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<mods:title id="544E0A92F0E2E189A3C9B8212E1E6571">Revision of the genus Tanycypris (Ostracoda, Cypricercinae) with the description of Tanycypris alfonsi n. sp., and an identification key to the genus</mods:title>
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<mods:namePart id="991E78C20ADD3C2D43D6B182DD943C8E">Nagler, Christina</mods:namePart>
<mods:affiliation id="46047516DC815E20401FA7590BECA8B6">Aquatic Systems Biology Unit, Department of Ecology and Ecosystem Management, Technische Universitaet Muenchen, Muehlenweg 22, 85350 Freising, Germany. Department of Earth and Environmental Sciences, Palaeontology and Geobiology, GeoBio-Center, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany.</mods:affiliation>
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<mods:affiliation id="4DA3BD4953E84588A9EFA4E73552ACAC">Aquatic Systems Biology Unit, Department of Ecology and Ecosystem Management, Technische Universitaet Muenchen, Muehlenweg 22, 85350 Freising, Germany. Department of Earth and Environmental Sciences, Palaeontology and Geobiology, GeoBio-Center, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany.</mods:affiliation>
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<treatment id="1231A341540A2E661DBBCE0BFBABCCD4" ID-GBIF-Taxon="183717270" LSID="urn:lsid:plazi:treatment:1231A341540A2E661DBBCE0BFBABCCD4" httpUri="http://treatment.plazi.org/id/1231A341540A2E661DBBCE0BFBABCCD4" lastPageNumber="417" pageId="16" pageNumber="417" scope_class="Ostracoda" scope_family="Cyprididae" scope_order="Podocopida" scope_subClass="Podocopa" scope_subFamily="Cypricercinae" scope_subOrder="Podocopina" scope_superFamily="Cypridoidea" scope_tribe="Cypricercini">
<subSubSection id="D28241DC540A2E661DBBCE0BFD85C9BD" box="[151,570,361,387]" pageId="16" pageNumber="417" type="nomenclature">
<paragraph id="9A271257540A2E661DBBCE0BFD85C9BD" blockId="16.[151,570,361,387]" box="[151,570,361,387]" pageId="16" pageNumber="417">
<heading id="C16FA53B540A2E661DBBCE0BFD85C9BD" bold="true" box="[151,570,361,387]" fontSize="11" level="1" pageId="16" pageNumber="417" reason="1">
<taxonomicName id="5D9869D4540A2E661DBBCE0BFD85C9BD" ID-CoL="CD396" authority="Brady, 1904" baseAuthorityName="Brady" baseAuthorityYear="1904" box="[151,570,361,387]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661DBBCE0BFD85C9BD" bold="true" box="[151,570,361,387]" pageId="16" pageNumber="417">
<emphasis id="A8ECCE45540A2E661DBBCE0BFE35C9BD" bold="true" box="[151,394,361,387]" italics="true" pageId="16" pageNumber="417">Cypricercus inermis</emphasis>
(
<bibRefCitation id="FE096FA6540A2E661CB5CE0BFD8DC9BD" author="Brady, G. W." box="[409,562,361,387]" pageId="16" pageNumber="417" pagination="121 - 128" refId="ref12636" refString="Brady, G. W. (1904) On Entomostraca collected in Natal by J. Gibson. Proceedings of the Zoological Society of London, 1904 121 - 128. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1904. tb 08317. x" type="book chapter" year="1904">Brady, 1904</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540A2E661DBBCED2FCF7CA7E" pageId="16" pageNumber="417" type="reference_group">
<paragraph id="9A271257540A2E661DBBCED2FCACC9F8" blockId="16.[151,852,432,576]" box="[151,787,432,454]" pageId="16" pageNumber="417">
* 1904
<taxonomicName id="5D9869D4540A2E661C02CED2FDB3C9F8" authority="Brady" authorityName="Brady" authorityYear="1904" box="[302,524,432,454]" class="Ostracoda" family="Cyprididae" genus="Cypris" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661C02CED2FE7BC9F8" box="[302,452,432,454]" italics="true" pageId="16" pageNumber="417">Cypris inermis</emphasis>
Brady
</taxonomicName>
: 125, pl. VIII. figs 4449.
</paragraph>
<paragraph id="9A271257540A2E661DD9CEACFCEBC9DA" blockId="16.[151,852,432,576]" box="[245,852,462,484]" pageId="16" pageNumber="417">
1910
<taxonomicName id="5D9869D4540A2E661C02CEADFE65C9DA" box="[302,474,463,484]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661C02CEADFE65C9DA" box="[302,474,463,484]" italics="true" pageId="16" pageNumber="417">Eucypris inermis</emphasis>
</taxonomicName>
—Daday: 167169, pl. 9. figs 1833.
</paragraph>
<paragraph id="9A271257540A2E661DD9CE8FFD61CA3D" blockId="16.[151,852,432,576]" box="[245,734,493,515]" pageId="16" pageNumber="417">
1971
<taxonomicName id="5D9869D4540A2E661C02CE8CFE51CA3D" box="[302,494,494,515]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661C02CE8CFE51CA3D" box="[302,494,494,515]" italics="true" pageId="16" pageNumber="417">Tanycypris inermis</emphasis>
</taxonomicName>
—McKenzie: 172, 208.
</paragraph>
<paragraph id="9A271257540A2E661DD9CD6EFD50CA1C" blockId="16.[151,852,432,576]" box="[245,751,524,546]" pageId="16" pageNumber="417">
2001
<taxonomicName id="5D9869D4540A2E661C02CD6EFDB6CA1F" authorityName="Sars" authorityYear="1895" box="[302,521,524,546]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="cuneatus">
<emphasis id="A8ECCE45540A2E661C02CD6EFDB6CA1F" box="[302,521,524,546]" italics="true" pageId="16" pageNumber="417">Cypricercus cuneatus</emphasis>
</taxonomicName>
—Martens: 6264, 70.
</paragraph>
<paragraph id="9A271257540A2E661DD9CD48FCF7CA7E" blockId="16.[151,852,432,576]" box="[245,840,554,576]" pageId="16" pageNumber="417">
2009b
<emphasis id="A8ECCE45540A2E661C17CD48FDA1CA7E" box="[315,542,554,576]" italics="true" pageId="16" pageNumber="417">
<taxonomicName id="5D9869D4540A2E661C17CD48FDBBCA7E" baseAuthorityName="Brady" baseAuthorityYear="1904" box="[315,516,554,576]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">Cypricercus inermis</taxonomicName>
</emphasis>
Savatenalinton &amp; Martens: 2.
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540A2E661DBBCD0FFA37CC40" pageId="16" pageNumber="417" type="diagnosis">
<paragraph id="9A271257540A2E661DBBCD0FFCDCCB60" blockId="16.[151,1437,621,1258]" pageId="16" pageNumber="417">
<emphasis id="A8ECCE45540A2E661DBBCD0FFEAECAB8" bold="true" box="[151,273,621,646]" inLineHeading="true" pageId="16" pageNumber="417" reason="1">Diagnosis.</emphasis>
L =
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; H = 0.450.47; carapace in lateral view elongated. Length = 2.4 times height. Valves greyish-white. LV overlapping RV ventrally, LV longer than RV anteriorly, RV longer than LV posteriorly. Valve surface with shallow pits and thin setae. A1 with small Rome organ. T1 with a-, b- and d-seta.T2 with d1, which is twice as long as d2. CR stout with four groups of setulae on the ventral margin. CR attachment with Triebel loop in the middle, and a long, well developed vb (
<bibRefCitation id="FE096FA6540A2E661E44CD9FFC49CB28" author="Brady, G. W." box="[872,1014,765,790]" pageId="16" pageNumber="417" pagination="121 - 128" refId="ref12636" refString="Brady, G. W. (1904) On Entomostraca collected in Natal by J. Gibson. Proceedings of the Zoological Society of London, 1904 121 - 128. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1904. tb 08317. x" type="book chapter" year="1904">Brady 1904</bibRefCitation>
;
<bibRefCitation id="FE096FA6540A2E66192BCD9FFA2ECB28" author="Savatenalinton, S. &amp; Martens, K." box="[1031,1425,765,790]" pageId="16" pageNumber="417" pagination="1 - 42" refId="ref14708" refString="Savatenalinton, S. &amp; Martens, K. (2009 b) Redescription of the type species of Strandesia Stuhlmann, 1888 and Cypricercus Sars, 1895 (Crustacea, Ostracoda, Cypricercinae), with a description of a new species of Cypricercus from South Africa. Zootaxa, 2007, 1 - 42." type="journal article" year="2009">Savatenalinton &amp; Martens 2009b</bibRefCitation>
). Hemipenis with triangular lateral shield and rounded distal and medial shields. Zenker organ long with 22 chitinous rosettes (for more details see
<bibRefCitation id="FE096FA6540A2E661CCCCC27FCE7CB60" author="Savatenalinton, S. &amp; Martens, K." box="[480,856,837,862]" pageId="16" pageNumber="417" pagination="1 - 42" refId="ref14708" refString="Savatenalinton, S. &amp; Martens, K. (2009 b) Redescription of the type species of Strandesia Stuhlmann, 1888 and Cypricercus Sars, 1895 (Crustacea, Ostracoda, Cypricercinae), with a description of a new species of Cypricercus from South Africa. Zootaxa, 2007, 1 - 42." type="journal article" year="2009">Savatenalinton &amp; Martens 2009b</bibRefCitation>
).
</paragraph>
<paragraph id="9A271257540A2E661DEBCC0BFA37CC40" blockId="16.[151,1437,621,1258]" pageId="16" pageNumber="417">
<emphasis id="A8ECCE45540A2E661DEBCC0BFE98CBBC" bold="true" box="[199,295,873,898]" inLineHeading="true" pageId="16" pageNumber="417" reason="1">History.</emphasis>
<bibRefCitation id="FE096FA6540A2E661C1DCC0BFE74CBBC" author="Brady, G. W." box="[305,459,873,898]" pageId="16" pageNumber="417" pagination="121 - 128" refId="ref12636" refString="Brady, G. W. (1904) On Entomostraca collected in Natal by J. Gibson. Proceedings of the Zoological Society of London, 1904 121 - 128. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1904. tb 08317. x" type="book chapter" year="1904">Brady (1904)</bibRefCitation>
named this species
<taxonomicName id="5D9869D4540A2E661F97CC08FCDBCBBF" authorityName="Brady" authorityYear="1904" box="[699,868,874,897]" class="Ostracoda" family="Cyprididae" genus="Cypris" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661F97CC08FCDBCBBF" box="[699,868,874,897]" italics="true" pageId="16" pageNumber="417">Cypris inermis</emphasis>
</taxonomicName>
and provided its first description.
<bibRefCitation id="FE096FA6540A2E6619D1CC0BFA23CBBC" author="Daday, E." box="[1277,1436,873,898]" pageId="16" pageNumber="417" pagination="113 - 176" refId="ref12970" refString="Daday, E. (1910) Die Susswasser-Mikrofauna Deutsch-Ost-Afrikas Lieferung 3. Zoologica Original-Abhandlungen aus dem Gesamtgebiete der Zoologie, 23, 113 - 176." type="journal article" year="1910">Daday (1910)</bibRefCitation>
transferred the species to the genus
<taxonomicName id="5D9869D4540A2E661F6ECCECFD19CB9B" authorityName="Vavra" authorityYear="1891" box="[578,678,910,933]" class="Ostracoda" family="Cyprididae" genus="Eucypris" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540A2E661F6ECCECFD19CB9B" box="[578,678,910,933]" italics="true" pageId="16" pageNumber="417">Eucypris</emphasis>
</taxonomicName>
.
<bibRefCitation id="FE096FA6540A2E661F94CCEFFC3DCB98" author="McKenzie, K. G." box="[696,898,909,934]" pageId="16" pageNumber="417" pagination="157 - 213" refId="ref13951" refString="McKenzie, K. G. (1971) Species List of South African freshwater Ostracoda with an appendix listing museum collections and some further determinations. Annals of the South African Museum, 57, 157 - 213." type="journal article" year="1971">McKenzie (1971)</bibRefCitation>
reported the existence of
<taxonomicName id="5D9869D4540A2E661990CCECFA2ACB9B" box="[1212,1429,910,933]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661990CCECFA2ACB9B" box="[1212,1429,910,933]" italics="true" pageId="16" pageNumber="417">Tanycypris inermis</emphasis>
</taxonomicName>
, which he found in a “small clear pool, formed by drippings of a spring in the near of (sic) Rydal Mount, Witsieshoek,
<collectingCountry id="E28F52C7540A2E661C03CCB7FE40CBD0" box="[303,511,981,1006]" name="South Africa" pageId="16" pageNumber="417">Orange Free State</collectingCountry>
”.
<bibRefCitation id="FE096FA6540A2E661F39CCB4FD7CCBD0" author="Martens, K." box="[533,707,981,1006]" pageId="16" pageNumber="417" pagination="9 - 77" refId="ref13672" refString="Martens, K. (2001) Ostracoda. In: Day, J., de Moor, I., Stewart, B. &amp; Louw, A. (Eds.), Freshwater Invertebrates of Southern Africa. Vol. 3. Crustacea II Ostracoda, Copepoda &amp; Branchiura. Water Research Commission, Pretoria, South Africa, 9 - 77." type="book chapter" year="2001">Martens (2001)</bibRefCitation>
transferred it to
<taxonomicName id="5D9869D4540A2E661EACCCB4FBB7CBD3" authorityName="Sars" authorityYear="1895" box="[896,1032,982,1005]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540A2E661EACCCB4FBB7CBD3" box="[896,1032,982,1005]" italics="true" pageId="16" pageNumber="417">Cypricercus</emphasis>
</taxonomicName>
under the assumption of synonymy with
<emphasis id="A8ECCE45540A2E661DE3CC98FE77CC2F" box="[207,456,1018,1041]" italics="true" pageId="16" pageNumber="417">
<taxonomicName id="5D9869D4540A2E661DE3CC98FE7CCC2F" authorityName="Sars" authorityYear="1895" box="[207,451,1018,1041]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="cuneatus">Cypricercus cuneatus</taxonomicName>
,
</emphasis>
a view, that was not followed in
<bibRefCitation id="FE096FA6540A2E661E1BCC9BFB01CC2C" author="Savatenalinton, S. &amp; Martens, K." box="[823,1214,1017,1042]" pageId="16" pageNumber="417" pagination="1 - 42" refId="ref14708" refString="Savatenalinton, S. &amp; Martens, K. (2009 b) Redescription of the type species of Strandesia Stuhlmann, 1888 and Cypricercus Sars, 1895 (Crustacea, Ostracoda, Cypricercinae), with a description of a new species of Cypricercus from South Africa. Zootaxa, 2007, 1 - 42." type="journal article" year="2009">Savatenalinton &amp; Martens (2009b)</bibRefCitation>
, where
<taxonomicName id="5D9869D4540A2E661838CC98FF54CC0B" baseAuthorityName="Brady" baseAuthorityYear="1904" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661838CC98FF54CC0B" italics="true" pageId="16" pageNumber="417">Cypricercus inermis</emphasis>
</taxonomicName>
and
<taxonomicName id="5D9869D4540A2E661C0ACB7CFDA4CC0B" authorityName="Sars" authorityYear="1895" box="[294,539,1054,1077]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="cuneatus">
<emphasis id="A8ECCE45540A2E661C0ACB7CFDA4CC0B" box="[294,539,1054,1077]" italics="true" pageId="16" pageNumber="417">Cypricercus cuneatus</emphasis>
</taxonomicName>
were treated as two separate species, which “can be distinguished from it [
<taxonomicName id="5D9869D4540A2E6618AECB7CFF44CC67" authorityName="Sars" authorityYear="1895" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="cuneatus">
<emphasis id="A8ECCE45540A2E6618AECB7CFF44CC67" italics="true" pageId="16" pageNumber="417">C. cuneatus</emphasis>
</taxonomicName>
] by the appearance of the unequal posterior valve margins, by the features of the caudal ramus and attachment and by the morphology of the hemipenis and the Zenkers organ“ (
<bibRefCitation id="FE096FA6540A2E66192FCB07FAC2CC40" author="Savatenalinton, S. &amp; Martens, K." box="[1027,1405,1125,1150]" pageId="16" pageNumber="417" pagination="1 - 42" refId="ref14708" refString="Savatenalinton, S. &amp; Martens, K. (2009 b) Redescription of the type species of Strandesia Stuhlmann, 1888 and Cypricercus Sars, 1895 (Crustacea, Ostracoda, Cypricercinae), with a description of a new species of Cypricercus from South Africa. Zootaxa, 2007, 1 - 42." type="journal article" year="2009">Savatenalinton &amp; Martens 2009b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D28241DC540A2E661DEBCBEBFBABCCD4" pageId="16" pageNumber="417" type="discussion">
<paragraph id="9A271257540A2E661DEBCBEBFBABCCD4" blockId="16.[151,1437,621,1258]" pageId="16" pageNumber="417">
<emphasis id="A8ECCE45540A2E661DEBCBEBFEEBCC9C" bold="true" box="[199,340,1161,1186]" inLineHeading="true" pageId="16" pageNumber="417" reason="1">Conclusion.</emphasis>
This species differs from
<taxonomicName id="5D9869D4540A2E661F55CBE8FD4CCC9F" authorityName="Triebel" authorityYear="1959" box="[633,755,1162,1185]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540A2E661F55CBE8FD4CCC9F" box="[633,755,1162,1185]" italics="true" pageId="16" pageNumber="417">Tanycypris</emphasis>
</taxonomicName>
in the presence of a well-developed vb and of a long groove in LV in
<taxonomicName id="5D9869D4540A2E661DD6CBCCFED0CCFB" baseAuthorityName="Brady" baseAuthorityYear="1904" box="[250,367,1198,1221]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661DD6CBCCFED0CCFB" box="[250,367,1198,1221]" italics="true" pageId="16" pageNumber="417">C. inermis</emphasis>
</taxonomicName>
. The width of the carapace is greater than one-third of the length in
<taxonomicName id="5D9869D4540A2E661941CBCCFB5ECCFB" baseAuthorityName="Brady" baseAuthorityYear="1904" box="[1133,1249,1198,1221]" class="Ostracoda" family="Cyprididae" genus="Cypricercus" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="A8ECCE45540A2E661941CBCCFB5ECCFB" box="[1133,1249,1198,1221]" italics="true" pageId="16" pageNumber="417">C. inermis</emphasis>
</taxonomicName>
and thus greater than in the genus
<taxonomicName id="5D9869D4540A2E661C70CBB0FE69CCD7" authorityName="Triebel" authorityYear="1959" box="[348,470,1234,1257]" class="Ostracoda" family="Cyprididae" genus="Tanycypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
<emphasis id="A8ECCE45540A2E661C70CBB0FE69CCD7" box="[348,470,1234,1257]" italics="true" pageId="16" pageNumber="417">Tanycypris</emphasis>
</taxonomicName>
(
<bibRefCitation id="FE096FA6540A2E661CC9CBB3FD3BCCD4" author="Martens, K." box="[485,644,1233,1258]" pageId="16" pageNumber="417" pagination="9 - 77" refId="ref13672" refString="Martens, K. (2001) Ostracoda. In: Day, J., de Moor, I., Stewart, B. &amp; Louw, A. (Eds.), Freshwater Invertebrates of Southern Africa. Vol. 3. Crustacea II Ostracoda, Copepoda &amp; Branchiura. Water Research Commission, Pretoria, South Africa, 9 - 77." type="book chapter" year="2001">Martens 2001</bibRefCitation>
;
<bibRefCitation id="FE096FA6540A2E661FBCCBB3FBB7CCD4" author="Savatenalinton, S. &amp; Martens, K." box="[656,1032,1233,1258]" pageId="16" pageNumber="417" pagination="1 - 42" refId="ref14708" refString="Savatenalinton, S. &amp; Martens, K. (2009 b) Redescription of the type species of Strandesia Stuhlmann, 1888 and Cypricercus Sars, 1895 (Crustacea, Ostracoda, Cypricercinae), with a description of a new species of Cypricercus from South Africa. Zootaxa, 2007, 1 - 42." type="journal article" year="2009">Savatenalinton &amp; Martens 2009b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>