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<mods:title id="ECCC2E3985907258FBFFAE973204C523">Revision of the Palaearctic Trachusa interrupta species complex (Apoidea: Anthidiini) with description of four new species</mods:title>
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<paragraph id="ED6A7506924FFFE97DDAFC98FD53FCB9" blockId="38.[151,672,826,888]" box="[151,672,826,853]" pageId="38" pageNumber="39">
<heading id="B622C26A924FFFE97DDAFC98FD53FCB9" bold="true" box="[151,672,826,853]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="DFA1A914924FFFE97DDAFC98FD53FCB9" bold="true" box="[151,672,826,853]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFC98FDFBFCB8" ID-CoL="8SSL3" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[151,520,826,853]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="DFA1A914924FFFE97DDAFC98FEBCFCB8" bold="true" box="[151,335,826,853]" italics="true" pageId="38" pageNumber="39">Trachusa varia</emphasis>
(
<bibRefCitation id="894408F7924FFFE97C13FC98FDF3FCB8" author="Olivier, G. A." box="[350,512,826,852]" pageId="38" pageNumber="39" pagination="57 - 84" refId="ref27496" refString="Olivier, G. A. (1789) Abeille. Apis. Lin. Geof. Fab. [Linnaeus, Geoffroy, Fabricius]. Encyclopedie Methodique ou par Ordre de Matieres, Par une Societe de Gens de Lettres, de Savans et d'Artistes, Livr. Histoire Naturelle, Insectes, 4, 57 - 84." type="journal article" year="1789">Olivier, 1789</bibRefCitation>
)
</taxonomicName>
stat. resurr.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5CF268D924FFFE97DDAFCFCFEA2FC94" box="[151,337,862,888]" pageId="38" pageNumber="39" type="description">
<paragraph id="ED6A7506924FFFE97DDAFCFCFEA2FC94" blockId="38.[151,672,826,888]" box="[151,337,862,888]" pageId="38" pageNumber="39">
(
<figureCitation id="75EE6983924FFFE97DD2FCFCFF05FC94" box="[159,246,862,888]" captionStart="FIGURE 33" captionStartId="33.[151,250,1947,1972]" captionTargetBox="[155,1432,1641,1922]" captionTargetId="figure@33.[155,1432,1641,1922]" captionTargetPageId="33" captionText="FIGURE 33. Comparison of the head colour patterns in the female of the Trachusa interrupta species complex. The light pattern over black ground colour is red or reddish in T. varia and yellow in all other species. The dark colour on figure shows the light colour on the bee." figureDoi="http://doi.org/10.5281/zenodo.3614499" httpUri="https://zenodo.org/record/3614499/files/figure.png" pageId="38" pageNumber="39">Figs 33</figureCitation>
,
<figureCitation id="75EE6983924FFFE97C4FFCFCFEBAFC94" box="[258,329,862,888]" captionStart-0="FIGURE 39" captionStart-1="FIGURE 40" captionStart-2="FIGURE 41" captionStartId-0="39.[151,250,1303,1328]" captionStartId-1="40.[151,250,1290,1315]" captionStartId-2="41.[151,250,1980,2005]" captionTargetBox-0="[151,1436,198,1226]" captionTargetBox-1="[151,1436,181,1265]" captionTargetBox-2="[164,1423,849,1955]" captionTargetId-0="figure@39.[151,1436,198,1226]" captionTargetId-1="figure@40.[151,1436,181,1265]" captionTargetId-2="figure@41.[164,1423,849,1955]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionText-0="FIGURE 39. Trachusa varia. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views." captionText-1="FIGURE 40. Trachusa varia. Female from Melilla, the Spanish enclave in Morocco, in the collection Mavromoustakis." captionText-2="FIGURE 41. Trachusa varia. Male. Paralectotype of Apis fulvipes from the Fabricius collection. A. Habitus in lateral view; B. metasoma in dorsal view; C. head in frontal view; D. apical terga with genitalia." figureDoi-0="http://doi.org/10.5281/zenodo.3614511" figureDoi-1="http://doi.org/10.5281/zenodo.3614513" figureDoi-2="http://doi.org/10.5281/zenodo.3614515" httpUri-0="https://zenodo.org/record/3614511/files/figure.png" httpUri-1="https://zenodo.org/record/3614513/files/figure.png" httpUri-2="https://zenodo.org/record/3614515/files/figure.png" pageId="38" pageNumber="39">3941</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A5CF268D924FFFE97DDAFC3CFAACFAEB" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="ED6A7506924FFFE97DDAFC3CFC13FC5A" blockId="38.[151,1436,925,1287]" box="[151,992,925,950]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="CDC55228924FFFE97DDAFC3CFC13FC5A" box="[151,992,925,950]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFC3CFDB1FC5A" ID-CoL="FN66" authority="Fabricius (1787: 303)" authorityName="Fabricius" authorityPageNumber="303" authorityYear="1787" box="[151,578,925,950]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97DDAFC3CFEE2FC59" box="[151,273,925,949]" italics="true" pageId="38" pageNumber="39">Apis rufipes</emphasis>
<treatmentCitation id="6C745317924FFFE97C5AFC3FFE00FC59" author="Fabricius, J. C." box="[279,499,925,950]" page="303" pageId="38" pageNumber="39" year="1787">
<bibRefCitation id="894408F7924FFFE97C5AFC3FFE00FC59" author="Fabricius, J. C." box="[279,499,925,950]" pageId="38" pageNumber="39" refId="ref26487" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom 1. Christ. Gottl. Proft, Hafniae (Kobenhavn), 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius (1787: 303)</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="95C23596924FFFE97F4DFC3FFDC8FC5A" box="[512,571,925,950]" name="Spain" pageId="38" pageNumber="39">Spain</collectingCountry>
)
</taxonomicName>
.—Synonymy by
<bibRefCitation id="894408F7924FFFE97FBAFC3FFC67FC59" author="Warncke, K." box="[759,916,925,950]" pageId="38" pageNumber="39" pagination="119 - 209" refId="ref28049" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980</bibRefCitation>
,
<bibRefCitation id="894408F7924FFFE97ED3FC3FFC26FC5A" author="Warncke, K." box="[926,981,925,950]" pageId="38" pageNumber="39" pagination="5 - 128" refId="ref28078" refString="Warncke, K. (1986) Die Wildbienen Mitteleuropas. Ihre gultigen Namen und ihre Verbreitung (Insecta: Hymenoptera). Entomofauna, 3 (Supplementum), 5 - 128." type="journal article" year="1986">1986</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFC1EFE8EFC1F" blockId="38.[151,1436,925,1287]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="CDC55228924FFFE97DDAFC1EFE8EFC1F" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFC1EFE4FFC38" authority="Olivier, 1789: 74" authorityName="Olivier" authorityPageNumber="74" authorityYear="1789" box="[151,444,955,980]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="DFA1A914924FFFE97DDAFC1EFEF1FC38" box="[151,258,956,980]" italics="true" pageId="38" pageNumber="39">Apis varia</emphasis>
<treatmentCitation id="6C745317924FFFE97C46FC19FE4FFC38" author="Olivier, G. A." box="[267,444,955,980]" page="74" pageId="38" pageNumber="39" year="1789">
<bibRefCitation id="894408F7924FFFE97C46FC19FE4FFC38" author="Olivier, G. A." box="[267,444,955,980]" pageId="38" pageNumber="39" pagination="57 - 84" refId="ref27496" refString="Olivier, G. A. (1789) Abeille. Apis. Lin. Geof. Fab. [Linnaeus, Geoffroy, Fabricius]. Encyclopedie Methodique ou par Ordre de Matieres, Par une Societe de Gens de Lettres, de Savans et d'Artistes, Livr. Histoire Naturelle, Insectes, 4, 57 - 84." type="journal article" year="1789">Olivier, 1789: 74</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.—Replacement name for
<taxonomicName id="2AD50E85924FFFE97F82FC1EFCB9FC38" ID-CoL="FN66" authorityName="Fabricius" authorityYear="1787" box="[719,842,956,980]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97F82FC1EFCB9FC38" box="[719,842,956,980]" italics="true" pageId="38" pageNumber="39">Apis rufipes</emphasis>
</taxonomicName>
F., not
<taxonomicName id="2AD50E85924FFFE97ED3FC1EFB30FC38" ID-CoL="FN66" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[926,1219,955,980]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97ED3FC1EFBE9FC38" box="[926,1050,956,980]" italics="true" pageId="38" pageNumber="39">Apis rufipes</emphasis>
<bibRefCitation id="894408F7924FFFE9796EFC19FB30FC38" author="Fabricius, J. C." box="[1059,1219,955,980]" pageId="38" pageNumber="39" refId="ref26605" refString="Fabricius, J. C. (1781) Species insectorum, exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus. Tom 1. Hamburg &amp; Kilonii, Bohnii, 552 pp. https: // doi. org / 10.5962 / bhl. title. 36509" type="book" year="1781">Fabricius, 1781</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="2AD50E85924FFFE979AFFC1EFE8AFC1F" ID-CoL="3YSBW" authority="(Fabricius, 1781)" baseAuthorityName="Fabricius" baseAuthorityYear="1781" class="Insecta" family="Megachilidae" genus="Megachile" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE979AFFC1EFA6FFC38" box="[1250,1436,956,980]" italics="true" pageId="38" pageNumber="39">Megachile rufipes</emphasis>
(
<bibRefCitation id="894408F7924FFFE97D83FC78FE82FC1F" author="Fabricius, J. C." box="[206,369,986,1011]" pageId="38" pageNumber="39" refId="ref26605" refString="Fabricius, J. C. (1781) Species insectorum, exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin, adjectis observationibus, descriptionibus. Tom 1. Hamburg &amp; Kilonii, Bohnii, 552 pp. https: // doi. org / 10.5962 / bhl. title. 36509" type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFC58FBCBFBFD" blockId="38.[151,1436,925,1287]" box="[151,1080,1017,1042]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="CDC55228924FFFE97DDAFC58FBCBFBFD" box="[151,1080,1017,1042]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFC58FE29FBFE" authority="Fabricius, 1789" authorityName="Fabricius" authorityYear="1789" box="[151,474,1017,1042]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="erythropus">
<emphasis id="DFA1A914924FFFE97DDAFC58FECBFBFD" box="[151,312,1017,1041]" italics="true" pageId="38" pageNumber="39">Apis erythropus</emphasis>
Fabricius, 1789
</taxonomicName>
.—Replacement name for
<taxonomicName id="2AD50E85924FFFE97FAAFC58FCBAFBFD" ID-CoL="3YSBW" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[743,841,1017,1041]" class="Insecta" family="Megachilidae" genus="Megachile" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97FAAFC58FCBAFBFD" box="[743,841,1017,1041]" italics="true" pageId="38" pageNumber="39">A. rufipes</emphasis>
</taxonomicName>
F. (
<treatmentCitation id="6C745317924FFFE97E22FC5BFBDEFBFD" author="Gmelin, J. F. &amp; von Linne, C." box="[879,1069,1017,1042]" page="278" pageId="38" pageNumber="39" year="1790">
<bibRefCitation id="894408F7924FFFE97E22FC5BFBDEFBFD" author="Gmelin, J. F. &amp; von Linne, C." box="[879,1069,1017,1042]" pageId="38" pageNumber="39" pagination="2770 - 2796" refId="ref26856" refString="Gmelin, J. F. &amp; von Linne, C. (1790) Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tom 1. Pars V. Georg Emanual Beer, Lipsiae (Leipzig), 27 pp. [pp. 2770 - 2796]" type="book chapter" year="1790">Gmelin 1790: 278</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFBBAFBE1FBA2" blockId="38.[151,1436,925,1287]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFBBAFE30FBDC" authority="Fabricius, 1793" authorityName="Fabricius" authorityYear="1793" box="[151,451,1047,1072]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97DDAFBBAFEE8FBDC" box="[151,283,1048,1072]" italics="true" pageId="38" pageNumber="39">Apis fulvipes</emphasis>
<bibRefCitation id="894408F7924FFFE97C6EFBB5FE30FBDC" author="Fabricius, J. C." box="[291,451,1047,1072]" pageId="38" pageNumber="39" refId="ref26545" refString="Fabricius, J. C. (1793) Entomologia Systematica Emendata et aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus. Christ. Gottl. Proft, Hafniae (Kobenhavn), 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius, 1793</bibRefCitation>
</taxonomicName>
.—Ent. system., Vol. 2: 333 (
<collectingCountry id="95C23596924FFFE97FBCFBB5FCD8FBDC" box="[753,811,1047,1072]" name="Spain" pageId="38" pageNumber="39">Spain</collectingCountry>
).—Without referring to
<bibRefCitation id="894408F7924FFFE97961FBB5FB31FBDC" author="Olivier, G. A." box="[1068,1218,1047,1072]" pageId="38" pageNumber="39" pagination="57 - 84" refId="ref27496" refString="Olivier, G. A. (1789) Abeille. Apis. Lin. Geof. Fab. [Linnaeus, Geoffroy, Fabricius]. Encyclopedie Methodique ou par Ordre de Matieres, Par une Societe de Gens de Lettres, de Savans et d'Artistes, Livr. Histoire Naturelle, Insectes, 4, 57 - 84." type="journal article" year="1789">Olivier (1789)</bibRefCitation>
,
<bibRefCitation id="894408F7924FFFE97983FBB5FA84FBDC" author="Fabricius, J. C." box="[1230,1399,1047,1072]" pageId="38" pageNumber="39" refId="ref26545" refString="Fabricius, J. C. (1793) Entomologia Systematica Emendata et aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus. Christ. Gottl. Proft, Hafniae (Kobenhavn), 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius (1793)</bibRefCitation>
repeated his description of
<taxonomicName id="2AD50E85924FFFE97C8BFB95FDDBFBA3" ID-CoL="3YSBW" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[454,552,1079,1103]" class="Insecta" family="Megachilidae" genus="Megachile" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97C8BFB95FDDBFBA3" box="[454,552,1079,1103]" italics="true" pageId="38" pageNumber="39">A. rufipes</emphasis>
</taxonomicName>
from 1787, but now under the name
<taxonomicName id="2AD50E85924FFFE97EECFB95FBFFFBA3" box="[929,1036,1079,1103]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97EECFB95FBFFFBA3" box="[929,1036,1079,1103]" italics="true" pageId="38" pageNumber="39">A. fulvipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFBF4FC0DFB81" blockId="38.[151,1436,925,1287]" box="[151,1022,1109,1134]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFBF4FE70FB82" authority="Fabricius." authorityName="Fabricius." box="[151,387,1109,1134]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97DDAFBF4FEE9FB81" box="[151,282,1109,1133]" italics="true" pageId="38" pageNumber="39">Apis fulvipes</emphasis>
Fabricius.
</taxonomicName>
—Listed by
<bibRefCitation id="894408F7924FFFE97F4EFBF7FD58FB81" author="Fabricius, J. C." box="[515,683,1109,1134]" pageId="38" pageNumber="39" refId="ref26661" refString="Fabricius, J. C. (1804) Systema Piezatorum Secundum Ordines, Genera, Species. Apud Carolum Reichard, Braunschweig (Brunsvigae), 439 + 30 pp." type="book" year="1804">Fabricius (1804)</bibRefCitation>
without referring to
<taxonomicName id="2AD50E85924FFFE97E33FBF4FC0BFB81" ID-CoL="FN66" authorityName="Fabricius" authorityYear="1781" box="[894,1016,1109,1133]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97E33FBF4FC0BFB81" box="[894,1016,1109,1133]" italics="true" pageId="38" pageNumber="39">Apis rufipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFBD6FEAEFB47" blockId="38.[151,1436,925,1287]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFBD6FDE9FB60" ID-CoL="5VFP2" authority="(Fabricius, 1787)" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[151,538,1139,1164]" class="Insecta" family="Apidae" genus="Anthophora" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97DDAFBD6FE95FB60" box="[151,358,1140,1164]" italics="true" pageId="38" pageNumber="39">Anthophora fulvipes</emphasis>
(
<bibRefCitation id="894408F7924FFFE97C39FBD1FDE0FB60" author="Fabricius, J. C." box="[372,531,1139,1164]" pageId="38" pageNumber="39" refId="ref26487" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom 1. Christ. Gottl. Proft, Hafniae (Kobenhavn), 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius, 1787</bibRefCitation>
)
</taxonomicName>
.—Listed by
<bibRefCitation id="894408F7924FFFE97FEEFBD1FCDDFB60" author="Illiger, K." box="[675,814,1139,1164]" pageId="38" pageNumber="39" pagination="28 - 175" refId="ref27085" refString="Illiger, K. (1806) William Kirby's Familien der bienenartigen Insekten mit Zusatzen nach Weisungen und Bemerkungen. Magazin fur Insektenkunde, 5, 28 - 175." type="journal article" year="1806">Illiger (1806)</bibRefCitation>
, who did not examine material of this taxon but refers to
<bibRefCitation id="894408F7924FFFE97837FBD6FEAAFB47" author="Fabricius, J. C." pageId="38" pageNumber="39" refId="ref26661" refString="Fabricius, J. C. (1804) Systema Piezatorum Secundum Ordines, Genera, Species. Apud Carolum Reichard, Braunschweig (Brunsvigae), 439 + 30 pp." type="book" year="1804">Fabricius (1804)</bibRefCitation>
.
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFB13FD43FB25" blockId="38.[151,1436,925,1287]" box="[151,688,1201,1226]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFB13FDF3FB26" ID-CoL="8SQVD" authority="(Fabricius, 1787)" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[151,512,1201,1226]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97DDAFB13FEBEFB25" box="[151,333,1201,1225]" italics="true" pageId="38" pageNumber="39">Anthidium rufipes</emphasis>
(
<bibRefCitation id="894408F7924FFFE97C16FB13FE0AFB26" author="Fabricius, J. C." box="[347,505,1201,1226]" pageId="38" pageNumber="39" refId="ref26487" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom 1. Christ. Gottl. Proft, Hafniae (Kobenhavn), 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius, 1787</bibRefCitation>
)
</taxonomicName>
.—
<bibRefCitation id="894408F7924FFFE97F53FB13FD5FFB25" author="Friese, H." box="[542,684,1201,1226]" pageId="38" pageNumber="39" refId="ref26828" refString="Friese, H. (1911) Das Tierreich. Hymenoptera. Apidae I. Megachilinae. R. Friedlander und Sohn, Berlin, 440 pp." type="book" year="1911">Friese (1911)</bibRefCitation>
.
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFB72FB8EFB04" blockId="38.[151,1436,925,1287]" box="[151,1149,1231,1256]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFB72FE0DFB04" ID-CoL="8SNWL" authority="Ferton, 1921" authorityName="Ferton" authorityYear="1921" box="[151,510,1231,1256]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="foliivolutor">
<emphasis id="DFA1A914924FFFE97DDAFB72FE8AFB04" box="[151,377,1232,1256]" italics="true" pageId="38" pageNumber="39">Anthidium foliivolutor</emphasis>
<bibRefCitation id="894408F7924FFFE97C32FB72FE0DFB04" author="Ferton, C." box="[383,510,1231,1256]" pageId="38" pageNumber="39" pagination="329 - 375" refId="ref26735" refString="Ferton, C. (1921) Notes detachees sur l'instinct des hymenopteres melliferes et ravisseurs avec la description de deux especes nouvelles. Annales de la Societe Entomologique de France, 89 (1920 / 1921), 329 - 375." type="journal article" year="1921">Ferton, 1921</bibRefCitation>
</taxonomicName>
.—Ghazaouet (formerly Nemours),
<collectingCountry id="95C23596924FFFE97E26FB6DFC44FB04" box="[875,951,1231,1256]" name="Algeria" pageId="38" pageNumber="39">Algeria</collectingCountry>
(male and female).
</paragraph>
<paragraph id="ED6A7506924FFFE97DDAFB4DFAACFAEB" blockId="38.[151,1436,925,1287]" box="[151,1375,1262,1287]" pageId="38" pageNumber="39">
<taxonomicName id="2AD50E85924FFFE97DDAFB4DFDD7FAEB" ID-CoL="8SR4R" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[151,548,1262,1287]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="interruptum">
<emphasis id="DFA1A914924FFFE97DDAFB4DFE73FAEA" box="[151,384,1263,1287]" italics="true" pageId="38" pageNumber="39">Anthidium interruptum</emphasis>
<bibRefCitation id="894408F7924FFFE97CCBFB4CFDD7FAEB" author="Fabricius, J. C." box="[390,548,1262,1287]" pageId="38" pageNumber="39" refId="ref26487" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom 1. Christ. Gottl. Proft, Hafniae (Kobenhavn), 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius, 1787</bibRefCitation>
</taxonomicName>
(partim).—Synonymy by
<bibRefCitation id="894408F7924FFFE97E7DFB4CFC20FAEB" author="Warncke, K." box="[816,979,1262,1287]" pageId="38" pageNumber="39" pagination="119 - 209" refId="ref28049" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
after examination of the
<typeStatus id="326ECBA4924FFFE9799EFB4DFB0CFAEB" box="[1235,1279,1263,1287]" pageId="38" pageNumber="39">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="A5CF268D924FFFE97DDAFA8DFC48F93A" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="ED6A7506924FFFE97DDAFA8DFB7AF9AA" blockId="38.[151,1437,1327,2038]" pageId="38" pageNumber="39">
<emphasis id="DFA1A914924FFFE97DDAFA8DFE17FAA6" bold="true" box="[151,484,1327,1354]" pageId="38" pageNumber="39">Material examined. Female</emphasis>
.
<emphasis id="DFA1A914924FFFE97CB9FA8DFD60FAA6" bold="true" box="[500,659,1327,1354]" pageId="38" pageNumber="39">
<typeStatus id="326ECBA4924FFFE97CB9FA8DFD60FAA6" box="[500,659,1327,1354]" pageId="38" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
: Female (
<collectionCode id="8BC4EDC3924FFFE97E45FA92FC97FAA6" box="[776,868,1328,1354]" pageId="38" pageNumber="39">NHMD</collectionCode>
). Original label
<emphasis id="DFA1A914924FFFE97967FA92FB79FAA6" box="[1066,1162,1328,1354]" italics="true" pageId="38" pageNumber="39">fulvipes</emphasis>
. Additional label in the handwriting of K. Warncke “
<typeStatus id="326ECBA4924FFFE97CA5FAF6FD94FA82" box="[488,615,1364,1390]" pageId="38" pageNumber="39" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="2AD50E85924FFFE97F3FFAF6FC3CFA82" authority="(Warncke 1977)" baseAuthorityName="Warncke" baseAuthorityYear="1977" box="[626,975,1364,1390]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97F3FFAF6FCF5FA82" box="[626,774,1364,1390]" italics="true" pageId="38" pageNumber="39">Apis fulvipes</emphasis>
(Warncke 1977)
</taxonomicName>
”. Head absent as already mentioned by
<bibRefCitation id="894408F7924FFFE97DDAFADAFEBFFA7E" author="Warncke, K." box="[151,332,1400,1426]" pageId="38" pageNumber="39" pagination="119 - 209" refId="ref28049" refString="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpalaarktis und im turkestanischen Becken. Entomofauna, 1, 119 - 209." type="journal article" year="1980">Warncke (1980)</bibRefCitation>
.—
<emphasis id="DFA1A914924FFFE97C22FAD5FD81FA7E" bold="true" box="[367,626,1399,1426]" pageId="38" pageNumber="39">
<typeStatus id="326ECBA4924FFFE97C22FAD5FD81FA7E" box="[367,626,1399,1426]" pageId="38" pageNumber="39" type="paralectotype">PARALECTOTYPEs</typeStatus>
</emphasis>
:
<specimenCount id="FBD3BE8F924FFFE97FCFFADAFD56FA7E" box="[642,677,1400,1426]" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
(
<collectionCode id="8BC4EDC3924FFFE97FF8FADAFCE2FA7E" box="[693,785,1400,1426]" pageId="38" pageNumber="39">NHMD</collectionCode>
). ZUMC 00241548, red label type and an additional red label in K. Warnckes handwriting
<typeStatus id="326ECBA4924FFFE97F56FA3EFD36FA5A" box="[539,709,1436,1462]" pageId="38" pageNumber="39" type="paralectotype">Paralectotypus</typeStatus>
<taxonomicName id="2AD50E85924FFFE97F86FA3EFBEBFA5B" authority="(Warncke 1977)" baseAuthorityName="Warncke" baseAuthorityYear="1977" box="[715,1048,1436,1463]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97F86FA3EFCAAFA5A" box="[715,857,1436,1462]" italics="true" pageId="38" pageNumber="39">Apis fulvipes</emphasis>
(Warncke1977)
</taxonomicName>
.—
<specimenCount id="FBD3BE8F924FFFE97972FA3EFBA8FA5B" box="[1087,1115,1436,1463]" pageId="38" pageNumber="39" type="male">1♂</specimenCount>
(
<collectionCode id="8BC4EDC3924FFFE97924FA3EFB31FA5A" box="[1129,1218,1436,1462]" pageId="38" pageNumber="39">NHMD</collectionCode>
) ZUMC 00241547.
<taxonomicName id="2AD50E85924FFFE97DECFA62FEE5FA36" baseAuthorityName="Warncke" baseAuthorityYear="1977" box="[161,278,1472,1498]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97DECFA62FEE5FA36" box="[161,278,1472,1498]" italics="true" pageId="38" pageNumber="39">A. fulvipes</emphasis>
</taxonomicName>
D. Vahl. Red label type and an additional red label in K. Warnckes handwriting
<typeStatus id="326ECBA4924FFFE979F7FA62FA97FA36" box="[1210,1380,1472,1498]" pageId="38" pageNumber="39" type="paralectotype">Paralectotypus</typeStatus>
<taxonomicName id="2AD50E85924FFFE97827FA62FE42FA12" authority="(Warncke 1977)" baseAuthorityName="Warncke" baseAuthorityYear="1977" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97827FA62FF1DFA12" italics="true" pageId="38" pageNumber="39">Apis fulvipes</emphasis>
(Warncke 1977)
</taxonomicName>
. All antennal segments available but badly damaged. Pinned through the head. This is the male of
<taxonomicName id="2AD50E85924FFFE97DBDF9AAFE85F9CE" authorityName="Fabricius" authorityYear="1781" box="[240,374,1544,1570]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97DBDF9AAFE85F9CE" box="[240,374,1544,1570]" italics="true" pageId="38" pageNumber="39">Apis rufipes</emphasis>
</taxonomicName>
in Fabricius collection.—
<emphasis id="DFA1A914924FFFE97FD0F9AAFC9CF9CE" bold="true" box="[669,879,1543,1570]" pageId="38" pageNumber="39">Further material:</emphasis>
<collectingCountry id="95C23596924FFFE97E38F9AAFC3BF9CE" box="[885,968,1544,1570]" name="Spain" pageId="38" pageNumber="39">SPAIN</collectingCountry>
:
<specimenCount id="FBD3BE8F924FFFE97E9CF9AAFC00F9CE" box="[977,1011,1544,1570]" pageId="38" pageNumber="39" type="female">2♀</specimenCount>
<collectingRegion id="2F11BBE4924FFFE97EB4F9AAFBB9F9CE" box="[1017,1098,1544,1570]" country="Spain" name="Melilla" pageId="38" pageNumber="39">Melilla</collectingRegion>
[Spanish enclave in Morocco]
<date id="996B53C6924FFFE97DDAF98EFEFEF9AA" box="[151,269,1580,1606]" pageId="38" pageNumber="39" value="1949-06">June 1949</date>
, B. Sincan leg., both
<taxonomicName id="2AD50E85924FFFE97CB6F98EFC1EF9AA" authority="Mavromoustakis" authorityName="Mavromoustakis" box="[507,1005,1580,1606]" class="Insecta" family="Megachilidae" genus="Paraanthidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="foliivolutor">
<emphasis id="DFA1A914924FFFE97CB6F98EFCD9F9AA" box="[507,810,1580,1606]" italics="true" pageId="38" pageNumber="39">Paraanthidium foliivolutor</emphasis>
Mavromoustakis
</taxonomicName>
det. (cMAV).
</paragraph>
<paragraph id="ED6A7506924FFFE97D8AF9EDFC48F93A" blockId="38.[151,1437,1327,2038]" pageId="38" pageNumber="39">
<emphasis id="DFA1A914924FFFE97D8AF9EDFE7DF986" bold="true" box="[199,398,1615,1642]" pageId="38" pageNumber="39">Taxonomic note.</emphasis>
<bibRefCitation id="894408F7924FFFE97CDBF9F2FDA1F986" author="Fabricius, J. C." box="[406,594,1616,1642]" pageId="38" pageNumber="39" refId="ref26487" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom 1. Christ. Gottl. Proft, Hafniae (Kobenhavn), 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius (1787)</bibRefCitation>
described
<taxonomicName id="2AD50E85924FFFE97F82F9F2FCABF986" authorityName="Fabricius" authorityYear="1781" box="[719,856,1616,1642]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="DFA1A914924FFFE97F82F9F2FCABF986" box="[719,856,1616,1642]" italics="true" pageId="38" pageNumber="39">Apis rufipes</emphasis>
</taxonomicName>
based on material collected in
<collectingCountry id="95C23596924FFFE979F2F9F2FAF1F986" box="[1215,1282,1616,1642]" name="Spain" pageId="38" pageNumber="39">Spain</collectingCountry>
. As the name was preoccupied, the following replacement names were suggested:
<taxonomicName id="2AD50E85924FFFE97EC1F9D6FC0CF961" authorityName="Olivier" authorityYear="1789" box="[908,1023,1652,1678]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="DFA1A914924FFFE97EC1F9D6FC0CF961" box="[908,1023,1652,1678]" italics="true" pageId="38" pageNumber="39">Apis varia</emphasis>
</taxonomicName>
by Oliver (1789),
<taxonomicName id="2AD50E85924FFFE97986F9D6FA89F962" authorityName="Fabricius" authorityYear="1789" box="[1227,1402,1652,1678]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="erythropus">
<emphasis id="DFA1A914924FFFE97986F9D6FA89F962" box="[1227,1402,1652,1678]" italics="true" pageId="38" pageNumber="39">Apis erythropus</emphasis>
</taxonomicName>
by
<bibRefCitation id="894408F7924FFFE97DDAF93AFECFF95E" author="Gmelin, J. F. &amp; von Linne, C." box="[151,316,1688,1714]" pageId="38" pageNumber="39" pagination="2770 - 2796" refId="ref26856" refString="Gmelin, J. F. &amp; von Linne, C. (1790) Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tom 1. Pars V. Georg Emanual Beer, Lipsiae (Leipzig), 27 pp. [pp. 2770 - 2796]" type="book chapter" year="1790">Gmelin (1790)</bibRefCitation>
and
<taxonomicName id="2AD50E85924FFFE97C3CF93AFDF3F95E" baseAuthorityName="Warncke" baseAuthorityYear="1977" box="[369,512,1688,1714]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="DFA1A914924FFFE97C3CF93AFDF3F95E" box="[369,512,1688,1714]" italics="true" pageId="38" pageNumber="39">Apis fulvipes</emphasis>
</taxonomicName>
by
<bibRefCitation id="894408F7924FFFE97F64F93AFD15F95E" author="Fabricius, J. C." box="[553,742,1688,1714]" pageId="38" pageNumber="39" refId="ref26545" refString="Fabricius, J. C. (1793) Entomologia Systematica Emendata et aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus. Christ. Gottl. Proft, Hafniae (Kobenhavn), 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius (1793)</bibRefCitation>
.
<taxonomicName id="2AD50E85924FFFE97FA2F93AFC91F95D" authorityName="Olivier" authorityYear="1789" box="[751,866,1688,1714]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="DFA1A914924FFFE97FA2F93AFC91F95D" box="[751,866,1688,1714]" italics="true" pageId="38" pageNumber="39">Apis varia</emphasis>
</taxonomicName>
has priority. This is not the only case known where Johann Christian Fabricius described a homonym (
<bibRefCitation id="894408F7924FFFE97F9DF91EFC5DF93A" author="van der Vecht, J." box="[720,942,1723,1750]" pageId="38" pageNumber="39" pagination="1 - 85" refId="ref28027" refString="van der Vecht, J. (1961) Hymenoptera Sphecoidea Fabriciana. Zoologische Verhandelingen, 48, 1 - 85." type="journal article" year="1961">van der Vecht 1961</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A5CF268D924FFFE97D8AF97DFB13F81A" pageId="38" pageNumber="39" type="description">
<paragraph id="ED6A7506924FFFE97D8AF97DFF09F866" blockId="38.[151,1437,1327,2038]" pageId="38" pageNumber="39">
<emphasis id="DFA1A914924FFFE97D8AF97DFEA2F916" bold="true" box="[199,337,1759,1786]" pageId="38" pageNumber="39">Description</emphasis>
.
<emphasis id="DFA1A914924FFFE97C2FF942FE45F916" box="[354,438,1760,1786]" italics="true" pageId="38" pageNumber="39">Female</emphasis>
: Colour of all maculations red (
<figureCitation id="75EE6983924FFFE97E67F942FC73F916" box="[810,896,1760,1786]" captionStart="FIGURE 39" captionStartId="39.[151,250,1303,1328]" captionTargetBox="[151,1436,198,1226]" captionTargetId="figure@39.[151,1436,198,1226]" captionTargetPageId="39" captionText="FIGURE 39. Trachusa varia. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.3614511" httpUri="https://zenodo.org/record/3614511/files/figure.png" pageId="38" pageNumber="39">Fig. 39</figureCitation>
); band along preoccipital ridge extending from lower gena to middle of vertex, and joining with maculation of paraocular area (
<figureCitation id="75EE6983924FFFE97963F8A6FB75F8F2" box="[1070,1158,1796,1822]" captionStart="FIGURE 33" captionStartId="33.[151,250,1947,1972]" captionTargetBox="[155,1432,1641,1922]" captionTargetId="figure@33.[155,1432,1641,1922]" captionTargetPageId="33" captionText="FIGURE 33. Comparison of the head colour patterns in the female of the Trachusa interrupta species complex. The light pattern over black ground colour is red or reddish in T. varia and yellow in all other species. The dark colour on figure shows the light colour on the bee." figureDoi="http://doi.org/10.5281/zenodo.3614499" httpUri="https://zenodo.org/record/3614499/files/figure.png" pageId="38" pageNumber="39">Figs 33</figureCitation>
,
<figureCitation id="75EE6983924FFFE979D9F8A6FB35F8F2" box="[1172,1222,1796,1822]" captionStart="FIGURE 39" captionStartId="39.[151,250,1303,1328]" captionTargetBox="[151,1436,198,1226]" captionTargetId="figure@39.[151,1436,198,1226]" captionTargetPageId="39" captionText="FIGURE 39. Trachusa varia. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.3614511" httpUri="https://zenodo.org/record/3614511/files/figure.png" pageId="38" pageNumber="39">39B</figureCitation>
); transversal stripe behind ocellus; scutum with anterolateral stripe; scutellum and axillae together with four red spots; all tergal bands uninterrupted; mid-femur with red stripe almost reaching distal end; inner side of hind tibia red (
<figureCitation id="75EE6983924FFFE97988F8EEFAD8F88A" box="[1221,1323,1868,1894]" captionStart="FIGURE 39" captionStartId="39.[151,250,1303,1328]" captionTargetBox="[151,1436,198,1226]" captionTargetId="figure@39.[151,1436,198,1226]" captionTargetPageId="39" captionText="FIGURE 39. Trachusa varia. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.3614511" httpUri="https://zenodo.org/record/3614511/files/figure.png" pageId="38" pageNumber="39">Fig. 39C</figureCitation>
); pronotal lobe red.
</paragraph>
<paragraph id="ED6A7506924FFFE97D8AF836FB13F81A" blockId="38.[151,1437,1327,2038]" pageId="38" pageNumber="39">
<emphasis id="DFA1A914924FFFE97D8AF836FEF2F842" box="[199,257,1940,1966]" italics="true" pageId="38" pageNumber="39">Male</emphasis>
(
<figureCitation id="75EE6983924FFFE97C5FF836FE9AF842" box="[274,361,1940,1966]" captionStart="FIGURE 41" captionStartId="41.[151,250,1980,2005]" captionTargetBox="[164,1423,849,1955]" captionTargetId="figure@41.[164,1423,849,1955]" captionTargetPageId="41" captionText="FIGURE 41. Trachusa varia. Male. Paralectotype of Apis fulvipes from the Fabricius collection. A. Habitus in lateral view; B. metasoma in dorsal view; C. head in frontal view; D. apical terga with genitalia." figureDoi="http://doi.org/10.5281/zenodo.3614515" httpUri="https://zenodo.org/record/3614515/files/figure.png" pageId="38" pageNumber="39">Fig. 41</figureCitation>
): Colour of all maculations yellow to rufous; bands on T1 and T2 interrupted, others uninterrupted (
<figureCitation id="75EE6983924FFFE97DA3F81AFEA5F83E" box="[238,342,1976,2002]" captionStart="FIGURE 41" captionStartId="41.[151,250,1980,2005]" captionTargetBox="[164,1423,849,1955]" captionTargetId="figure@41.[164,1423,849,1955]" captionTargetPageId="41" captionText="FIGURE 41. Trachusa varia. Male. Paralectotype of Apis fulvipes from the Fabricius collection. A. Habitus in lateral view; B. metasoma in dorsal view; C. head in frontal view; D. apical terga with genitalia." figureDoi="http://doi.org/10.5281/zenodo.3614515" httpUri="https://zenodo.org/record/3614515/files/figure.png" pageId="38" pageNumber="39">Fig. 41B</figureCitation>
); scutum with narrow, hardly discernible rufous lateral stripe; scutellum and axillae black; rufous stripe of mid-femur reaching middle of femur; inner side of hind tibia black; pronotal lobe rufous.
</paragraph>
</subSubSection>
<caption id="B9AA258E924EFFE87DDAFAB5FD45FAB8" ID-DOI="http://doi.org/10.5281/zenodo.3614511" ID-Zenodo-Dep="3614511" httpUri="https://zenodo.org/record/3614511/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1303,1328]" targetBox="[151,1436,198,1226]" targetPageId="39">
<paragraph id="ED6A7506924EFFE87DDAFAB5FD45FAB8" blockId="39.[151,1436,1303,1364]" pageId="39" pageNumber="40">
<emphasis id="DFA1A914924EFFE87DDAFAB5FED2FADC" bold="true" box="[151,289,1303,1328]" pageId="39" pageNumber="40">FIGURE 39.</emphasis>
<taxonomicName id="2AD50E85924EFFE87C65FAB5FE30FAC3" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[296,451,1303,1327]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="DFA1A914924EFFE87C65FAB5FE30FAC3" box="[296,451,1303,1327]" italics="true" pageId="39" pageNumber="40">Trachusa varia</emphasis>
</taxonomicName>
. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views.
</paragraph>
</caption>
<subSubSection id="A5CF268D924EFFE87D8AFA23FEFFF8A0" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="ED6A7506924EFFE87D8AFA23FBC7F9BC" blockId="39.[151,1437,1409,2012]" pageId="39" pageNumber="40">
<emphasis id="DFA1A914924EFFE87D8AFA23FE39FA70" bold="true" box="[199,458,1409,1436]" pageId="39" pageNumber="40">Differential Diagnosis</emphasis>
. Habitus and colouration pattern of the female similar to
<taxonomicName id="2AD50E85924EFFE87909FA21FB23FA70" box="[1092,1232,1411,1436]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="interrupta">
<emphasis id="DFA1A914924EFFE87909FA21FB23FA70" box="[1092,1232,1411,1436]" italics="true" pageId="39" pageNumber="40">T. interrupta</emphasis>
</taxonomicName>
but all pale maculations red or reddish instead of bright yellow. Additionally, the female is distinguished from all other species of the complex by (1) a red band extending from the genae to the middle of the vertex along the preoccipital ridges; (2) red maculation of the paraocular areas merged with the bands along the preoccipital ridges, and (3) a transverse red band behind the ocelli (
<figureCitation id="75EE6983924EFFE87C2CF9B0FE44F9C0" box="[353,439,1554,1580]" captionStart="FIGURE 33" captionStartId="33.[151,250,1947,1972]" captionTargetBox="[155,1432,1641,1922]" captionTargetId="figure@33.[155,1432,1641,1922]" captionTargetPageId="33" captionText="FIGURE 33. Comparison of the head colour patterns in the female of the Trachusa interrupta species complex. The light pattern over black ground colour is red or reddish in T. varia and yellow in all other species. The dark colour on figure shows the light colour on the bee." figureDoi="http://doi.org/10.5281/zenodo.3614499" httpUri="https://zenodo.org/record/3614499/files/figure.png" pageId="39" pageNumber="40">Figs 33</figureCitation>
,
<figureCitation id="75EE6983924EFFE87C8EF9B0FE12F9C0" box="[451,481,1554,1580]" captionStart="FIGURE 39" captionStartId="39.[151,250,1303,1328]" captionTargetBox="[151,1436,198,1226]" captionTargetId="figure@39.[151,1436,198,1226]" captionTargetPageId="39" captionText="FIGURE 39. Trachusa varia. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.3614511" httpUri="https://zenodo.org/record/3614511/files/figure.png" pageId="39" pageNumber="40">39</figureCitation>
,
<figureCitation id="75EE6983924EFFE87CA0F9B0FDFFF9C0" box="[493,524,1554,1580]" captionStart="FIGURE 40" captionStartId="40.[151,250,1290,1315]" captionTargetBox="[151,1436,181,1265]" captionTargetId="figure@40.[151,1436,181,1265]" captionTargetPageId="40" captionText="FIGURE 40. Trachusa varia. Female from Melilla, the Spanish enclave in Morocco, in the collection Mavromoustakis." figureDoi="http://doi.org/10.5281/zenodo.3614513" httpUri="https://zenodo.org/record/3614513/files/figure.png" pageId="39" pageNumber="40">40</figureCitation>
,
<figureCitation id="75EE6983924EFFE87F5AF9B0FDC5F9C0" box="[535,566,1554,1580]" captionStart="FIGURE 41" captionStartId="41.[151,250,1980,2005]" captionTargetBox="[164,1423,849,1955]" captionTargetId="figure@41.[164,1423,849,1955]" captionTargetPageId="41" captionText="FIGURE 41. Trachusa varia. Male. Paralectotype of Apis fulvipes from the Fabricius collection. A. Habitus in lateral view; B. metasoma in dorsal view; C. head in frontal view; D. apical terga with genitalia." figureDoi="http://doi.org/10.5281/zenodo.3614515" httpUri="https://zenodo.org/record/3614515/files/figure.png" pageId="39" pageNumber="40">41</figureCitation>
). The female is further distinguished from all other species of the complex by the uninterrupted band on T1, which has some rufous inclusion medially (
<figureCitation id="75EE6983924EFFE87E9EF994FBD4F9BC" box="[979,1063,1590,1616]" captionStart="FIGURE 39" captionStartId="39.[151,250,1303,1328]" captionTargetBox="[151,1436,198,1226]" captionTargetId="figure@39.[151,1436,198,1226]" captionTargetPageId="39" captionText="FIGURE 39. Trachusa varia. Female paralectotype from the Fabricius collection. AB. Head in frontal view and at an angle from above; CD. habitus in dorsal and lateral views." figureDoi="http://doi.org/10.5281/zenodo.3614511" httpUri="https://zenodo.org/record/3614511/files/figure.png" pageId="39" pageNumber="40">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="ED6A7506924EFFE87D8AF9F8FB46F90C" blockId="39.[151,1437,1409,2012]" pageId="39" pageNumber="40">
The maculations of the male are yellow to rufous, hereby paler than in the female and closer to the other males of the complex. The hind femora are black with the distal one-thirds being reddish-yellow. A similar pattern only occurs in
<taxonomicName id="2AD50E85924EFFE87C4EF901FE4AF950" authorityName="Kasparek" authorityYear="2020" box="[259,441,1698,1724]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="maghrebensis">
<emphasis id="DFA1A914924EFFE87C4EF901FE4AF950" box="[259,441,1698,1724]" italics="true" pageId="39" pageNumber="40">T. maghrebensis</emphasis>
</taxonomicName>
from
<collectingCountry id="95C23596924EFFE87CB6F900FDA1F950" box="[507,594,1698,1724]" name="Tunisia" pageId="39" pageNumber="40">Tunisia</collectingCountry>
. Tergal bands are broad, bands on T1T2 interrupted and band on T3 with a deep notch. A similar pattern has been found in one out of nine specimens of
<taxonomicName id="2AD50E85924EFFE87EB5F965FB5DF90C" authorityName="Kasparek" authorityYear="2020" box="[1016,1198,1734,1760]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="maghrebensis">
<emphasis id="DFA1A914924EFFE87EB5F965FB5DF90C" box="[1016,1198,1734,1760]" italics="true" pageId="39" pageNumber="40">T. maghrebensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="ED6A7506924EFFE87D8AF948FEFFF8A0" blockId="39.[151,1437,1409,2012]" pageId="39" pageNumber="40">While the female is well characterised and can easily be identified by both colouration and colour pattern, the single more than 230 years old male is in bad condition and distinguishing morphological characters need to be examined.</paragraph>
</subSubSection>
<subSubSection id="A5CF268D924EFFE77D8AF8F4FA67FA3B" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="discussion">
<paragraph id="ED6A7506924EFFE87D8AF8F4FC47F830" blockId="39.[151,1437,1409,2012]" pageId="39" pageNumber="40">
<emphasis id="DFA1A914924EFFE87D8AF8F4FEC7F89C" bold="true" box="[199,308,1878,1904]" pageId="39" pageNumber="40">Remarks</emphasis>
. The description by
<bibRefCitation id="894408F7924EFFE87F69F8F4FD32F89C" author="Ferton, C." box="[548,705,1878,1904]" pageId="39" pageNumber="40" pagination="329 - 375" refId="ref26735" refString="Ferton, C. (1921) Notes detachees sur l'instinct des hymenopteres melliferes et ravisseurs avec la description de deux especes nouvelles. Annales de la Societe Entomologique de France, 89 (1920 / 1921), 329 - 375." type="journal article" year="1921">Ferton (1921)</bibRefCitation>
of
<taxonomicName id="2AD50E85924EFFE87FA6F8F4FC14F89C" authorityName="Ferton" authorityYear="1921" box="[747,999,1878,1904]" class="Insecta" family="Megachilidae" genus="Anthidium" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="foliivolutor">
<emphasis id="DFA1A914924EFFE87FA6F8F4FC14F89C" box="[747,999,1878,1904]" italics="true" pageId="39" pageNumber="40">Anthidium foliivolutor</emphasis>
</taxonomicName>
largely matches the description given here. He particularly described the colour of the male maculae as yellow, and those of the female as red, thus confirming the colour dimorphism. The male in Feltons (1921) material has the band on T1T3 interrupted, whereas they are interrupted in the male in Fabricius collection only on T1T2.
</paragraph>
<caption id="B9AA258E9241FFE77DDAFAA8FAABFACF" ID-DOI="http://doi.org/10.5281/zenodo.3614513" ID-Zenodo-Dep="3614513" box="[151,1368,1290,1315]" httpUri="https://zenodo.org/record/3614513/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1290,1315]" targetBox="[151,1436,181,1265]" targetPageId="40">
<paragraph id="ED6A75069241FFE77DDAFAA8FAABFACF" blockId="40.[151,1368,1290,1315]" box="[151,1368,1290,1315]" pageId="40" pageNumber="41">
<emphasis id="DFA1A9149241FFE77DDAFAA8FED3FACF" bold="true" box="[151,288,1290,1315]" pageId="40" pageNumber="41">FIGURE 40.</emphasis>
<taxonomicName id="2AD50E859241FFE77C6BFAA9FE33FACF" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[294,448,1291,1315]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="DFA1A9149241FFE77C6BFAA9FE33FACF" box="[294,448,1291,1315]" italics="true" pageId="40" pageNumber="41">Trachusa varia</emphasis>
</taxonomicName>
. Female from Melilla, the Spanish enclave in Morocco, in the collection Mavromoustakis.
</paragraph>
</caption>
<paragraph id="ED6A75069241FFE77D8AFAF3FA67FA3B" blockId="40.[151,1437,1361,1712]" pageId="40" pageNumber="41">
Given the great age of Fabricius
<typeStatus id="326ECBA49241FFE77F00FAF0FD8EFA80" box="[589,637,1362,1388]" pageId="40" pageNumber="41">type</typeStatus>
material (over 230 years old), it was considered whether colouration might have changed, possibly as a consequence of treatment e.g. with arsenic. While some change cannot be ruled out, the colour tone (red, ochreous, yellow) is well correlated with the colour design and the same unique combination of colour and colour design was found in material collected in the same region approximately 150 years later.
</paragraph>
</subSubSection>
<subSubSection id="A5CF268D9241FFE77D8AFA43FC29F943" pageId="40" pageNumber="41" type="distribution">
<paragraph id="ED6A75069241FFE77D8AFA43FC29F943" blockId="40.[151,1437,1361,1712]" pageId="40" pageNumber="41">
<emphasis id="DFA1A9149241FFE77D8AFA43FE92FA10" bold="true" box="[199,353,1505,1532]" pageId="40" pageNumber="41">Distribution.</emphasis>
<collectingCountry id="95C235969241FFE77C2AFA43FE54FA17" box="[359,423,1505,1531]" name="Spain" pageId="40" pageNumber="41">Spain</collectingCountry>
and North Africa.
<bibRefCitation id="894408F79241FFE77F37FA43FCDEFA10" author="Fabricius, J. C." box="[634,813,1505,1532]" pageId="40" pageNumber="41" refId="ref26487" refString="Fabricius, J. C. (1787) Mantissa insectorum sistens eorum species nuper detectas adiectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom 1. Christ. Gottl. Proft, Hafniae (Kobenhavn), 348 pp. https: // doi. org / 10.5962 / bhl. title. 11657" type="book" year="1787">Fabricius (1787</bibRefCitation>
,
<bibRefCitation id="894408F79241FFE77E75FA40FC80FA10" author="Fabricius, J. C." box="[824,883,1506,1532]" pageId="40" pageNumber="41" refId="ref26545" refString="Fabricius, J. C. (1793) Entomologia Systematica Emendata et aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus. Christ. Gottl. Proft, Hafniae (Kobenhavn), 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">1793</bibRefCitation>
,
<bibRefCitation id="894408F79241FFE77E33FA40FC44FA10" author="Fabricius, J. C." box="[894,951,1506,1532]" pageId="40" pageNumber="41" refId="ref26661" refString="Fabricius, J. C. (1804) Systema Piezatorum Secundum Ordines, Genera, Species. Apud Carolum Reichard, Braunschweig (Brunsvigae), 439 + 30 pp." type="book" year="1804">1804</bibRefCitation>
) gives
<collectingCountry id="95C235969241FFE77945FA43FBBBFA17" box="[1032,1096,1505,1531]" name="Spain" pageId="40" pageNumber="41">Spain</collectingCountry>
as the location where the
<typeStatus id="326ECBA49241FFE77821FA40FA6FFA10" box="[1388,1436,1506,1532]" pageId="40" pageNumber="41">type</typeStatus>
material was collected by Prof. Martin Vahl (
<date id="996B53C69241FFE77FDCF9A4FCE7F9CC" box="[657,788,1542,1568]" pageId="40" pageNumber="41" value="1749" valueMax="1804">17491804</date>
), who was a botanist who collected in
<collectingCountry id="95C235969241FFE7798AF9A7FAF4F9F3" box="[1223,1287,1541,1567]" name="Spain" pageId="40" pageNumber="41">Spain</collectingCountry>
and northern Africa (e.g.
<collectingCountry id="95C235969241FFE77C53F98BFE91F9A8" box="[286,354,1577,1604]" name="Tunisia" pageId="40" pageNumber="41">Tunis</collectingCountry>
). Information on the exact collecting site or on the travel itinerary was not available but it cannot be ruled out that the material actually came from northern Africa and not from
<collectingCountry id="95C235969241FFE77962F9EFFB81F98B" box="[1071,1138,1613,1639]" name="Spain" pageId="40" pageNumber="41">Spain</collectingCountry>
. Further material is available from
<collectingCountry id="95C235969241FFE77C43F9D3FE91F967" box="[270,354,1649,1675]" name="Spain" pageId="40" pageNumber="41">Melilla</collectingCountry>
, the Spanish enclave in
<collectingCountry id="95C235969241FFE77F3AF9D0FD2CF960" box="[631,735,1650,1676]" name="Morocco" pageId="40" pageNumber="41">Morocco</collectingCountry>
, and the material of
<bibRefCitation id="894408F79241FFE77E81F9D0FB9BF960" author="Ferton, C." box="[972,1128,1650,1676]" pageId="40" pageNumber="41" pagination="329 - 375" refId="ref26735" refString="Ferton, C. (1921) Notes detachees sur l'instinct des hymenopteres melliferes et ravisseurs avec la description de deux especes nouvelles. Annales de la Societe Entomologique de France, 89 (1920 / 1921), 329 - 375." type="journal article" year="1921">Ferton (1921)</bibRefCitation>
from Ghazaouet (formerly Nemours) in West
<collectingCountry id="95C235969241FFE77C2AF937FE48F943" box="[359,443,1685,1711]" name="Algeria" pageId="40" pageNumber="41">Algeria</collectingCountry>
can apparently also be attributed to this species.
</paragraph>
</subSubSection>
</treatment>
</document>