFFA7FFAAFF558D26FE6AF9FA" blockId="11.[151,1437,1182,1963]" box="[199,603,1542,1567]" pageId="11" pageNumber="456">Wing. Minimum value for VR 1.25.</paragraph>
<paragraph id="8BC8368AFFA7FFAAFF558D0BFE04F982" blockId="11.[151,1437,1182,1963]" pageId="11" pageNumber="456">
Legs. Width at apex of fore tibia 2529, mid tibia 2430, of hind tibia 3742. Comb of 1316 setae. Length and proportions of legs as in
<tableCitation id="C6F50331FFA7FFAAFE4E8D6EFE03F982" box="[476,562,1614,1639]" captionStart="TABLE 6" captionStartId="12.[151,239,152,175]" captionTargetBox="[151,1340,234,710]" captionTargetPageId="12" captionText="TABLE 6. Lengths (in µm) and proportions of legs of Allocladius quadrus (male) (n = 3, except when otherwise stated). For abbreviation explanations see Table 1." httpUri="http://table.plazi.org/id/DF086602FFA0FFADFF058BB8FE6BFF2A" pageId="11" pageNumber="456" tableUuid="DF086602FFA0FFADFF058BB8FE6BFF2A">Table 6</tableCitation>
.
</paragraph>
<paragraph id="8BC8368AFFA7FFAAFF558D52FE44F8FF" blockId="11.[151,1437,1182,1963]" pageId="11" pageNumber="456">Hypopygium. Anal point 4041, including 26 long basal sclerotized part with 8 setae and 1415 long bare, hyaline distal part. Tergite IX with 68 setae in addition to those on anal point. Phallapodeme 4446 long. Transverse sternapodeme 6176 long. Virga 5 long, 1014 wide. Pars ventralis (Fig. 44) single, subquadrangular, 7593 (2) long, 80103 (2) wide. Gonocoxite 166181 long. Inferior volsella reaching to 0.430.46 gonocoxite length. Gonostylus 7173; megaseta 710.</paragraph>
</subSubSection>
<subSubSection id="C36D6501FFA7FFADFF558C06FEF1FB07" lastPageId="12" lastPageNumber="457" pageId="11" pageNumber="456" type="discussion">
<paragraph id="8BC8368AFFA7FFAAFF558C06FD2AF84E" blockId="11.[151,1437,1182,1963]" pageId="11" pageNumber="456">
<emphasis id="B903EA98FFA7FFAAFF558C06FD05F8DA" bold="true" box="[199,308,1830,1855]" pageId="11" pageNumber="456">Remarks</emphasis>
. The specimens from Quebrada del agua in Salta province were collected at a Punean spring and the specimen collected at laguna Los Juncos in Rio Negro province at a small temporal lagoon (
<figureCitation id="134C2A0FFFA7FFAAFB0A8C6AF8DAF887" box="[1176,1259,1866,1891]" captionStart="FIGURE 13" captionStartId="8.[151,250,1975,1997]" captionTargetBox="[228,1355,193,1960]" captionTargetId="figure@8.[228,1356,193,1961]" captionTargetPageId="8" captionText="FIGURE 13. Map showing the new records of Allocladius globosus and A. quadrus. A. globosus: square, known records; triangle, new records. A. quadrus: star, known records; circle, new records." httpUri="https://zenodo.org/record/223458/files/figure.png" pageId="11" pageNumber="456">Fig. 13</figureCitation>
). This lagoon is located in the Patagonian steppe. These new records define the distributional pattern of
<taxonomicName id="4C774D09FFA7FFAAFB088C4FF928F862" box="[1178,1305,1902,1927]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="456" phylum="Arthropoda" rank="species" species="quadrus">
<emphasis id="B903EA98FFA7FFAAFB088C4FF928F862" box="[1178,1305,1902,1927]" italics="true" pageId="11" pageNumber="456">A. quadrus</emphasis>
</taxonomicName>
as Andean Patagonian.
</paragraph>
<caption id="DF086602FFA0FFADFF058BB8FE6BFF2A" ID-Table-UUID="DF086602FFA0FFADFF058BB8FE6BFF2A" httpUri="http://table.plazi.org/id/DF086602FFA0FFADFF058BB8FE6BFF2A" pageId="12" pageNumber="457" targetBox="[151,1340,234,710]" targetIsTable="true" targetPageId="12">
<paragraph id="8BC8368AFFA0FFADFF058BB8FE6BFF2A" blockId="12.[151,1437,151,207]" pageId="12" pageNumber="457">
<emphasis id="B903EA98FFA0FFADFF058BB8FD38FF4A" bold="true" box="[151,265,152,175]" pageId="12" pageNumber="457">TABLE 6.</emphasis>
Lengths (in µm) and proportions of legs of
<taxonomicName id="4C774D09FFA0FFADFD4F8BB7FF85FF4A" box="[733,948,151,175]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="quadrus">
<emphasis id="B903EA98FFA0FFADFD4F8BB7FF85FF4A" box="[733,948,151,175]" italics="true" pageId="12" pageNumber="457">Allocladius quadrus</emphasis>
</taxonomicName>
(male) (
<emphasis id="B903EA98FFA0FFADFB828BB8F82CFF4A" box="[1040,1053,152,175]" italics="true" pageId="12" pageNumber="457">n</emphasis>
= 3, except when otherwise stated). For abbreviation explanations see Table 1.
</paragraph>
</caption>
<paragraph id="8BC8368AFFA0FFADFF4F8BCAFD41FD23" pageId="12" pageNumber="457">
<table id="F977C42AFFA0005EFF058BCAF90DFD23" box="[151,1340,234,710]" gridcols="5" gridrows="9" pageId="12" pageNumber="457">
<tr id="354734C8FFA0005EFF058BCAF90DFEE1" box="[151,1340,234,260]" gridrow="0" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058BCAFD41FEE1" box="[151,368,234,260]" gridcol="0" gridrow="0" pageId="12" pageNumber="457">fe</th>
<th id="76965DB4FFA0005EFE408BCAFE07FEE1" box="[466,566,234,260]" gridcol="1" gridrow="0" pageId="12" pageNumber="457">ti</th>
<th id="76965DB4FFA0005EFD5A8BCAFF60FEE1" box="[712,849,234,260]" gridcol="2" gridrow="0" pageId="12" pageNumber="457">ta1</th>
<th id="76965DB4FFA0005EFC2C8BCAF876FEE1" box="[958,1095,234,260]" gridcol="3" gridrow="0" pageId="12" pageNumber="457">ta2</th>
<th id="76965DB4FFA0005EFB218BCAF90DFEE1" box="[1203,1340,234,260]" gridcol="4" gridrow="0" pageId="12" pageNumber="457">ta3</th>
</tr>
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<th id="76965DB4FFA0005EFF058A36FD41FED5" box="[151,368,278,304]" gridcol="0" gridrow="1" pageId="12" pageNumber="457">P1 470510</th>
<td id="76965DB4FFA0005EFE408A36FE07FED5" box="[466,566,278,304]" gridcol="1" gridrow="1" pageId="12" pageNumber="457">570620</td>
<td id="76965DB4FFA0005EFD5A8A36FF60FED5" box="[712,849,278,304]" gridcol="2" gridrow="1" pageId="12" pageNumber="457">260270 (2)</td>
<td id="76965DB4FFA0005EFC2C8A36F876FED5" box="[958,1095,278,304]" gridcol="3" gridrow="1" pageId="12" pageNumber="457">160180 (2)</td>
<td id="76965DB4FFA0005EFB218A36F90DFED5" box="[1203,1340,278,304]" gridcol="4" gridrow="1" pageId="12" pageNumber="457">110 (1)</td>
</tr>
<tr id="354734C8FFA0005EFF058A63F90DFEB8" box="[151,1340,323,349]" gridrow="2" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058A63FD41FEB8" box="[151,368,323,349]" gridcol="0" gridrow="2" pageId="12" pageNumber="457">P2 550600</th>
<td id="76965DB4FFA0005EFE408A63FE07FEB8" box="[466,566,323,349]" gridcol="1" gridrow="2" pageId="12" pageNumber="457">570610</td>
<td id="76965DB4FFA0005EFD5A8A63FF60FEB8" box="[712,849,323,349]" gridcol="2" gridrow="2" pageId="12" pageNumber="457">220250</td>
<td id="76965DB4FFA0005EFC2C8A63F876FEB8" box="[958,1095,323,349]" gridcol="3" gridrow="2" pageId="12" pageNumber="457">130140</td>
<td id="76965DB4FFA0005EFB218A63F90DFEB8" box="[1203,1340,323,349]" gridcol="4" gridrow="2" pageId="12" pageNumber="457">110 (2)</td>
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<tr id="354734C8FFA0005EFF058A52F90DFE69" box="[151,1340,370,396]" gridrow="3" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058A52FD41FE69" box="[151,368,370,396]" gridcol="0" gridrow="3" pageId="12" pageNumber="457">P3 570610</th>
<td id="76965DB4FFA0005EFE408A52FE07FE69" box="[466,566,370,396]" gridcol="1" gridrow="3" pageId="12" pageNumber="457">600630</td>
<td id="76965DB4FFA0005EFD5A8A52FF60FE69" box="[712,849,370,396]" gridcol="2" gridrow="3" pageId="12" pageNumber="457">350350 (1)</td>
<td id="76965DB4FFA0005EFC2C8A52F876FE69" box="[958,1095,370,396]" gridcol="3" gridrow="3" pageId="12" pageNumber="457">180180 (1)</td>
<td id="76965DB4FFA0005EFB218A52F90DFE69" box="[1203,1340,370,396]" gridcol="4" gridrow="3" pageId="12" pageNumber="457">150 (1)</td>
</tr>
<tr id="354734C8FFA0005EFF058A80F90DFE5F" box="[151,1340,416,442]" gridrow="4" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058A80FD41FE5F" box="[151,368,416,442]" gridcol="0" gridrow="4" pageId="12" pageNumber="457">ta4</th>
<td id="76965DB4FFA0005EFE408A80FE07FE5F" box="[466,566,416,442]" gridcol="1" gridrow="4" pageId="12" pageNumber="457">ta5</td>
<td id="76965DB4FFA0005EFD5A8A80FF60FE5F" box="[712,849,416,442]" gridcol="2" gridrow="4" pageId="12" pageNumber="457">LR</td>
<td id="76965DB4FFA0005EFC2C8A80F876FE5F" box="[958,1095,416,442]" gridcol="3" gridrow="4" pageId="12" pageNumber="457">BV</td>
<td id="76965DB4FFA0005EFB218A80F90DFE5F" box="[1203,1340,416,442]" gridcol="4" gridrow="4" pageId="12" pageNumber="457">SV</td>
</tr>
<tr id="354734C8FFA0005EFF058AECF90DFE00" box="[151,1340,460,485]" gridrow="5" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058AECFD41FE00" box="[151,368,460,485]" gridcol="0" gridrow="5" pageId="12" pageNumber="457">P1 70 (1)</th>
<td id="76965DB4FFA0005EFE408AECFE07FE00" box="[466,566,460,485]" gridcol="1" gridrow="5" pageId="12" pageNumber="457">70 (1)</td>
<td id="76965DB4FFA0005EFD5A8AECFF60FE00" box="[712,849,460,485]" gridcol="2" gridrow="5" pageId="12" pageNumber="457">0.430.44 (2)</td>
<td id="76965DB4FFA0005EFC2C8AECF876FE00" box="[958,1095,460,485]" gridcol="3" gridrow="5" pageId="12" pageNumber="457">3.323.32 (1)</td>
<td id="76965DB4FFA0005EFB218AECF90DFE00" box="[1203,1340,460,485]" gridcol="4" gridrow="5" pageId="12" pageNumber="457">4.194.23 (2)</td>
</tr>
<tr id="354734C8FFA0005EFF058ADAF90DFDF1" box="[151,1340,506,532]" gridrow="6" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058ADAFD41FDF1" box="[151,368,506,532]" gridcol="0" gridrow="6" pageId="12" pageNumber="457">P2 6080 (2)</th>
<td id="76965DB4FFA0005EFE408ADAFE07FDF1" box="[466,566,506,532]" gridcol="1" gridrow="6" pageId="12" pageNumber="457">6070 (2)</td>
<td id="76965DB4FFA0005EFD5A8ADAFF60FDF1" box="[712,849,506,532]" gridcol="2" gridrow="6" pageId="12" pageNumber="457">0.390.41</td>
<td id="76965DB4FFA0005EFC2C8ADAF876FDF1" box="[958,1095,506,532]" gridcol="3" gridrow="6" pageId="12" pageNumber="457">3.653.83 (2)</td>
<td id="76965DB4FFA0005EFB218ADAF90DFDF1" box="[1203,1340,506,532]" gridcol="4" gridrow="6" pageId="12" pageNumber="457">4.845.09</td>
</tr>
<tr id="354734C8FFA0005EFF058908F90DFDA4" box="[151,1340,552,577]" gridrow="7" pageId="12" pageNumber="457">
<th id="76965DB4FFA0005EFF058908FD41FDA4" box="[151,368,552,577]" gridcol="0" gridrow="7" pageId="12" pageNumber="457">P3 80 (1)</th>
<td id="76965DB4FFA0005EFE408908FE07FDA4" box="[466,566,552,577]" gridcol="1" gridrow="7" pageId="12" pageNumber="457">70 (1)</td>
<td id="76965DB4FFA0005EFD5A8908FF60FDA4" box="[712,849,552,577]" gridcol="2" gridrow="7" pageId="12" pageNumber="457">0.560.56 (1)</td>
<td id="76965DB4FFA0005EFC2C8908F876FDA4" box="[958,1095,552,577]" gridcol="3" gridrow="7" pageId="12" pageNumber="457">3.313.31 (1)</td>
<td id="76965DB4FFA0005EFB218908F90DFDA4" box="[1203,1340,552,577]" gridcol="4" gridrow="7" pageId="12" pageNumber="457">3.543.54 (1)</td>
</tr>
<tr id="354734C8FFA0005EFF05898CF90DFD23" box="[151,1340,684,710]" gridrow="8" pageId="12" pageNumber="457" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76965DB4FFA0005EFF05898CFD41FD23" box="[151,368,684,710]" gridcol="0" gridrow="8" pageId="12" pageNumber="457">Cladistic analysis</th>
</tr>
</table>
</paragraph>
<paragraph id="8BC8368AFFA0FFADFF0589D3FE0FFC26" blockId="12.[151,1436,755,1251]" pageId="12" pageNumber="457">
The analyses of the data matrix using only male characters as defined by Ferrington &amp; Saether (2011) under concavities K=2 to K=20 showed the best group support under K=5. In this concavity the analysis yielded 7 trees (Fit= 63.62, CI= 0.17, RI= 0.56) and the resulting strict consensus tree is shown in
<figureCitation id="134C2A0FFFA0FFADFBAD881BF89FFCB1" box="[1087,1198,827,852]" captionStart="FIGURE 14" captionStartId="13.[151,250,1965,1987]" captionTargetBox="[432,1137,202,1929]" captionTargetId="figure@13.[412,1175,185,1945]" captionTargetPageId="13" captionText="FIGURE 14. Strict consensus tree obtained from the 7 trees yielded under K = 5 (Fit = 63.62, CI = 0.17, RI = 0.56). Number above branches indicates absolute Bremer support and number below branches indicates relative Bremer support." httpUri="https://zenodo.org/record/223459/files/figure.png" pageId="12" pageNumber="457">Figure 14</figureCitation>
. The species
<taxonomicName id="4C774D09FFA0FFADFAD6881CF9ADFCB1" box="[1348,1436,827,852]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="larga">
<emphasis id="B903EA98FFA0FFADFAD6881CF9ADFCB1" box="[1348,1436,827,852]" italics="true" pageId="12" pageNumber="457">P. larga</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D09FFA0FFADFF578840FD03FC9D" box="[197,306,863,888]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="saetheri">
<emphasis id="B903EA98FFA0FFADFF578840FD03FC9D" box="[197,306,863,888]" italics="true" pageId="12" pageNumber="457">P. saetheri</emphasis>
</taxonomicName>
conform a clade supported by the characters Cu1 straight [21 (0)]; superior volsella with strong orally curved microtrichia [57 (1)] and inferior volsella without accessory lobe [63 (1)]. This clade is located in a major clade in the
<emphasis id="B903EA98FFA0FFADFE8D888BFD49FC27" box="[287,376,939,962]" italics="true" pageId="12" pageNumber="457">angusta</emphasis>
group of species.
</paragraph>
<paragraph id="8BC8368AFFA0FFADFF5588EEFEA9FBB6" blockId="12.[151,1436,755,1251]" pageId="12" pageNumber="457">
The species
<taxonomicName id="4C774D09FFA0FFADFEC388EFFDC0FC02" box="[337,497,974,999]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="trapezoidea">
<emphasis id="B903EA98FFA0FFADFEC388EFFDC0FC02" box="[337,497,974,999]" italics="true" pageId="12" pageNumber="457">P. trapezoidea</emphasis>
</taxonomicName>
is the sister group of the clade ((
<taxonomicName id="4C774D09FFA0FFADFCCD88EFFFCAFC03" box="[863,1019,975,998]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="propetropis">
<emphasis id="B903EA98FFA0FFADFCCD88EFFFCAFC03" box="[863,1019,975,998]" italics="true" pageId="12" pageNumber="457">P. propetropis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C774D09FFA0FFADFB9988EFF857FC03" box="[1035,1126,975,998]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="tropis">
<emphasis id="B903EA98FFA0FFADFB9988EFF857FC03" box="[1035,1126,975,998]" italics="true" pageId="12" pageNumber="457">P. tropis</emphasis>
</taxonomicName>
<taxonomicName id="4C774D09FFA0FFADFB1088EFF936FC02" box="[1154,1287,974,999]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="lamellata">
<emphasis id="B903EA98FFA0FFADFB1088EFF936FC02" box="[1154,1287,974,999]" italics="true" pageId="12" pageNumber="457">P. lamellata</emphasis>
</taxonomicName>
)) sharing the characters inner verticals of temporal setae absent [5 (1)] and virga 3557 long [46 (1)]. This species is part of a major clade constituted by all the species of the
<taxonomicName id="4C774D09FFA0FFADFD248F36FF0FFBCA" box="[694,830,1046,1071]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="brevifurcata">
<emphasis id="B903EA98FFA0FFADFD248F36FF0FFBCA" box="[694,830,1046,1071]" italics="true" pageId="12" pageNumber="457">brevifurcata</emphasis>
</taxonomicName>
group except
<taxonomicName id="4C774D09FFA0FFADFC728F37F8B4FBCA" box="[992,1157,1046,1071]" class="Insecta" family="Chironomidae" genus="Pseudosmittia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="brevifurcata">
<emphasis id="B903EA98FFA0FFADFC728F37F8B4FBCA" box="[992,1157,1046,1071]" italics="true" pageId="12" pageNumber="457">P. brevifurcata</emphasis>
</taxonomicName>
that is located in a clade with several species of the
<emphasis id="B903EA98FFA0FFADFE578F1AFE79FBB6" box="[453,584,1082,1107]" italics="true" pageId="12" pageNumber="457">xanthostola</emphasis>
group.
</paragraph>
<paragraph id="8BC8368AFFA0FFADFF558F7FFEF1FB07" blockId="12.[151,1436,755,1251]" pageId="12" pageNumber="457">
In the strict consensus tree
<taxonomicName id="4C774D09FFA0FFADFE6D8F7EFF24FB93" box="[511,789,1118,1143]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="fortispinatus">
<emphasis id="B903EA98FFA0FFADFE6D8F7EFF24FB93" box="[511,789,1118,1143]" italics="true" pageId="12" pageNumber="457">Allocladius fortispinatus</emphasis>
</taxonomicName>
is place at the base of the
<taxonomicName id="4C774D09FFA0FFADFBD88F7EF8FBFB92" box="[1098,1226,1118,1143]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA98FFA0FFADFBD88F7EF8FBFB92" box="[1098,1226,1118,1143]" italics="true" pageId="12" pageNumber="457">Allocladius</emphasis>
</taxonomicName>
clade as the sister species of the rest of the species of the genus except
<taxonomicName id="4C774D09FFA0FFADFD638FA3FF65FB7E" box="[753,852,1154,1179]" class="Insecta" family="Chironomidae" genus="Allocladius" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="B903EA98FFA0FFADFD638FA3FF65FB7E" box="[753,852,1154,1179]" italics="true" pageId="12" pageNumber="457">A. wirthi</emphasis>
</taxonomicName>
sharing the synapomorphies postorbitals present [7 (0)], anal point at least extreme apex without microtrichia or anal point absent [34 (0)], and oral projections of transverse sternapodeme well developed [41 (0)].
</paragraph>
</subSubSection>
</treatment>
</document>