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<document id="4C1692C13A3B85F3E918DA9022CCBC97" ID-DOI="10.5281/zenodo.10081384" ID-ISSN="2345-7600" ID-RefBankID="FFD5FFA3FFB6FF90FFF2A2107A6E6F66" ID-Zenodo-Dep="10081384" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616095200931" checkinUser="carolina" docAuthor="Chen, I-Shiung &amp; Kottelat, Maurice" docDate="2003" docId="747E878FFFFAFFD9EAAAFA6AFE89F9CD" docLanguage="en" docName="RafflesBZool.51.1.87-95.pdf" docOrigin="Raffles Bulletin of Zoology 51 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Rhinogobius vermiculatus Chen &amp; Kottelat 2003, new species" docType="treatment" docVersion="4" lastPageNumber="94" masterDocId="8847FFF7FFFEFFDEEA43FFABFF88FFB4" masterDocTitle="Three New Freshwater Gobies Of The Genus Rhinogobius (Teleostei: Gobiidae) From Northeastern Laos" masterLastPageNumber="95" masterPageNumber="87" pageNumber="91" updateTime="1700277477268" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CB1E9EFDEE750A174EFA9C7CF8FC6896" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CA123ABFFDE64B1E4AEAF1F4F50514B3">
<mods:title id="2C4B612680FD9546F4EEFA358B807A04">Three New Freshwater Gobies Of The Genus Rhinogobius (Teleostei: Gobiidae) From Northeastern Laos</mods:title>
</mods:titleInfo>
<mods:name id="AB6D10FA3524687BE865026286A55CF9" type="personal">
<mods:role id="42E5B4190D22F54B7630B9A394E3FF64">
<mods:roleTerm id="D7564753CC60F03F65D2C9EE9E5CA7D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="402771FD4BA004F55CEA6F6AA5E5837B">Chen, I-Shiung</mods:namePart>
</mods:name>
<mods:name id="ADAE9BCC93E04390DC953B952C630C75" type="personal">
<mods:role id="829896BF1263DC5B6488EC9147C16C6F">
<mods:roleTerm id="1B9CA21FC7375EBFB3F3244DE0B907EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC8762E575A579DB5084D66BD53E7C6D">Kottelat, Maurice</mods:namePart>
</mods:name>
<mods:typeOfResource id="4E35ED0C8C56EE5D7417770E0F44E098">text</mods:typeOfResource>
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<mods:titleInfo id="8C156C669B420A1BAC5FE4C722E0B877">
<mods:title id="BC987FEF2AE1DE92C4ABF9C93A140B48">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="FF1294C27F6CB28445F1A54C20413B9D">
<mods:date id="2DE6BF461EFB17D00DB1CD1B1FC7167E">2003</mods:date>
<mods:detail id="76DD25E923B26E8EDB8A963CF9C717C3" type="volume">
<mods:number id="F4A41502C810DD9222B097C3F5AE2BC6">51</mods:number>
</mods:detail>
<mods:detail id="6DBAA4F69A01A5F4EF9A14D1B531A9E1" type="issue">
<mods:number id="92760F5912B9848DE4C61D4328F64E46">1</mods:number>
</mods:detail>
<mods:extent id="D82DDF2B4F979571199F37A9A4328EDE" unit="page">
<mods:start id="1B8DDF662E93D6CE36D62274B7F3A429">87</mods:start>
<mods:end id="88B70A791EF4F9C483C64D8830EC045E">95</mods:end>
</mods:extent>
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<mods:classification id="E3C2C46C8D5CA90CB8ACE89C4CECDC48">journal article</mods:classification>
<mods:identifier id="308F58BEF060CE9151CFC607FF544690" type="DOI">10.5281/zenodo.10081384</mods:identifier>
<mods:identifier id="E5340C492132AA4FCFE4DCFBE64C9F74" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="465F12B1DA23BE0FCA3AE938646E5827" type="RefBankID">FFD5FFA3FFB6FF90FFF2A2107A6E6F66</mods:identifier>
<mods:identifier id="49695E26EAF9AAE9F79381C72D381A4C" type="Zenodo-Dep">10081384</mods:identifier>
</mods:mods>
<treatment id="747E878FFFFAFFD9EAAAFA6AFE89F9CD" LSID="urn:lsid:plazi:treatment:747E878FFFFAFFD9EAAAFA6AFE89F9CD" httpUri="http://treatment.plazi.org/id/747E878FFFFAFFD9EAAAFA6AFE89F9CD" lastPageId="7" lastPageNumber="94" pageId="4" pageNumber="91">
<subSubSection id="B4CD6512FFFAFFDAEAAAFA6AFD2EFA6C" box="[233,678,1472,1497]" pageId="4" pageNumber="91" type="nomenclature">
<paragraph id="FC683699FFFAFFDAEAAAFA6AFD2EFA6C" blockId="4.[233,678,1472,1528]" box="[233,678,1472,1497]" pageId="4" pageNumber="91">
<heading id="A72081F5FFFAFFDAEAAAFA6AFD2EFA6C" bold="true" box="[233,678,1472,1497]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="91" reason="2">
<emphasis id="CEA3EA8BFFFAFFDAEAAAFA6AFD2EFA6C" bold="true" box="[233,678,1472,1497]" pageId="4" pageNumber="91">
<taxonomicName id="3BD74D1AFFFAFFDAEAAAFA6AFD98FA6D" ID-CoL="4S937" authority="Chen &amp; Kottelat, 2003" authorityName="Chen &amp; Kottelat" authorityYear="2003" box="[233,528,1473,1497]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="vermiculatus" status="sp. nov.">
<emphasis id="CEA3EA8BFFFAFFDAEAAAFA6AFD98FA6D" bold="true" box="[233,528,1473,1497]" italics="true" pageId="4" pageNumber="91">Rhinogobius vermiculatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D59057F0FFFAFFDAE863FA6BFD2EFA6C" box="[544,678,1472,1496]" pageId="4" pageNumber="91" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFFAFFDAEB3FFA4AFD9BFA4C" box="[380,531,1505,1528]" pageId="4" pageNumber="91" type="description">
<paragraph id="FC683699FFFAFFDAEB3FFA4AFD9BFA4C" blockId="4.[233,678,1472,1528]" box="[380,531,1505,1528]" pageId="4" pageNumber="91">
(
<figureCitation id="64EC2A1CFFFAFFDAEBC7FA4AFE5AFA4C" box="[388,466,1505,1528]" captionStart="Fig" captionStartId="5.[113,148,2065,2086]" captionTargetBox="[112,753,1522,2048]" captionTargetId="figure-82@5.[112,753,1522,1788]" captionTargetPageId="5" captionText="Fig. 3. Rhinogobius vermiculatus, CMK 15306, paratypes, (a) male, 34.1 mm SL, and (b) female, 41.1 mm SL; Laos: Nam Hang." figureDoi="http://doi.org/10.5281/zenodo.10081390" httpUri="https://zenodo.org/record/10081390/files/figure.png" pageId="4" pageNumber="91">Figs. 3</figureCitation>
,
<figureCitation id="64EC2A1CFFFAFFDAEBA3FA4AFE67FA4C" box="[480,495,1505,1528]" captionStart="Fig" captionStartId="6.[136,171,1100,1121]" captionTargetBox="[211,1475,212,1067]" captionTargetId="figure-10@6.[824,1485,275,1052]" captionTargetPageId="6" captionText="Fig. 6. Rhinogobius vermiculatus, CMK 15306, 47.7 mm SL; head lateral-line system. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10081394" httpUri="https://zenodo.org/record/10081394/files/figure.png" pageId="4" pageNumber="91">6</figureCitation>
,
<figureCitation id="64EC2A1CFFFAFFDAEBBEFA4AFD83FA4C" box="[509,523,1505,1528]" captionStart="Fig" captionStartId="6.[835,871,2065,2086]" captionText="Fig. 9. Rhinogobius vermiculatus, CMK 15306, 47.7 mm SL; cephalic colour pattern of male. Scale bar = 1 mm." pageId="4" pageNumber="91">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFFAFFDAEACBF98BFECEF8C6" pageId="4" pageNumber="91" type="materials_examined">
<paragraph id="FC683699FFFAFFDAEACBF98BFECEF938" blockId="4.[136,776,1568,1676]" pageId="4" pageNumber="91">
<emphasis id="CEA3EA8BFFFAFFDAEACBF98BFED4F981" bold="true" box="[136,348,1568,1589]" italics="true" pageId="4" pageNumber="91">Material examined. –</emphasis>
<materialsCitation id="4CBF3CC4FFFAFFDAEB21F98BFECEF938" accessionNumber="ZRC 46583" collectingDate="1999-05-05" collectorName="M. Kottelat" country="Laos" latitude="20.156666" location="Nam Xam in Ban Houtangoua" longLatPrecision="21" longitude="104.54722" municipality="Houay Tangoua" pageId="4" pageNumber="91" specimenCount="1" stateProvince="Houaphan" typeStatus="holotype">
<typeStatus id="236C883BFFFAFFDAEB21F98BFE35F981" box="[354,445,1568,1589]" pageId="4" pageNumber="91" type="holotype">Holotype</typeStatus>
– ZRC 46583, 39.0 mm SL,
<collectingCountry id="84C07609FFFAFFDAE892F98BFC8CF981" box="[721,772,1568,1589]" name="Laos" pageId="4" pageNumber="91">Laos</collectingCountry>
:
<collectingRegion id="3E13F87BFFFAFFDAEACBF996FEAFF9E6" box="[136,295,1597,1618]" country="Laos" name="Houaphan" pageId="4" pageNumber="91">Houaphan Prov.</collectingRegion>
,
<collectingMunicipality id="1C0CACE3FFFAFFDAEB6DF996FE42F9E6" box="[302,458,1597,1618]" pageId="4" pageNumber="91">Houay Tangoua</collectingMunicipality>
, small stream entering
<location id="F9086042FFFAFFDAE8E7F996FECBF9DB" LSID="urn:lsid:plazi:treatment:747E878FFFFAFFD9EAAAFA6AFE89F9CD:F9086042FFFAFFDAE8E7F996FECBF9DB" country="Laos" latitude="20.156666" longLatPrecision="21" longitude="104.54722" municipality="Houay Tangoua" name="Nam Xam in Ban Houtangoua" pageId="4" pageNumber="91" stateProvince="Houaphan">Nam Xam in Ban Houtangoua</location>
,
<geoCoordinate id="99E3505EFFFAFFDAEB0FF9F1FE46F9DB" box="[332,462,1625,1647]" degrees="20" direction="north" minutes="09" orientation="latitude" pageId="4" pageNumber="91" precision="15" seconds="24" value="20.156666">
20
<superScript id="0BA29BD1FFFAFFDAEB27F9F2FEE3F9D1" attach="left" box="[356,363,1625,1637]" fontSize="5" pageId="4" pageNumber="91">o</superScript>
09’24’’ N
</geoCoordinate>
<geoCoordinate id="99E3505EFFFAFFDAEB90F9F1FDE9F9DB" box="[467,609,1625,1647]" degrees="104" direction="east" minutes="32" orientation="longitude" pageId="4" pageNumber="91" precision="15" seconds="50" value="104.54722">
104
<superScript id="0BA29BD1FFFAFFDAEBB4F9F2FE76F9D1" attach="left" box="[503,510,1625,1637]" fontSize="5" pageId="4" pageNumber="91">o</superScript>
32’50’’ E
</geoCoordinate>
, coll.
<collectorName id="5122534FFFFAFFDAE8DAF9F1FC8FF9DB" box="[665,775,1626,1647]" pageId="4" pageNumber="91">M. Kottelat</collectorName>
et al.,
<date id="88691059FFFAFFDAEA8BF9DCFECBF938" box="[200,323,1655,1676]" pageId="4" pageNumber="91" value="1999-05-05">
<collectingDate id="982DE9B1FFFAFFDAEA8BF9DCFECBF938" box="[200,323,1655,1676]" pageId="4" pageNumber="91" value="1999-05-05">5 May.1999</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="FC683699FFFAFFDAEACBF91BFECEF8C6" blockId="4.[136,775,1712,1906]" pageId="4" pageNumber="91">
<materialsCitation id="4CBF3CC4FFFAFFDAEACBF91BFDCAF956" accessionNumber="ZRC 47676" pageId="4" pageNumber="91" specimenCount="6" typeStatus="paratype">
<typeStatus id="236C883BFFFAFFDAEACBF91BFF66F971" box="[136,238,1712,1733]" pageId="4" pageNumber="91" type="paratype">Paratypes</typeStatus>
– ZRC 47676,
<specimenCount id="EAD1FD10FFFAFFDAEBE3F91BFDAFF971" box="[416,551,1712,1733]" pageId="4" pageNumber="91" type="generic">2 specimens</specimenCount>
, LARReC, uncat.,
<specimenCount id="EAD1FD10FFFAFFDAE8B8F91BFF74F956" pageId="4" pageNumber="91" type="generic">2 specimens</specimenCount>
, NMMBA 464,
<specimenCount id="EAD1FD10FFFAFFDAEBF5F966FDCAF956" box="[438,578,1741,1762]" pageId="4" pageNumber="91" type="generic">2 specimens</specimenCount>
</materialsCitation>
,
<materialsCitation id="4CBF3CC4FFFAFFDAE811F966FD32F94B" accessionNumber="CMK 15306" collectingDate="1999-05-05" collectorName="M. Kottelat" country="Laos" latitude="20.156666" location="Nam Xam in Ban Houtangoua" longLatPrecision="21" longitude="104.54722" municipality="Houay Tangoua" pageId="4" pageNumber="91" specimenCount="10" stateProvince="Houaphan" typeStatus="paratype">
CMK 15306,
<specimenCount id="EAD1FD10FFFAFFDAE8AEF966FF78F94B" pageId="4" pageNumber="91" type="generic">10 specimens</specimenCount>
,
<quantity id="3B2F9B7CFFFAFFDAEABEF941FE29F94B" box="[253,417,1770,1791]" metricMagnitude="-2" metricUnit="m" metricValue="3.2150000000000007" metricValueMax="4.7700000000000005" metricValueMin="1.6600000000000001" pageId="4" pageNumber="91" unit="mm" value="32.15" valueMax="47.7" valueMin="16.6">16.6 – 47.7 mm</quantity>
SL, same data as holotype
</materialsCitation>
.
<materialsCitation id="4CBF3CC4FFFAFFDAE893F941FECBF8C6" accessionNumber="CMK 15321" collectingDate="1999-05-06" collectorName="M. Kottelat" country="Laos" county="Nam Hang" latitude="19.985" location="Ban Tao" longLatPrecision="21" longitude="104.674995" municipality="Xam Tai" pageId="4" pageNumber="91" specimenCount="4" stateProvince="Houaphan" typeStatus="paratype">
CMK 15321,
<specimenCount id="EAD1FD10FFFAFFDAEA91F8ACFEC5F8A8" box="[210,333,1799,1820]" pageId="4" pageNumber="91" type="generic">4 specimens</specimenCount>
,
<quantity id="3B2F9B7CFFFAFFDAEB1BF8ACFE7CF8A8" box="[344,500,1799,1820]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" metricValueMax="3.41" metricValueMin="2.95" pageId="4" pageNumber="91" unit="mm" value="31.8" valueMax="34.1" valueMin="29.5">29.5 – 34.1 mm</quantity>
SL,
<collectingCountry id="84C07609FFFAFFDAE867F8ACFDDFF8A8" box="[548,599,1799,1820]" name="Laos" pageId="4" pageNumber="91">Laos</collectingCountry>
:
<collectingRegion id="3E13F87BFFFAFFDAE822F8ACFC8CF8A8" box="[609,772,1799,1820]" country="Laos" name="Houaphan" pageId="4" pageNumber="91">Houaphan Prov.</collectingRegion>
,
<collectingCounty id="15094E15FFFAFFDAEACBF88FFF7BF88D" box="[136,243,1828,1849]" pageId="4" pageNumber="91">Nam Hang</collectingCounty>
, small stream on road from
<collectingMunicipality id="1C0CACE3FFFAFFDAE843F88FFDDDF88D" box="[512,597,1828,1849]" pageId="4" pageNumber="91">Xam Tai</collectingMunicipality>
to
<location id="F9086042FFFAFFDAE830F88FFD4FF88D" LSID="urn:lsid:plazi:treatment:747E878FFFFAFFD9EAAAFA6AFE89F9CD:F9086042FFFAFFDAE830F88FFD4FF88D" box="[627,711,1828,1849]" country="Laos" county="Nam Hang" latitude="19.985" longLatPrecision="21" longitude="104.674995" municipality="Xam Tai" name="Ban Tao" pageId="4" pageNumber="91" stateProvince="Houaphan">Ban Tao</location>
, about
<quantity id="3B2F9B7CFFFAFFDAEACBF8EBFF30F8E1" box="[136,184,1856,1877]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="91" unit="km" value="6.0">6 km</quantity>
from
<location id="F9086042FFFAFFDAEAB2F8EBFEC0F8E1" LSID="urn:lsid:plazi:treatment:747E878FFFFAFFD9EAAAFA6AFE89F9CD:F9086042FFFAFFDAEAB2F8EBFEC0F8E1" box="[241,328,1856,1877]" country="Laos" county="Nam Hang" latitude="19.985" longLatPrecision="21" longitude="104.674995" municipality="Xam Tai" name="Xam Tai" pageId="4" pageNumber="91" stateProvince="Houaphan">Xam Tai</location>
,
<geoCoordinate id="99E3505EFFFAFFDAEB12F8EBFE58F8E1" box="[337,464,1855,1877]" degrees="19" direction="north" minutes="59" orientation="latitude" pageId="4" pageNumber="91" precision="15" seconds="06" value="19.985">
19
<superScript id="0BA29BD1FFFAFFDAEB2AF894FEF8F8FF" attach="left" box="[361,368,1855,1867]" fontSize="5" pageId="4" pageNumber="91">o</superScript>
59’06’’N
</geoCoordinate>
<geoCoordinate id="99E3505EFFFAFFDAEB96F8EBFDD7F8E1" box="[469,607,1855,1877]" degrees="104" direction="east" minutes="40" orientation="longitude" pageId="4" pageNumber="91" precision="15" seconds="30" value="104.674995">
104
<superScript id="0BA29BD1FFFAFFDAEBBAF894FD88F8FF" attach="left" box="[505,512,1855,1867]" fontSize="5" pageId="4" pageNumber="91">o</superScript>
40’30’’E
</geoCoordinate>
, coll.
<collectorName id="5122534FFFFAFFDAE8DBF8EBFC8EF8E1" box="[664,774,1856,1877]" pageId="4" pageNumber="91">M. Kottelat</collectorName>
et al.,
<date id="88691059FFFAFFDAEA8BF8F6FECBF8C6" box="[200,323,1885,1906]" pageId="4" pageNumber="91" value="1999-05-06">
<collectingDate id="982DE9B1FFFAFFDAEA8BF8F6FECBF8C6" box="[200,323,1885,1906]" pageId="4" pageNumber="91" value="1999-05-06">6 May.1999</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFFAFFDAEACBF831FA4BFEEC" pageId="4" pageNumber="91" type="diagnosis">
<paragraph id="FC683699FFFAFFDAEACBF831FC8FF785" blockId="4.[136,775,1946,2097]" pageId="4" pageNumber="91">
<emphasis id="CEA3EA8BFFFAFFDAEACBF831FDA3F806" box="[136,555,1946,1970]" italics="true" pageId="4" pageNumber="91">
<emphasis id="CEA3EA8BFFFAFFDAEACBF831FE87F806" bold="true" box="[136,271,1946,1970]" italics="true" pageId="4" pageNumber="91">Diagnosis. –</emphasis>
<taxonomicName id="3BD74D1AFFFAFFDAEB57F830FDA3F806" authorityName="Chen &amp; Kottelat" authorityYear="2003" box="[276,555,1947,1970]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="vermiculatus">Rhinogobius vermiculatus</taxonomicName>
</emphasis>
can be distinguished from all other species of the genus by the combination of the following characters: second dorsal fin rays modally I/ 9; anal fin rays I/7; pectoral fin rays modally 16; longitudinal scale series 32-34; vertebral counts 10 + 18 = 28; first dorsal
</paragraph>
<paragraph id="FC683699FFFAFFDAE900FF6AFA4BFEEC" blockId="4.[835,1475,193,344]" pageId="4" pageNumber="91">fin with rays III and IV longest; body with 6-7 dark brown blotches and female with 5 longitudinal dark brown lines. Opercle with vermiculate lines against brown background and branchiostegal membrane with small dark greyish brown spots in male; and caudal fin base with a dark brown mark.</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFFAFFD9E900FE2AFF68FC4D" lastPageId="7" lastPageNumber="94" pageId="4" pageNumber="91" type="description">
<paragraph id="FC683699FFFAFFDAE900FE2AFA4BFD4C" blockId="4.[835,1475,385,760]" pageId="4" pageNumber="91">
<emphasis id="CEA3EA8BFFFAFFDAE900FE2AFC57FE2D" bold="true" box="[835,991,385,409]" italics="true" pageId="4" pageNumber="91">Description. –</emphasis>
Body cylindrical anteriorly and compressed posteriorly. Body proportions in Table 1. Head moderately large, and depressed in adult male. Eye large, dorsolateral. Snout pointed and longer in male. Bony interorbital quite narrow. Cheek very fleshy in male. Lips thick, and with projecting lateral side of upper lip (especially in male). Mouth oblique, maxillary extending to vertical through middle of orbit. Both jaws with 3-4 rows of conical teeth, outer rows enlarged. Tongue tip rounded, entire. Anterior nostril a short tube, posterior one merely a very shallow tube. Gill-opening extending to vertical through posterior extremity of preopercle. Isthmus broad. 10 + 18 = 28 (in
<specimenCount id="EAD1FD10FFFAFFDAEF63FD4AFA3FFD4C" box="[1312,1463,737,760]" pageId="4" pageNumber="91" type="generic">10 specimens</specimenCount>
).
</paragraph>
<paragraph id="FC683699FFFAFFDAE900FC8AFA9DFBEC" blockId="4.[835,1475,801,1112]" pageId="4" pageNumber="91">Fins. D1 V-VII (mode VI); D2 I/8-9 (mode 9); A I/7; P 15- 18 (mode 16); V I/5+I/5 (frequency distribution in Table 2). D1 rays III and IV longest and somewhat filamentous in male, with D1 rear tip reaching 1st or 2nd branched rays of D2 when depressed in male, not extending to D2 origin in female. D2 and A rear tips not reaching procurrent rays of C. Origin of A inserted at vertical line between 2nd and 3rd branched rays of D2. P oblong, rear tip not extending to anus. V disc rounded, spinous rays with pointed membranous lobes. C elliptical with rounded rear edge.</paragraph>
<paragraph id="FC683699FFFAFFDAE900FB2AFAEBFA6C" blockId="4.[835,1475,1153,1496]" pageId="4" pageNumber="91">Scales. Lateral body with moderately large ctenoid scales, anterior predorsal area naked, posterior predorsal region and belly scales cycloid, with 7-11 middle series of predorsal scales; scales in longitudinal series 32-34 (mode 33); transverse series 9-10; series between D1 origin and upper P origin 6-7 (frequency distribution in Table 2). Head including opercle, preopercle, and prepelvic area naked. Predorsal squamation with trifurcate anterior edge, anterior tip of middle series reaching vertical through middle of opercle; both tips of lateral squamation extending forward to above pore of posterior oculoscapular canal.</paragraph>
<paragraph id="FC683699FFFAFFDAE900F9AAFA4AF9CC" blockId="4.[835,1474,1537,1976]" pageId="4" pageNumber="91">Head lateral-line system. Canals. Nasal extension of anterior oculoscapular canal with terminal pores located at middle between anterior and posterior nostril. Anterior interorbital sections of oculoscapular canal separated, with paired pores</paragraph>
<paragraph id="FC683699FFFAFFDAE911F92AFB3BF80C" blockId="4.[835,1474,1537,1976]" pageId="4" pageNumber="91">
. A single pore
<emphasis id="CEA3EA8BFFFAFFDAEE41F92AFB86F92C" box="[1026,1038,1665,1688]" italics="true" pageId="4" pageNumber="91">k</emphasis>
in posterior interorbital region. Pore present at posteriodorsal edge of eye. Gap between anterior and posterior oculoscapular canals somewhat smaller than length of posterior oculoscapular canal. Preopercular canal present with pores
<emphasis id="CEA3EA8BFFFAFFDAEE61F8AAFBA4F8AC" box="[1058,1068,1793,1816]" italics="true" pageId="4" pageNumber="91">r</emphasis>
, and. Sensory papillae. Row
<emphasis id="CEA3EA8BFFFAFFDAEFF6F8AAFA4AF8AC" box="[1461,1474,1793,1816]" italics="true" pageId="4" pageNumber="91">a</emphasis>
extending to vertical through middle of orbit. Length of row
<emphasis id="CEA3EA8BFFFAFFDAE900F8EAFCD8F8EC" box="[835,848,1857,1880]" italics="true" pageId="4" pageNumber="91">b</emphasis>
slightly shorter than orbit. Rows
<emphasis id="CEA3EA8BFFFAFFDAEE98F8EAFB6FF8EC" box="[1243,1255,1857,1880]" italics="true" pageId="4" pageNumber="91">c</emphasis>
,
<emphasis id="CEA3EA8BFFFAFFDAEEBBF8EAFA8DF8EC" box="[1272,1285,1857,1880]" italics="true" pageId="4" pageNumber="91">d</emphasis>
long, and row
<emphasis id="CEA3EA8BFFFAFFDAEFF5F8EAFA4AF8EC" box="[1462,1474,1857,1880]" italics="true" pageId="4" pageNumber="91">c</emphasis>
reaching or slightly beyond vertical of pore a. A single
<emphasis id="CEA3EA8BFFFAFFDAEFEAF8CAFA4AF8CC" box="[1449,1474,1889,1912]" italics="true" pageId="4" pageNumber="91">cp</emphasis>
papilla. Row
<emphasis id="CEA3EA8BFFFAFFDAE99DF82AFC6DF82C" box="[990,997,1921,1944]" italics="true" pageId="4" pageNumber="91">f</emphasis>
paired. Opercular rows
<emphasis id="CEA3EA8BFFFAFFDAEEBDF82AFA9BF82C" box="[1278,1299,1921,1944]" italics="true" pageId="4" pageNumber="91">ot</emphasis>
and
<emphasis id="CEA3EA8BFFFAFFDAEF13F82AFAEDF82C" box="[1360,1381,1921,1944]" italics="true" pageId="4" pageNumber="91">oi</emphasis>
slightly separated. Other details in
<figureCitation id="64EC2A1CFFFAFFDAEE2FF80AFB38F80C" box="[1132,1200,1953,1976]" captionStart="Fig" captionStartId="5.[813,849,2064,2085]" captionTargetBox="[897,1368,1158,2011]" captionTargetId="figure-125@5.[884,1383,1158,2019]" captionTargetPageId="5" captionText="Fig. 5. Rhinogobius nammaensis, ZRC 46582, 43.0 mm SL; head lateral-line system. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10081392" httpUri="https://zenodo.org/record/10081392/files/figure.png" pageId="4" pageNumber="91">Fig. 5</figureCitation>
.
</paragraph>
<paragraph id="FC683699FFFAFFDAE900F84AFA4AF78C" blockId="4.[835,1474,2017,2104]" pageId="4" pageNumber="91">
<emphasis id="CEA3EA8BFFFAFFDAE900F84AFBD6F84D" bold="true" box="[835,1118,2017,2041]" italics="true" pageId="4" pageNumber="91">Colouration in alcohol. –</emphasis>
Body yellowish brown with 6-7 dark brown blotches; 5 longitudinal dark brown lines in female, indistinct markings except darker margins of lateral
</paragraph>
<caption id="A8A86611FFFBFFDBEA32FD4AFD3BFCA6" ID-DOI="http://doi.org/10.5281/zenodo.10081386" ID-Zenodo-Dep="10081386" httpUri="https://zenodo.org/record/10081386/files/figure.png" pageId="5" pageNumber="92" startId="5.[113,148,737,758]" targetBox="[113,753,190,720]" targetPageId="5">
<paragraph id="FC683699FFFBFFDBEA32FD4AFD3BFCA6" blockId="5.[113,752,682,786]" pageId="5" pageNumber="92">
Fig. 1.
<taxonomicName id="3BD74D1AFFFBFFDBEAFBFD4AFE32FD42" authority=", CMK" authorityName="CMK" box="[184,442,737,758]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="milleri">
<emphasis id="CEA3EA8BFFFBFFDBEAFBFD4AFEFFFD42" box="[184,375,737,758]" italics="true" pageId="5" pageNumber="92">Rhinogobius milleri</emphasis>
, CMK
</taxonomicName>
15249, paratypes, (a) male, 39.2 mm SL, and (b) female, 45.8 mm SL; Laos: Nam Kuang.
</paragraph>
</caption>
<caption id="A8A86611FFFBFFDBE96EFBB0FB6AFBF9" ID-DOI="http://doi.org/10.5281/zenodo.10081388" ID-Zenodo-Dep="10081388" httpUri="https://zenodo.org/record/10081388/files/figure.png" pageId="5" pageNumber="92" startId="5.[813,849,1051,1072]" targetBox="[905,1350,186,1018]" targetPageId="5">
<paragraph id="FC683699FFFBFFDBE96EFBB0FB6AFBF9" blockId="5.[813,1452,1051,1101]" pageId="5" pageNumber="92">
Fig. 4.
<taxonomicName id="3BD74D1AFFFBFFDBE937FBB0FBB0FB84" box="[884,1080,1051,1072]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="milleri">
<emphasis id="CEA3EA8BFFFBFFDBE937FBB0FBB0FB84" box="[884,1080,1051,1072]" italics="true" pageId="5" pageNumber="92">Rhinogobius milleri</emphasis>
</taxonomicName>
, holotype, ZRC 46581, 39.8 mm SL; head lateral-line system. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="FC683699FFFBFFDBE88EFA9CFD69FAE7" blockId="5.[113,753,1335,1467]" box="[717,737,1335,1363]" pageId="5" pageNumber="92">
<emphasis id="CEA3EA8BFFFBFFDBE88EFA9CFD69FAE7" bold="true" box="[717,737,1335,1363]" pageId="5" pageNumber="92">b</emphasis>
</paragraph>
<paragraph id="FC683699FFFBFFDBEA32FAC6FE83FA0F" blockId="5.[113,753,1335,1467]" pageId="5" pageNumber="92">
Fig. 2.
<taxonomicName id="3BD74D1AFFFBFFDBEAF5FAC6FE27FA36" authorityName="Chen &amp; Kottelat" authorityYear="2003" box="[182,431,1389,1410]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="nammaensis">
<emphasis id="CEA3EA8BFFFBFFDBEAF5FAC6FE27FA36" box="[182,431,1389,1410]" italics="true" pageId="5" pageNumber="92">Rhinogobius nammaensis</emphasis>
</taxonomicName>
, (a) ZRC 46582,
<typeStatus id="236C883BFFFBFFDBE81AFAC6FD39FA36" box="[601,689,1389,1410]" pageId="5" pageNumber="92" type="holotype">holotype</typeStatus>
, male, 43.0 mm SL, and (b) CMK 15370, female,
<quantity id="3B2F9B7CFFFBFFDBE86FFA22FD0EFA2A" box="[556,646,1417,1438]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="5" pageNumber="92" unit="mm" value="36.1">36.1 mm</quantity>
SL;
<collectingCountry id="84C07609FFFBFFDBE8F9FA22FD65FA2A" box="[698,749,1417,1438]" name="Laos" pageId="5" pageNumber="92">Laos</collectingCountry>
: Nam Ma basin.
</paragraph>
<caption id="A8A86611FFFBFFDBEA32F7BAFD52F7F7" ID-DOI="http://doi.org/10.5281/zenodo.10081390" ID-Zenodo-Dep="10081390" httpUri="https://zenodo.org/record/10081390/files/figure.png" pageId="5" pageNumber="92" startId="5.[113,148,2065,2086]" targetBox="[112,753,1522,2048]" targetPageId="5">
<paragraph id="FC683699FFFBFFDBEA32F7BAFD52F7F7" blockId="5.[113,753,2011,2115]" pageId="5" pageNumber="92">
Fig. 3.
<taxonomicName id="3BD74D1AFFFBFFDBEAF0F7BAFE64F792" authority=", CMK" authorityName="CMK" box="[179,492,2065,2086]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="CEA3EA8BFFFBFFDBEAF0F7BAFE23F792" box="[179,427,2065,2086]" italics="true" pageId="5" pageNumber="92">Rhinogobius vermiculatus</emphasis>
, CMK
</taxonomicName>
15306, paratypes, (a) male, 34.1 mm SL, and (b) female, 41.1 mm SL; Laos: Nam Hang.
</paragraph>
</caption>
<caption id="A8A86611FFFBFFDBE96EF7BBFB24F7F6" ID-DOI="http://doi.org/10.5281/zenodo.10081392" ID-Zenodo-Dep="10081392" httpUri="https://zenodo.org/record/10081392/files/figure.png" pageId="5" pageNumber="92" startId="5.[813,849,2064,2085]" targetBox="[897,1368,1158,2011]" targetPageId="5">
<paragraph id="FC683699FFFBFFDBE96EF7BBFB24F7F6" blockId="5.[813,1452,2064,2114]" pageId="5" pageNumber="92">
Fig. 5.
<taxonomicName id="3BD74D1AFFFBFFDBE931F7BAFB20F791" authority=", ZRC" authorityName="ZRC" box="[882,1192,2064,2086]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="nammaensis">
<emphasis id="CEA3EA8BFFFBFFDBE931F7BAFBE4F792" box="[882,1132,2065,2086]" italics="true" pageId="5" pageNumber="92">Rhinogobius nammaensis</emphasis>
, ZRC
</taxonomicName>
46582, 43.0 mm SL; head lateral-line system. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="A8A86611FFF8FFD8EACBFBE7FD8FFBCA" ID-DOI="http://doi.org/10.5281/zenodo.10081394" ID-Zenodo-Dep="10081394" httpUri="https://zenodo.org/record/10081394/files/figure.png" pageId="6" pageNumber="93" startId="6.[136,171,1100,1121]" targetBox="[211,1475,212,1067]" targetPageId="6">
<paragraph id="FC683699FFF8FFD8EACBFBE7FD8FFBCA" blockId="6.[136,775,1100,1150]" pageId="6" pageNumber="93">
Fig. 6.
<taxonomicName id="3BD74D1AFFF8FFD8EA88FBE7FD82FBD5" authority=", CMK" authorityName="CMK" box="[203,522,1100,1121]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="CEA3EA8BFFF8FFD8EA88FBE7FE4FFBD5" box="[203,455,1100,1121]" italics="true" pageId="6" pageNumber="93">Rhinogobius vermiculatus</emphasis>
, CMK
</taxonomicName>
15306, 47.7 mm SL; head lateral-line system. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="A8A86611FFF8FFD8E900FBE7FAE7FBC9" pageId="6" pageNumber="93">
<paragraph id="FC683699FFF8FFD8E900FBE7FAE7FBC9" blockId="6.[835,1475,1100,1149]" pageId="6" pageNumber="93">
Fig. 8.
<taxonomicName id="3BD74D1AFFF8FFD8E9CBFBE7FB09FBD5" box="[904,1153,1100,1121]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="nammaensis">
<emphasis id="CEA3EA8BFFF8FFD8E9CBFBE7FB09FBD5" box="[904,1153,1100,1121]" italics="true" pageId="6" pageNumber="93">Rhinogobius nammaensis</emphasis>
</taxonomicName>
, holotype, ZRC 46582, 43.0 mm SL; cephalic colour pattern of male. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="A8A86611FFF8FFD8EACBF7BAFD01F7F7" ID-DOI="http://doi.org/10.5281/zenodo.10081396" ID-Zenodo-Dep="10081396" httpUri="https://zenodo.org/record/10081396/files/figure.png" pageId="6" pageNumber="93" startId="6.[136,172,2065,2086]" targetBox="[139,1404,1179,2039]" targetPageId="6">
<paragraph id="FC683699FFF8FFD8EACBF7BAFD01F7F7" blockId="6.[136,775,2065,2115]" pageId="6" pageNumber="93">
Fig. 7.
<taxonomicName id="3BD74D1AFFF8FFD8EA93F7BAFE1BF792" box="[208,403,2065,2086]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="milleri">
<emphasis id="CEA3EA8BFFF8FFD8EA93F7BAFE1BF792" box="[208,403,2065,2086]" italics="true" pageId="6" pageNumber="93">Rhinogobius milleri</emphasis>
</taxonomicName>
, holotype, ZRC 46581, 39.8 mm SL; cephalic colour pattern of male. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="A8A86611FFF8FFD8E900F7BAFACCF7F7" pageId="6" pageNumber="93">
<paragraph id="FC683699FFF8FFD8E900F7BAFACCF7F7" blockId="6.[835,1474,2065,2115]" pageId="6" pageNumber="93">
Fig. 9.
<taxonomicName id="3BD74D1AFFF8FFD8E9D2F7BAFB6AF792" authority=", CMK" authorityName="CMK" box="[913,1250,2065,2086]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="CEA3EA8BFFF8FFD8E9D2F7BAFB12F792" box="[913,1178,2065,2086]" italics="true" pageId="6" pageNumber="93">Rhinogobius vermiculatus</emphasis>
, CMK
</taxonomicName>
15306, 47.7 mm SL; cephalic colour pattern of male. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="FC683699FFF9FFD9EA32FF69FE76FDED" blockId="7.[113,753,194,601]" pageId="7" pageNumber="94">scales in larger male. Predorsal region with several short, dark brown bars. Cheek with many irregular brown stripes or spots and a horizontal brownish black bar crossing anterior part, that of lower half near preopercle with few white rounded spots. Opercle with 8-10 white vermiculations on brown background on lower half in male, usually indistinct with 3-4 larger spots or plain in female. Snout with a pair of brownish black lines uniting at anterior tip. A brownish black line behind orbit along lateral section of anterior oculoscapular canal. A somewhat squarish, brownish grey blotch below eye. Branchiostegal membrane scattered with dark greyish spots (which may have been brownish red when fresh) in male, unmarked in female.</paragraph>
<paragraph id="FC683699FFF9FFD9EA32FD29FF68FC4D" blockId="7.[113,753,642,1017]" pageId="7" pageNumber="94">First dorsal fin membranes between three anterior rays with a black spot, rest of fin with 2-3 rows of dark brown spots in male, with 3-4 rows of dark spots in female. Caudal fin greyish with 4-6 vertical rows of dark brown spots in male, with 4-5 rows of greyish to dark brown spots in female. Caudal fin base with a vertical, dark brown curved bar. Anal fin greyish black on distal half with narrow white margin in male, uniform whitish in female. Pectoral fin with a conspicuous black spot on upper part of base, and a brown blotch on lower portion; basal region with a semicircular mark. Pelvic fin greyish near basal portion in male whitish in female.</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFF9FFD9EA32FB89FDC8FB2D" pageId="7" pageNumber="94" type="distribution">
<paragraph id="FC683699FFF9FFD9EA32FB89FDC8FB2D" blockId="7.[113,752,1058,1177]" pageId="7" pageNumber="94">
<emphasis id="CEA3EA8BFFF9FFD9EA32FB89FDA3FB8E" box="[113,555,1058,1082]" italics="true" pageId="7" pageNumber="94">
<emphasis id="CEA3EA8BFFF9FFD9EA32FB89FE87FB8E" bold="true" box="[113,271,1058,1082]" italics="true" pageId="7" pageNumber="94">Distribution. –</emphasis>
<taxonomicName id="3BD74D1AFFF9FFD9EB56FB88FDA3FB8E" box="[277,555,1059,1082]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="species" species="vermiculatus">Rhinogobius vermiculatus</taxonomicName>
</emphasis>
has been collected only in small forest streams, with stony substrate and rapid current of the Nan Xam basin, northeastern
<collectingCountry id="84C07609FFF9FFD9E876FBC9FDE4FBCD" box="[565,620,1122,1145]" name="Laos" pageId="7" pageNumber="94">Laos</collectingCountry>
. Its presence is expected in the same basin in
<collectingCountry id="84C07609FFF9FFD9EB9FFB29FDB4FB2D" box="[476,572,1154,1177]" name="Vietnam" pageId="7" pageNumber="94">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFF9FFD9EA32FB69FDE6FB4D" pageId="7" pageNumber="94" type="etymology">
<paragraph id="FC683699FFF9FFD9EA32FB69FDE6FB4D" blockId="7.[113,752,1218,1273]" pageId="7" pageNumber="94">
<emphasis id="CEA3EA8BFFF9FFD9EA32FB69FE8AFB6E" bold="true" box="[113,258,1218,1242]" italics="true" pageId="7" pageNumber="94">Etymology. –</emphasis>
From the Latin,
<taxonomicName id="3BD74D1AFFF9FFD9EBFAFB68FDCCFB6E" box="[441,580,1219,1242]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="CEA3EA8BFFF9FFD9EBFAFB68FDCCFB6E" box="[441,580,1219,1242]" italics="true" pageId="7" pageNumber="94">vermiculatus</emphasis>
</taxonomicName>
(vermiculated), refers to the coloration pattern on the opercle.
</paragraph>
</subSubSection>
<subSubSection id="B4CD6512FFF9FFD9EA32FA89FE89F9CD" pageId="7" pageNumber="94" type="discussion">
<paragraph id="FC683699FFF9FFD9EA32FA89FE89F9CD" blockId="7.[113,753,1314,1657]" pageId="7" pageNumber="94">
<emphasis id="CEA3EA8BFFF9FFD9EA32FA89FD87FA8E" box="[113,527,1314,1338]" italics="true" pageId="7" pageNumber="94">
<emphasis id="CEA3EA8BFFF9FFD9EA32FA89FF66FA8E" bold="true" box="[113,238,1314,1338]" italics="true" pageId="7" pageNumber="94">Remarks. –</emphasis>
<taxonomicName id="3BD74D1AFFF9FFD9EAB6FA88FD87FA8E" box="[245,527,1315,1338]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="species" species="vermiculatus">Rhinogobius vermiculatus</taxonomicName>
</emphasis>
is more similar to
<taxonomicName id="3BD74D1AFFF9FFD9E89AFA88FDD8FAED" authority="Chen. Kottelat &amp; Miller, 1999" authorityName="Chen. Kottelat &amp; Miller" authorityYear="1999" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="species" species="albimaculatus">
<emphasis id="CEA3EA8BFFF9FFD9E89AFA88FE81FAEE" italics="true" pageId="7" pageNumber="94">R. albimaculatus</emphasis>
Chen. Kottelat &amp; Miller, 1999
</taxonomicName>
, than any other species of
<taxonomicName id="3BD74D1AFFF9FFD9EAA3FAC8FEE0FACE" authorityName="Gill" authorityYear="1859" box="[224,360,1379,1402]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="genus">
<emphasis id="CEA3EA8BFFF9FFD9EAA3FAC8FEE0FACE" box="[224,360,1379,1402]" italics="true" pageId="7" pageNumber="94">Rhinogobius</emphasis>
</taxonomicName>
. They share the similar development of predorsal scales. However,
<taxonomicName id="3BD74D1AFFF9FFD9EB9CFA28FD1FFA2E" box="[479,663,1411,1434]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="species" species="vermiculatus">
<emphasis id="CEA3EA8BFFF9FFD9EB9CFA28FD1FFA2E" box="[479,663,1411,1434]" italics="true" pageId="7" pageNumber="94">R. vermiculatus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="3BD74D1AFFF9FFD9EB0AFA08FD8BFA0E" authorityName="Chen. Kottelat &amp; Miller" authorityYear="1999" box="[329,515,1443,1466]" class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="94" phylum="Chordata" rank="species" species="albimaculatus">
<emphasis id="CEA3EA8BFFF9FFD9EB0AFA08FD8BFA0E" box="[329,515,1443,1466]" italics="true" pageId="7" pageNumber="94">R. albimaculatus</emphasis>
</taxonomicName>
by more pectoral fin rays (modally 16 vs. 15), more scales on longitudinal rows (modally 32-33 vs. 30), few large, vermiculated lines on opercle (vs. many tiny round spots on cheek and opercle); branchiostegal membrane with rounded dark greyish brown spots in male (vs. with white spots on a dark brown background).
</paragraph>
</subSubSection>
</treatment>
</document>
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