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<document id="8E4C553F736E54A03B43FDB934D70EC0" ID-DOI="10.5281/zenodo.293434" ID-GBIF-Dataset="90a8372c-5dc2-4af5-abc6-d6974ab05a11" ID-ISSN="1175-5326" ID-Zenodo-Dep="293434" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460479489793" checkinUser="plazi" docAuthor="Theron, Elizabeth Omukunda Pieter D &amp; Ueckermann, E A" docDate="2012" docId="03FCB14BFFB1FFF2E2D2FD93FA0DFBAB" docLanguage="en" docName="zt03304p024.pdf" docOrigin="Zootaxa 3304" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Spinibdella pongolensis Theron &amp; Ueckermann, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FFC5C933FFBAFFFCE245FFFFFF98FFB0" masterDocTitle="Spinibdella Thor (Acari: Bdellidae) from southern Africa: descriptions of five new species and the redescription of S. thori (Meyer &amp; Ryke)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="12" updateTime="1698307322454" updateUser="plazi">
<mods:mods id="23DDCD9B8DAD89A28894611297709535" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D58F62236554EDDD335523FC0AC94D96">
<mods:title id="56CB6807D195328EBD9C3F0BFF0C372F">Spinibdella Thor (Acari: Bdellidae) from southern Africa: descriptions of five new species and the redescription of S. thori (Meyer &amp; Ryke)</mods:title>
</mods:titleInfo>
<mods:name id="B3011AA33B9FCC1934217D1D39EEA0A5" type="personal">
<mods:role id="03E17FA20E679C03B9B51C7D2D216EB9">
<mods:roleTerm id="2E90CE05D49D13D88C1C65EDD862114D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2DF4CD1B098D89690FFCADC59C75C35A">Theron, Elizabeth Omukunda Pieter D</mods:namePart>
</mods:name>
<mods:name id="596A70BBCCB55EF7AEE8C79003960073" type="personal">
<mods:role id="E88E905CFB5F9438F7B5D0EACB25F0EC">
<mods:roleTerm id="ADDFF6D1AE8B4F4E7E975C85699EEBE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A36030C0BEE3EF226FBF7287E32714A9">Ueckermann, E A</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D987028BBA042ADA47B8FBAFE308750">text</mods:typeOfResource>
<mods:relatedItem id="6A7B9387B485932B5007C93CD1077E62" type="host">
<mods:titleInfo id="3B22DF6D5261494406917E9D305659FB">
<mods:title id="F86A4195FCBE179CF62390A6BDD876D5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6BCE04D736A7BF67A67AE73335DD51FB">
<mods:date id="B71CB575EFB0B5E079F22CD721518C2C">2012</mods:date>
<mods:detail id="CEE095AF41AE2BC4770F0B374169FE2F" type="volume">
<mods:number id="0C9B6A99F6BC3A429759DF54E1253599">3304</mods:number>
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<mods:extent id="E71B99913E8742EBF580D134B84B5FCB" unit="page">
<mods:start id="90A286E42D03E293BA0E7697328FDF1E">1</mods:start>
<mods:end id="D398E677F7F03147930FACAFBD048493">24</mods:end>
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<mods:classification id="0800377A6BE16BADD87D21123310AE72">journal article</mods:classification>
<mods:identifier id="64F5F48CBCA98C7A3C5A1648A37D5AB1" type="DOI">10.5281/zenodo.293434</mods:identifier>
<mods:identifier id="42D119CCA85ACEE00ABC4CBC6D60920F" type="GBIF-Dataset">90a8372c-5dc2-4af5-abc6-d6974ab05a11</mods:identifier>
<mods:identifier id="1BDB82B9DF3F8D128B3E9CEB23F26F36" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3A8B49EDC67D70420A95BC1DBF946B60" type="Zenodo-Dep">293434</mods:identifier>
</mods:mods>
<treatment id="03FCB14BFFB1FFF2E2D2FD93FA0DFBAB" ID-DOI="http://doi.org/10.5281/zenodo.6168965" ID-GBIF-Taxon="119516922" ID-Zenodo-Dep="6168965" LSID="urn:lsid:plazi:treatment:03FCB14BFFB1FFF2E2D2FD93FA0DFBAB" httpUri="http://treatment.plazi.org/id/03FCB14BFFB1FFF2E2D2FD93FA0DFBAB" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C34F53D6FFB1FFF7E2D2FD93FE8CFD16" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BEA005DFFB1FFF7E2D2FD93FC7DFD35" blockId="11.[151,997,619,678]" box="[151,997,619,646]" pageId="11" pageNumber="12">
<heading id="D0A2B731FFB1FFF7E2D2FD93FC7DFD35" bold="true" box="[151,997,619,646]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B921DC4FFFB1FFF7E2D2FD93FC7DFD35" bold="true" box="[151,997,619,646]" pageId="11" pageNumber="12">
<taxonomicName id="4C557BDEFFB1FFF7E2D2FD93FE2EFD36" box="[151,438,620,646]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pongolensis" status="sp. nov.">
<emphasis id="B921DC4FFFB1FFF7E2D2FD93FE2EFD36" bold="true" box="[151,438,620,646]" italics="true" pageId="11" pageNumber="12">Spinibdella pongolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2126134FFB1FFF7E3F8FD94FD83FD35" box="[445,539,619,645]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
Omukunda, Theron &amp; Ueckermann
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA005DFFB1FFF7E2D2FD71FE8CFD16" blockId="11.[151,997,619,678]" box="[151,276,654,678]" pageId="11" pageNumber="12">
<figureCitation id="136E1CD8FFB1FFF7E2D2FD71FE8CFD16" box="[151,276,654,678]" captionStart-0="FIGURES 28 – 34" captionStart-1="FIGURES 35 – 36" captionStartId-0="12.[151,264,1648,1671]" captionStartId-1="13.[151,264,1968,1991]" captionTargetBox-0="[216,1382,372,1617]" captionTargetBox-1="[152,1390,568,1967]" captionTargetId-0="figure@12.[151,1436,340,1648]" captionTargetId-1="figure@13.[151,1436,521,1989]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 28 – 34. Spinibdella pongolensis sp. n.: 28. Female chelicera; 29. Female palp; 30. Female hypostome; 31. Female prodorsum; 32. Female dorsal view; 33. Female ventral view; 34. Female ovipositor." captionText-1="FIGURES 35 – 36. Spinibdella pongolensis sp. n.: 35. Female leg I; 36. Female leg II." httpUri-0="https://zenodo.org/record/293441/files/figure.png" httpUri-1="https://zenodo.org/record/293442/files/figure.png" pageId="11" pageNumber="12">Figs 28–36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F53D6FFB1FFF2E2D2FD29FBB2FD4B" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
<paragraph id="8BEA005DFFB1FFF7E2D2FD29FECDFC7C" blockId="11.[151,1437,726,2025]" pageId="11" pageNumber="12">
<emphasis id="B921DC4FFFB1FFF7E2D2FD29FE14FD5F" bold="true" box="[151,396,726,751]" pageId="11" pageNumber="12">Description. Female</emphasis>
(n = 2). Dimensions: Dimensions: length including gnathosoma 800–944 (800); width 290–382 (290); length of gnathosoma 220–244 (224); length of chelicerae 210–216 (210);
<emphasis id="B921DC4FFFB1FFF7E6D3FD04FB55FCA2" box="[1174,1229,763,786]" italics="true" pageId="11" pageNumber="12">DHS</emphasis>
21; length of palp segments:
<date id="FFEB269DFFB1FFF7E355FCE0FEF0FC86" box="[272,360,798,822]" pageId="11" pageNumber="12" value="2010-01-13">I 10–13</date>
(13); II + III 191–210 (191);
<date id="FFEB269DFFB1FFF7E081FCE0FCA8FC86" box="[708,816,798,822]" pageId="11" pageNumber="12" value="1923-04-25">IV 23–25</date>
(23);
<date id="FFEB269DFFB1FFF7E13FFCE0FC2BFC86" box="[890,947,798,822]" pageId="11" pageNumber="12" value="1934-05">V 34</date>
–38 (34);
<emphasis id="B921DC4FFFB1FFF7E663FCE1FBD3FC86" box="[1062,1099,798,822]" italics="true" pageId="11" pageNumber="12">des</emphasis>
258–267 (258);
<emphasis id="B921DC4FFFB1FFF7E748FCDFFAA9FC87" box="[1293,1329,800,823]" italics="true" pageId="11" pageNumber="12">ves</emphasis>
176–191 (191); length of legs: I 226–362 (226); II 281–342 (281); III 394–416 (394); IV 426–483 (426); length of setae:
<date id="FFEB269DFFB1FFF7E7CDFCBBFF2EFCCE" pageId="11" pageNumber="12" value="1955-06">
<emphasis id="B921DC4FFFB1FFF7E7CDFCBBFA04FCEB" box="[1416,1436,836,859]" italics="true" pageId="11" pageNumber="12">vi</emphasis>
55
</date>
;
<emphasis id="B921DC4FFFB1FFF7E280FC97FF46FCCF" box="[197,222,872,895]" italics="true" pageId="11" pageNumber="12">ve</emphasis>
95–108 (108);
<emphasis id="B921DC4FFFB1FFF7E3D3FC98FE2DFCCE" box="[406,437,871,894]" italics="true" pageId="11" pageNumber="12">sci</emphasis>
48;
<emphasis id="B921DC4FFFB1FFF7E3ABFC97FD8AFCCF" box="[494,530,872,895]" italics="true" pageId="11" pageNumber="12">sce</emphasis>
77–93 (93);
<emphasis id="B921DC4FFFB1FFF7E0EBFC97FD5BFC33" box="[686,707,872,899]" italics="true" pageId="11" pageNumber="12">c1</emphasis>
75–79 (75);
<emphasis id="B921DC4FFFB1FFF7E11AFC97FCEBFC33" box="[863,883,872,899]" italics="true" pageId="11" pageNumber="12">c2</emphasis>
137–147 (137);
<emphasis id="B921DC4FFFB1FFF7E67CFC99FBDFFCCE" box="[1081,1095,870,894]" italics="true" pageId="11" pageNumber="12">d</emphasis>
68–73 (73);
<emphasis id="B921DC4FFFB1FFF7E6A1FC97FB68FCCF" box="[1252,1264,872,895]" italics="true" pageId="11" pageNumber="12">e</emphasis>
63–66 (66);
<emphasis id="B921DC4FFFB1FFF7E7C9FC99FA04FC33" box="[1420,1436,870,899]" italics="true" pageId="11" pageNumber="12">f1</emphasis>
114–121 (114);
<emphasis id="B921DC4FFFB1FFF7E30EFC72FEC2FC19" box="[331,346,909,937]" italics="true" pageId="11" pageNumber="12">f2</emphasis>
85–108 (85);
<emphasis id="B921DC4FFFB1FFF7E3B9FC72FD8AFC19" box="[508,530,909,937]" italics="true" pageId="11" pageNumber="12">h1</emphasis>
146–158 (185);
<emphasis id="B921DC4FFFB1FFF7E08AFC72FD7DFC19" box="[719,741,909,937]" italics="true" pageId="11" pageNumber="12">h2</emphasis>
125–139 (139); distance
<emphasis id="B921DC4FFFB1FFF7E643FC71FBA4FC15" box="[1030,1084,910,933]" italics="true" pageId="11" pageNumber="12">
vi–
<date id="FFEB269DFFB1FFF7E66DFC71FBA4FC15" box="[1064,1084,910,933]" pageId="11" pageNumber="12" value="1946-06">vi</date>
</emphasis>
46–48 (46); first interspace (
<emphasis id="B921DC4FFFB1FFF7E7CDFC71FA04FC19" box="[1416,1436,910,937]" italics="true" pageId="11" pageNumber="12">c1</emphasis>
to
<emphasis id="B921DC4FFFB1FFF7E2F1FC4BFF5AFC7C" box="[180,194,948,972]" italics="true" pageId="11" pageNumber="12">d</emphasis>
) 69–75 (75).
</paragraph>
<paragraph id="8BEA005DFFB1FFF7E28CFC28FB7CFB14" blockId="11.[151,1437,726,2025]" pageId="11" pageNumber="12">
<emphasis id="B921DC4FFFB1FFF7E28CFC28FEFAFC40" bold="true" box="[201,354,983,1008]" pageId="11" pageNumber="12">Gnathosoma</emphasis>
(
<figureCitation id="136E1CD8FFB1FFF7E335FC27FE6CFC40" box="[368,500,984,1008]" captionStart="FIGURES 28 – 34" captionStartId="12.[151,264,1648,1671]" captionTargetBox="[216,1382,372,1617]" captionTargetId="figure@12.[151,1436,340,1648]" captionTargetPageId="12" captionText="FIGURES 28 – 34. Spinibdella pongolensis sp. n.: 28. Female chelicera; 29. Female palp; 30. Female hypostome; 31. Female prodorsum; 32. Female dorsal view; 33. Female ventral view; 34. Female ovipositor." httpUri="https://zenodo.org/record/293441/files/figure.png" pageId="11" pageNumber="12">Figs 28–30</figureCitation>
). Chelicerae have weak continuous to sparsely broken longitudinal striations, a pair of strong setae inserted on each as shown in figure 28. Fixed digit rather short and straight, movable digit slightly longer. Palp extends beyond hypostome. Palp tibiotarsi have one long blunt solenidion on ventral side as shown in figure 29. Palp chaetotaxy from trochanter to tibiotarsus as follows: 0-9-1-4-4 sts, 1 ω,
<emphasis id="B921DC4FFFB1FFF7E63CFBBBFB06FBEC" box="[1145,1182,1092,1116]" italics="true" pageId="11" pageNumber="12">des</emphasis>
,
<emphasis id="B921DC4FFFB1FFF7E6E8FBBAFB48FBEC" box="[1197,1232,1093,1116]" italics="true" pageId="11" pageNumber="12">ves</emphasis>
. Hypostome with continuous to rough longitudinal striations, base with fine transverse striations. Hypostome has two pairs of strong setae and adoral setae as shown in figure 30. Dorsal hypostomal setae exceptionally long (21 µm).
</paragraph>
<paragraph id="8BEA005DFFB1FFF7E28CFB4FFB46FA13" blockId="11.[151,1437,726,2025]" pageId="11" pageNumber="12">
<emphasis id="B921DC4FFFB1FFF7E28CFB4FFEB2FB78" bold="true" box="[201,298,1200,1224]" pageId="11" pageNumber="12">Dorsum</emphasis>
(
<figureCitation id="136E1CD8FFB1FFF7E37DFB4FFE23FB78" box="[312,443,1200,1224]" captionStart="FIGURES 28 – 34" captionStartId="12.[151,264,1648,1671]" captionTargetBox="[216,1382,372,1617]" captionTargetId="figure@12.[151,1436,340,1648]" captionTargetPageId="12" captionText="FIGURES 28 – 34. Spinibdella pongolensis sp. n.: 28. Female chelicera; 29. Female palp; 30. Female hypostome; 31. Female prodorsum; 32. Female dorsal view; 33. Female ventral view; 34. Female ovipositor." httpUri="https://zenodo.org/record/293441/files/figure.png" pageId="11" pageNumber="12">Figs 31–32</figureCitation>
). Naso simple, detectable median eye posterior to naso. All dorsal setae simple. Setae
<emphasis id="B921DC4FFFB1FFF7E7C6FB4EFA04FB78" box="[1411,1436,1201,1224]" italics="true" pageId="11" pageNumber="12">ve</emphasis>
straight, located in anterior half of prodorsum and longer than
<emphasis id="B921DC4FFFB1FFF7E110FB2AFCE1FB5C" box="[853,889,1237,1260]" italics="true" pageId="11" pageNumber="12">sce</emphasis>
. Setae
<emphasis id="B921DC4FFFB1FFF7E189FB2AFC78FB5C" box="[972,992,1237,1260]" italics="true" pageId="11" pageNumber="12">vi</emphasis>
and
<emphasis id="B921DC4FFFB1FFF7E652FB2AFBAEFB5C" box="[1047,1078,1237,1260]" italics="true" pageId="11" pageNumber="12">sci</emphasis>
lost in this specimen. Center of prodorsum with sparsely broken to continuous transverse striations that strongly curve outward towards anterior. Posterior striations slightly curve outward posteriorly as seen in
<figureCitation id="136E1CD8FFB1FFF7E132FAE3FC70FA84" box="[887,1000,1308,1332]" captionStart="FIGURES 28 – 34" captionStartId="12.[151,264,1648,1671]" captionTargetBox="[216,1382,372,1617]" captionTargetId="figure@12.[151,1436,340,1648]" captionTargetPageId="12" captionText="FIGURES 28 – 34. Spinibdella pongolensis sp. n.: 28. Female chelicera; 29. Female palp; 30. Female hypostome; 31. Female prodorsum; 32. Female dorsal view; 33. Female ventral view; 34. Female ovipositor." httpUri="https://zenodo.org/record/293441/files/figure.png" pageId="11" pageNumber="12">Figure 31</figureCitation>
. Five detectable eyes present, anterior pair separated from posterior pair by transverse striations. Distance between them equals diameter of anterior pair. Sejugal groove conspicuous. Setae
<emphasis id="B921DC4FFFB1FFF7E060FA9AFDA1FA30" box="[549,569,1381,1408]" italics="true" pageId="11" pageNumber="12">c1</emphasis>
about half first interspace. Setae
<emphasis id="B921DC4FFFB1FFF7E1F7FA9AFC5FFA30" box="[946,967,1381,1408]" italics="true" pageId="11" pageNumber="12">c2</emphasis>
, twice length of
<emphasis id="B921DC4FFFB1FFF7E6C3FA9AFB02FA30" box="[1158,1178,1381,1408]" italics="true" pageId="11" pageNumber="12">c1</emphasis>
as shown in figure 32. Internal clunals (
<emphasis id="B921DC4FFFB1FFF7E311FA74FEF3FA17" box="[340,363,1419,1447]" italics="true" pageId="11" pageNumber="12">h1</emphasis>
) long. Posterior annals (
<emphasis id="B921DC4FFFB1FFF7E03BFA73FD07FA17" box="[638,671,1420,1447]" italics="true" pageId="11" pageNumber="12">ps1</emphasis>
) near anal cleft and two pairs of anal setae present.
</paragraph>
<paragraph id="8BEA005DFFB1FFF7E28CFA4DFBCEF9E9" blockId="11.[151,1437,726,2025]" pageId="11" pageNumber="12">
<emphasis id="B921DC4FFFB1FFF7E28CFA4DFE81FA7A" bold="true" box="[201,281,1458,1482]" pageId="11" pageNumber="12">Venter</emphasis>
(
<figureCitation id="136E1CD8FFB1FFF7E362FA4EFE32FA79" box="[295,426,1457,1481]" captionStart="FIGURES 28 – 34" captionStartId="12.[151,264,1648,1671]" captionTargetBox="[216,1382,372,1617]" captionTargetId="figure@12.[151,1436,340,1648]" captionTargetPageId="12" captionText="FIGURES 28 – 34. Spinibdella pongolensis sp. n.: 28. Female chelicera; 29. Female palp; 30. Female hypostome; 31. Female prodorsum; 32. Female dorsal view; 33. Female ventral view; 34. Female ovipositor." httpUri="https://zenodo.org/record/293441/files/figure.png" pageId="11" pageNumber="12">Figs 33–34</figureCitation>
). All ventral setae simple. Twenty-eight to 34 long blunt aggenital setae, first two unpaired between coxae IV. Some may be slightly branched distally. Ten to 11 pairs of blunt linearly arranged genital setae equally long as shown in figure 33. Three pairs of equal genital discs, first pair in anterior half and two pairs in posterior half. Genital tracheae well defined. Ovipositor with 18 setae and a characteristic lip as seen in figure 34. Ovipositor has longitudinal striations. Anal cleft has two pairs of anal setae (
<emphasis id="B921DC4FFFB1FFF7E18DF9BDFC74F9EE" box="[968,1004,1602,1630]" italics="true" pageId="11" pageNumber="12">ad2</emphasis>
and
<emphasis id="B921DC4FFFB1FFF7E667F9BCFBDEF9EE" box="[1058,1094,1603,1630]" italics="true" pageId="11" pageNumber="12">an1</emphasis>
).
</paragraph>
<paragraph id="8BEA005DFFB1FFF7E28CF996FEF0F858" blockId="11.[151,1437,726,2025]" pageId="11" pageNumber="12">
<emphasis id="B921DC4FFFB1FFF7E28CF996FE99F931" bold="true" box="[201,257,1641,1665]" pageId="11" pageNumber="12">Legs</emphasis>
(
<figureCitation id="136E1CD8FFB1FFF7E355F997FE0CF930" box="[272,404,1640,1664]" captionStart="FIGURES 35 – 36" captionStartId="13.[151,264,1968,1991]" captionTargetBox="[152,1390,568,1967]" captionTargetId="figure@13.[151,1436,521,1989]" captionTargetPageId="13" captionText="FIGURES 35 – 36. Spinibdella pongolensis sp. n.: 35. Female leg I; 36. Female leg II." httpUri="https://zenodo.org/record/293442/files/figure.png" pageId="11" pageNumber="12">Figs 35–36</figureCitation>
). Legs shorter than idiosoma. Leg II shorter than leg I while rest increase progressively. All tactile setae simple. Genu I with an attenuate solenidion, tibia I has a distal trichobothrium, solid attenuate famulus and two closely associated attenuate solenidia as shown in figure 35. Tarsi I have one attenuate, two blunt solenidia. Claws nude and shorter than pretarsi. Tibiae II have one attenuate and one reduced blunt solenidion each. Tarsi II have only one blunt solenidion in proximal half as shown in figure 36. Genua, telofemora and basifemora each have a macroseta. Tibiae III have a distal attenuate solenidion while tarsi III have a proximal trichobothrium. Tibiae IV have a trichobothrium in distal half. Tarsi IV have an attenuate solenidion anterior to a proximal trichobothrium. Leg chaetotaxy as follows: coxae
<date id="FFEB269DFFB1FFF7E0C0F89AFD57F8CC" box="[645,719,1892,1916]" pageId="11" pageNumber="12">I–IV 7</date>
(8-9)-6-9-4(3) sts; trochanters
<date id="FFEB269DFFB1FFF7E667F89AFBE7F8CC" box="[1058,1151,1892,1916]" pageId="11" pageNumber="12">I–IV 1-1</date>
-2-1 sts; basifemora
<date id="FFEB269DFFB1FFF7E722F89AFF52F810" pageId="11" pageNumber="12" value="2011-01-07" valueMax="2011-04-07" valueMin="2011-01-07">I–IV 11-7</date>
(8-9)-6-3(2) sts; telofemora
<date id="FFEB269DFFB1FFF7E047F876FDD4F810" box="[514,588,1928,1952]" pageId="11" pageNumber="12">I–IV 7</date>
(8)-8(7)-5(6)-7(6) sts; genua
<date id="FFEB269DFFB1FFF7E1CAF876FC40F810" box="[911,984,1929,1952]" pageId="11" pageNumber="12">I–IV 7</date>
sts, lσ -6(7)-5(7)-5(6) sts; tibiae
<date id="FFEB269DFFB1FFF7E700F876FA04F810" box="[1349,1436,1928,1952]" pageId="11" pageNumber="12" value="1916-01" valueMax="1916-04-31" valueMin="1916-01-16">I–IV 16</date>
(15) sts, 2φ, 1ĸ, 1tr -13(12) sts, 2φ -12 sts, 1φ -14(15) sts, 1tr; tarsi
<date id="FFEB269DFFB1FFF7E1CBF852FC7FF874" box="[910,999,1964,1988]" pageId="11" pageNumber="12" value="1929-01" valueMax="1929-04-31" valueMin="1929-01-29">I–IV 29</date>
(30) sts, 3ω -29 sts, lω, 1 ĸ -28(27) sts, 1tr -26 sts, 1tr, 1ω.
</paragraph>
<paragraph id="8BEA005DFFB6FFF0E28CFF68FCCAFEF2" blockId="12.[151,1436,151,327]" pageId="12" pageNumber="13">
<emphasis id="B921DC4FFFB6FFF0E28CFF68FECFFF00" bold="true" box="[201,343,151,176]" pageId="12" pageNumber="13">Tritonymph</emphasis>
(n = 3). Dimensions: length including gnathosoma 714–756; width 277–382; length of gnathosoma 191–210; length of chelicerae 164–182; length of palp segments:
<date id="FFEB269DFFB6FFF0E190FF42FBA9FF64" box="[981,1073,188,212]" pageId="12" pageNumber="13" value="2010-01-12">I 10–12</date>
; II + III 143–162;
<date id="FFEB269DFFB6FFF0E755FF42FAE6FF64" box="[1296,1406,188,212]" pageId="12" pageNumber="13" value="1919-04-21">IV 19–21</date>
<date id="FFEB269DFFB6FFF0E7CDFF42FF78FF47" pageId="12" pageNumber="13" value="1938-05-31">V 31–38</date>
;
<emphasis id="B921DC4FFFB6FFF0E2AEFF1FFE88FF48" box="[235,272,224,248]" italics="true" pageId="12" pageNumber="13">des</emphasis>
204–229;
<emphasis id="B921DC4FFFB6FFF0E3C2FF1EFE33FF48" box="[391,427,225,248]" italics="true" pageId="12" pageNumber="13">ves</emphasis>
134–162; length of legs: I 273–309; II 210–298; III 258–361; IV 317–407; length of setae
<emphasis id="B921DC4FFFB6FFF0E2D2FEFAFF33FEAC" box="[151,171,261,284]" italics="true" pageId="12" pageNumber="13">vi</emphasis>
103–113;
<emphasis id="B921DC4FFFB6FFF0E361FEFAFEA5FEAC" box="[292,317,261,284]" italics="true" pageId="12" pageNumber="13">ve</emphasis>
336–46;
<emphasis id="B921DC4FFFB6FFF0E3EFFEFAFE51FEAC" box="[426,457,261,284]" italics="true" pageId="12" pageNumber="13">sci</emphasis>
broken off;
<emphasis id="B921DC4FFFB6FFF0E01DFEFAFDE4FEAC" box="[600,636,261,284]" italics="true" pageId="12" pageNumber="13">sce</emphasis>
46;
<emphasis id="B921DC4FFFB6FFF0E0F4FEFAFD25FEAC" box="[689,701,261,284]" italics="true" pageId="12" pageNumber="13">c</emphasis>
<date id="FFEB269DFFB6FFF0E0F8FEECFC8FFEAC" box="[701,791,260,288]" pageId="12" pageNumber="13" value="1942-01-29">
<emphasis id="B921DC4FFFB6FFF0E0F8FEECFD5DFE90" box="[701,709,275,288]" italics="true" pageId="12" pageNumber="13">1</emphasis>
29–42
</date>
;
<emphasis id="B921DC4FFFB6FFF0E161FEFAFCA0FE90" box="[804,824,261,288]" italics="true" pageId="12" pageNumber="13">c2</emphasis>
61–80;
<emphasis id="B921DC4FFFB6FFF0E1D3FEFBFC3CFEAC" box="[918,932,260,284]" italics="true" pageId="12" pageNumber="13">d</emphasis>
32;
<emphasis id="B921DC4FFFB6FFF0E19CFEFAFC7DFEAC" box="[985,997,261,284]" italics="true" pageId="12" pageNumber="13">e</emphasis>
34;
<emphasis id="B921DC4FFFB6FFF0E65FFEFBFBA9FEAC" box="[1050,1073,260,284]" italics="true" pageId="12" pageNumber="13">f1</emphasis>
64;
<emphasis id="B921DC4FFFB6FFF0E621FEFBFBE2FEAC" box="[1124,1146,260,284]" italics="true" pageId="12" pageNumber="13">f2</emphasis>
49;
<emphasis id="B921DC4FFFB6FFF0E6EAFEFBFB53FEAC" box="[1199,1227,260,284]" italics="true" pageId="12" pageNumber="13">h1</emphasis>
71–76;
<emphasis id="B921DC4FFFB6FFF0E76FFEFBFADEFEAC" box="[1322,1350,260,284]" italics="true" pageId="12" pageNumber="13">h2</emphasis>
61–66;
<emphasis id="B921DC4FFFB6FFF0E2D2FED4FF56FEF2" box="[151,206,299,322]" italics="true" pageId="12" pageNumber="13">DHS</emphasis>
13; distance:
<emphasis id="B921DC4FFFB6FFF0E32FFED3FE38FEF3" box="[362,416,299,323]" italics="true" pageId="12" pageNumber="13">
vi–
<date id="FFEB269DFFB6FFF0E3C9FED3FE38FEF3" box="[396,416,300,323]" pageId="12" pageNumber="13" value="1936-06">vi</date>
</emphasis>
36–48; first interspace (
<emphasis id="B921DC4FFFB6FFF0E0F7FED3FD5EFEF7" box="[690,710,300,327]" italics="true" pageId="12" pageNumber="13">c1</emphasis>
to
<emphasis id="B921DC4FFFB6FFF0E0A3FED4FD6CFEF3" box="[742,756,299,323]" italics="true" pageId="12" pageNumber="13">d</emphasis>
) 57–76.
</paragraph>
<caption id="DF2A50D5FFB6FFF0E2D2F98FFC69F915" httpUri="https://zenodo.org/record/293441/files/figure.png" pageId="12" pageNumber="13" targetBox="[216,1382,372,1617]" targetPageId="12">
<paragraph id="8BEA005DFFB6FFF0E2D2F98FFC69F915" blockId="12.[151,1436,1648,1701]" pageId="12" pageNumber="13">
<emphasis id="B921DC4FFFB6FFF0E2D2F98FFECDF935" bold="true" box="[151,341,1648,1671]" pageId="12" pageNumber="13">FIGURES 28–34.</emphasis>
<taxonomicName id="4C557BDEFFB6FFF0E319F98FFDC8F936" box="[348,592,1648,1670]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pongolensis" status="sp. nov.">
<emphasis id="B921DC4FFFB6FFF0E319F98FFDC8F936" box="[348,592,1648,1670]" italics="true" pageId="12" pageNumber="13">Spinibdella pongolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2126134FFB6FFF0E012F98EFD09F936" box="[599,657,1649,1670]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
: 28. Female chelicera; 29. Female palp; 30. Female hypostome; 31. Female prodorsum; 32. Female dorsal view; 33. Female ventral view; 34. Female ovipositor.
</paragraph>
</caption>
<paragraph id="8BEA005DFFB6FFF0E28CF933FB3AF872" blockId="12.[151,1437,1740,2022]" pageId="12" pageNumber="13">
Three pairs of genital discs, anterior pair in proximal half and posterior two pairs in distal half. Six pairs of blunt genital setae linearly arranged, of approximate length. Twenty two aggenital setae surround genital region. Anterior aggenital setae unpaired and occur between coxae IV. Ventrally, three pairs of anal setae (
<emphasis id="B921DC4FFFB6FFF0E742F8EBFAC7F880" box="[1287,1375,1812,1840]" italics="true" pageId="12" pageNumber="13">ad2, an1</emphasis>
,
<emphasis id="B921DC4FFFB6FFF0E72AF8EAFA0BF880" box="[1391,1427,1813,1840]" italics="true" pageId="12" pageNumber="13">an2</emphasis>
) occur. Setae
<emphasis id="B921DC4FFFB6FFF0E36CF8C4FEA6F8E7" box="[297,318,1851,1879]" italics="true" pageId="12" pageNumber="13">h1</emphasis>
are very long and flank anal cleft. Palp basifemora have seven setae each. Leg chaetotaxy differs from that of female as follows: coxae
<date id="FFEB269DFFB6FFF0E007F89CFD15F8CA" box="[578,653,1890,1914]" pageId="12" pageNumber="13">I–IV 7</date>
(6) -5(4)-5-3 sts; trochanters
<date id="FFEB269DFFB6FFF0E197F89CFBA8F8CA" box="[978,1072,1890,1914]" pageId="12" pageNumber="13">I–IV 1-1</date>
-2-1 sts; basifemora
<date id="FFEB269DFFB6FFF0E75FF89CFAE4F8CA" box="[1306,1404,1890,1914]" pageId="12" pageNumber="13">I–IV 7-7</date>
-4- 3 sts; telofemora 1–
<date id="FFEB269DFFB6FFF0E336F879FE27F82E" box="[371,447,1926,1950]" pageId="12" pageNumber="13">IV 5-5</date>
(4)-5(4)-4 sts; genua
<date id="FFEB269DFFB6FFF0E0EBF878FD60F82E" box="[686,760,1926,1950]" pageId="12" pageNumber="13">I–IV 5</date>
sts, 1σ-5-5-4 sts; tibiae
<date id="FFEB269DFFB6FFF0E642F878FBC7F82E" box="[1031,1119,1926,1950]" pageId="12" pageNumber="13" value="1912-01" valueMax="1912-04-31" valueMin="1912-01-12">I–IV 12</date>
sts, 2φ, 1ε, 1tr -8 sts, 2φ - 8 sts, 1φ - 11 sts, 1tr; tarsi
<date id="FFEB269DFFB6FFF0E3EEF854FD9CF872" box="[427,516,1962,1986]" pageId="12" pageNumber="13" value="1925-01" valueMax="1925-04-31" valueMin="1925-01-25">I–IV 25</date>
(24) sts, 3ω - 25 sts, 1ω, 1 ĸ -23 sts, 1tr - 22(21) sts, 1ω, 1tr.
</paragraph>
<paragraph id="8BEA005DFFB6FFF1E28CF831FDCDFEAF" blockId="12.[151,1437,1740,2022]" lastBlockId="13.[151,1437,151,508]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B921DC4FFFB6FFF0E28CF831FEFDF856" bold="true" box="[201,357,1998,2022]" pageId="12" pageNumber="13">Deutonymph</emphasis>
(n = 2). Dimensions: Length including gnathosoma 622–648; width 227–238; length of gnathosoma 153–157 length of chelicerae 138–141; length of palp segments:
<date id="FFEB269DFFB7FFF1E193FF67FBBDFF1F" box="[982,1061,151,175]" pageId="13" pageNumber="14" value="1913-01-08">I 8–13</date>
; II + III 113–117;
<date id="FFEB269DFFB7FFF1E742FF67FAE0FF1F" box="[1287,1400,151,175]" pageId="13" pageNumber="14" value="1915-04-17">IV 15–17</date>
;
<date id="FFEB269DFFB7FFF1E7CDFF67FF78FF64" pageId="13" pageNumber="14" value="1927-05-29">V 27–29</date>
;
<emphasis id="B921DC4FFFB7FFF1E2ABFF43FE8CFF64" box="[238,276,188,212]" italics="true" pageId="13" pageNumber="14">des</emphasis>
176–185;
<emphasis id="B921DC4FFFB7FFF1E3CAFF42FE2BFF64" box="[399,435,189,212]" italics="true" pageId="13" pageNumber="14">ves</emphasis>
124–128; length of legs: I 224; II 197–225; III 224–237; IV 227–269; length of setae:
<emphasis id="B921DC4FFFB7FFF1E2D2FF1FFF56FF47" box="[151,206,224,247]" italics="true" pageId="13" pageNumber="14">DHS</emphasis>
13–15;
<emphasis id="B921DC4FFFB7FFF1E36FFF1EFEDBFF48" box="[298,323,225,248]" italics="true" pageId="13" pageNumber="14">ve</emphasis>
33–34;
<emphasis id="B921DC4FFFB7FFF1E3E5FF1EFE27FF48" box="[416,447,225,248]" italics="true" pageId="13" pageNumber="14">sci</emphasis>
broken off;
<emphasis id="B921DC4FFFB7FFF1E00EFF1EFDF7FF48" box="[587,623,225,248]" italics="true" pageId="13" pageNumber="14">sce</emphasis>
, 29–31;
<emphasis id="B921DC4FFFB7FFF1E096FF1EFD47FF48" box="[723,735,225,248]" italics="true" pageId="13" pageNumber="14">c</emphasis>
<date id="FFEB269DFFB7FFF1E09AFF10FCAFFF47" box="[735,823,223,252]" pageId="13" pageNumber="14" value="1925-01-29">
<emphasis id="B921DC4FFFB7FFF1E09AFF10FD7FFF4C" box="[735,743,239,252]" italics="true" pageId="13" pageNumber="14">1</emphasis>
25–29
</date>
,
<emphasis id="B921DC4FFFB7FFF1E106FF1EFCCFFF4C" box="[835,855,225,252]" italics="true" pageId="13" pageNumber="14">c2</emphasis>
57;
<emphasis id="B921DC4FFFB7FFF1E1CFFF1FFC00FF48" box="[906,920,224,248]" italics="true" pageId="13" pageNumber="14">d</emphasis>
29;
<emphasis id="B921DC4FFFB7FFF1E18FFF1EFC4EFF48" box="[970,982,225,248]" italics="true" pageId="13" pageNumber="14">e</emphasis>
33;
<emphasis id="B921DC4FFFB7FFF1E64FFF1FFB87FF48" box="[1034,1055,224,248]" italics="true" pageId="13" pageNumber="14">f1</emphasis>
57;
<emphasis id="B921DC4FFFB7FFF1E617FF1FFBFFFF48" box="[1106,1127,224,248]" italics="true" pageId="13" pageNumber="14">f2</emphasis>
40;
<emphasis id="B921DC4FFFB7FFF1E6DEFF1FFB2EFF48" box="[1179,1206,224,248]" italics="true" pageId="13" pageNumber="14">h1</emphasis>
64;
<emphasis id="B921DC4FFFB7FFF1E6AFFF1FFA9EFF48" box="[1258,1286,224,248]" italics="true" pageId="13" pageNumber="14">h2</emphasis>
52; distance:
<emphasis id="B921DC4FFFB7FFF1E2D2FEF7FF55FEAF" box="[151,205,264,287]" italics="true" pageId="13" pageNumber="14">
vi–
<date id="FFEB269DFFB7FFF1E2FCFEF7FF55FEAF" box="[185,205,264,287]" pageId="13" pageNumber="14" value="1933-06-31">vi</date>
</emphasis>
31–33; first interspace (
<emphasis id="B921DC4FFFB7FFF1E39AFEF7FE6BFE93" box="[479,499,264,291]" italics="true" pageId="13" pageNumber="14">c1</emphasis>
to
<emphasis id="B921DC4FFFB7FFF1E051FEF8FDBAFEAF" box="[532,546,263,287]" italics="true" pageId="13" pageNumber="14">d</emphasis>
) 54.
</paragraph>
<paragraph id="8BEA005DFFB7FFF1E28CFED2FDA0FE4C" blockId="13.[151,1437,151,508]" pageId="13" pageNumber="14">
This developmental stage has two pairs of genital discs; two pairs of genital setae and 12 aggenital setae, first pair between coxae IV. Three pairs of anal setae (
<emphasis id="B921DC4FFFB7FFF1E0FFFEAEFCD8FEDE" box="[698,832,337,366]" italics="true" pageId="13" pageNumber="14">ad2, an1, an2</emphasis>
) ventrally. Dorsally, anal cleft not yet visible and only posterior anal setae present. Palp basifemora have four setae each. Leg chaetotaxy differs from that of female as follows: coxae
<date id="FFEB269DFFB7FFF1E307FE62FE3CFE04" box="[322,420,412,436]" pageId="13" pageNumber="14">I–IV 5-4</date>
-4(3)-2 sts; trochanters
<date id="FFEB269DFFB7FFF1E0ECFE62FC9DFE04" box="[681,773,412,436]" pageId="13" pageNumber="14">I–IV 1-1</date>
-2-1 sts; basifemora
<date id="FFEB269DFFB7FFF1E1ABFE62FBCBFE04" box="[1006,1107,412,436]" pageId="13" pageNumber="14">I–IV 6-5</date>
-3-1 sts; telofemora
<date id="FFEB269DFFB7FFF1E777FE62FA0CFE04" box="[1330,1428,412,436]" pageId="13" pageNumber="14">I–IV 5-4</date>
- 3-4 sts; genua
<date id="FFEB269DFFB7FFF1E37DFE3EFE1AFE68" box="[312,386,448,472]" pageId="13" pageNumber="14">I–IV 5</date>
sts, 1σ - 5 -5-4 sts; tibiae
<date id="FFEB269DFFB7FFF1E0E1FE3EFD75FE68" box="[676,749,448,472]" pageId="13" pageNumber="14">I–IV 6</date>
sts, 2φ, 1ĸ-6 sts, 2φ -5 sts, 1φ -7 sts, 1tr; tarsi
<date id="FFEB269DFFB7FFF1E6B7FE3EFAD1FE68" box="[1266,1353,448,472]" pageId="13" pageNumber="14" value="1920-01" valueMax="1920-04-31" valueMin="1920-01-20">I–IV 20</date>
sts, 3ω -21 sts, 1ω-20 sts, 1tr -18 sts, 1ω, 1tr.
</paragraph>
<caption id="DF2A50D5FFB7FFF1E2D2F84FFC64F876" box="[151,1020,1968,1991]" httpUri="https://zenodo.org/record/293442/files/figure.png" pageId="13" pageNumber="14" targetBox="[152,1390,568,1967]" targetPageId="13">
<paragraph id="8BEA005DFFB7FFF1E2D2F84FFC64F876" blockId="13.[151,1020,1968,1991]" box="[151,1020,1968,1991]" pageId="13" pageNumber="14">
<emphasis id="B921DC4FFFB7FFF1E2D2F84FFECBF876" bold="true" box="[151,339,1968,1991]" pageId="13" pageNumber="14">FIGURES 35–36.</emphasis>
<taxonomicName id="4C557BDEFFB7FFF1E31FF84FFDD4F876" box="[346,588,1968,1990]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pongolensis" status="sp. nov.">
<emphasis id="B921DC4FFFB7FFF1E31FF84FFDD4F876" box="[346,588,1968,1990]" italics="true" pageId="13" pageNumber="14">Spinibdella pongolensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2126134FFB7FFF1E016F84EFD13F876" box="[595,651,1969,1990]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
: 35. Female leg I; 36. Female leg II.
</paragraph>
</caption>
<paragraph id="8BEA005DFFB4FFF2E28CFF67FA08FEAF" blockId="14.[151,1436,151,1055]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CFF67FEC7FF00" bold="true" box="[201,351,152,176]" pageId="14" pageNumber="15">Protonymph</emphasis>
(n = 2). Dimensions: length including gnathosoma 459–631; width 191–243; length of gnathosoma 134–136; length of chelicerae 113–120; length of palp segments:
<date id="FFEB269DFFB4FFF2E19CFF42FBBFFF64" box="[985,1063,188,212]" pageId="14" pageNumber="15" value="2010-01-08">I 8–10</date>
; II + III 103–113;
<date id="FFEB269DFFB4FFF2E74DFF42FAE0FF64" box="[1288,1400,188,212]" pageId="14" pageNumber="15" value="1913-04-17">IV 13–17</date>
;
<date id="FFEB269DFFB4FFF2E7CDFF42FF78FF47" pageId="14" pageNumber="15" value="1923-05-29">V 23–29</date>
;
<emphasis id="B921DC4FFFB4FFF2E2ABFF1FFE8CFF48" box="[238,276,224,248]" italics="true" pageId="14" pageNumber="15">des</emphasis>
141–159;
<emphasis id="B921DC4FFFB4FFF2E3D5FF1EFE2BFF48" box="[400,435,225,248]" italics="true" pageId="14" pageNumber="15">ves</emphasis>
86–103; length of setae:
<emphasis id="B921DC4FFFB4FFF2E099FF1FFC8AFF47" box="[732,786,224,247]" italics="true" pageId="14" pageNumber="15">DHS</emphasis>
12–13;
<date id="FFEB269DFFB4FFF2E131FF1EFC35FF47" box="[884,941,223,248]" pageId="14" pageNumber="15" value="1963-06">
<emphasis id="B921DC4FFFB4FFF2E131FF1EFC10FF48" box="[884,904,225,248]" italics="true" pageId="14" pageNumber="15">vi</emphasis>
63
</date>
–84;
<emphasis id="B921DC4FFFB4FFF2E1ACFF1EFB9AFF48" box="[1001,1026,225,248]" italics="true" pageId="14" pageNumber="15">ve</emphasis>
27–29;
<emphasis id="B921DC4FFFB4FFF2E627FF1EFB19FF48" box="[1122,1153,225,248]" italics="true" pageId="14" pageNumber="15">sci</emphasis>
75;
<emphasis id="B921DC4FFFB4FFF2E6FDFF1EFB44FF48" box="[1208,1244,225,248]" italics="true" pageId="14" pageNumber="15">sce</emphasis>
21–27;
<emphasis id="B921DC4FFFB4FFF2E778FF1EFAC9FF4C" box="[1341,1361,225,252]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
29;
<emphasis id="B921DC4FFFB4FFF2E7CDFF1EFA04FF4C" box="[1416,1436,225,252]" italics="true" pageId="14" pageNumber="15">c2</emphasis>
44–57;
<emphasis id="B921DC4FFFB4FFF2E2A9FEF8FF62FEAF" box="[236,250,263,287]" italics="true" pageId="14" pageNumber="15">d</emphasis>
27–28;
<emphasis id="B921DC4FFFB4FFF2E310FEF7FEF9FEAF" box="[341,353,264,287]" italics="true" pageId="14" pageNumber="15">e</emphasis>
29–31;
<emphasis id="B921DC4FFFB4FFF2E3F8FEF8FE4CFEAF" box="[445,468,263,287]" italics="true" pageId="14" pageNumber="15">f1</emphasis>
65–68;
<emphasis id="B921DC4FFFB4FFF2E06AFEF8FDDCFEAF" box="[559,580,263,287]" italics="true" pageId="14" pageNumber="15">f2</emphasis>
41–42;
<emphasis id="B921DC4FFFB4FFF2E0E4FEF8FD24FEAF" box="[673,700,263,287]" italics="true" pageId="14" pageNumber="15">h1</emphasis>
63;
<emphasis id="B921DC4FFFB4FFF2E0ABFEF8FC93FEAF" box="[750,779,263,287]" italics="true" pageId="14" pageNumber="15">h2</emphasis>
50; distance:
<emphasis id="B921DC4FFFB4FFF2E1E2FEF7FC45FEAF" box="[935,989,264,287]" italics="true" pageId="14" pageNumber="15">
vi–
<date id="FFEB269DFFB4FFF2E18CFEF7FC45FEAF" box="[969,989,264,287]" pageId="14" pageNumber="15" value="1923-06-27">vi</date>
</emphasis>
23–27; first interspace (
<emphasis id="B921DC4FFFB4FFF2E6AAFEF7FA9BFE93" box="[1263,1283,264,291]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
to
<emphasis id="B921DC4FFFB4FFF2E761FEF8FAAAFEAF" box="[1316,1330,263,287]" italics="true" pageId="14" pageNumber="15">d</emphasis>
) 42–67.
</paragraph>
<paragraph id="8BEA005DFFB4FFF2E28CFED2FCB1FE65" blockId="14.[151,1436,151,1055]" pageId="14" pageNumber="15">
This ontogenetic stage has one pair of genital discs, one pair of genital setae and seven aggenital setae, first unpaired seta occurring between coxae IV. Two pairs of anal setae ventrally. Tibiae have one seta each. Leg chaetotaxy differs from that of female as follows: coxae
<date id="FFEB269DFFB4FFF2E084FE89FCBBFE3D" box="[705,803,373,397]" pageId="14" pageNumber="15">I–IV 4-2</date>
-3-0 sts; trochanters
<date id="FFEB269DFFB4FFF2E640FE89FBFAFE3D" box="[1029,1122,373,397]" pageId="14" pageNumber="15">I–IV 1-1</date>
-1-0 sts; basifemora
<date id="FFEB269DFFB4FFF2E70CFE89FF3FFE01" pageId="14" pageNumber="15">I–IV 2- 2</date>
-1-0 sts; telofemora
<date id="FFEB269DFFB4FFF2E3C3FE65FE7FFE01" box="[390,487,409,433]" pageId="14" pageNumber="15">I–IV 5-4</date>
-4-0 sts; genua
<date id="FFEB269DFFB4FFF2E0D7FE65FD44FE01" box="[658,732,409,433]" pageId="14" pageNumber="15">I–IV 5</date>
sts, 1σ -5-5-0 sts; tibiae I–IV, 4 sts, 2φ, 1ĸ, 1tr -5 sts; 2φ -5 sts, 1φ -1 sts; tarsi
<date id="FFEB269DFFB4FFF2E37BFE41FE0EFE65" box="[318,406,445,469]" pageId="14" pageNumber="15" value="1916-01" valueMax="1916-04-31" valueMin="1916-01-16">I–IV 16</date>
sts, 2ω -16 sts, 1ω-16 sts, 1tr -6 sts.
</paragraph>
<paragraph id="8BEA005DFFB4FFF2E28CFE1DFBB2FD4B" blockId="14.[151,1436,151,1055]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CFE1DFE8AFE4A" bold="true" box="[201,274,482,506]" pageId="14" pageNumber="15">Larva</emphasis>
(n = 3). Dimensions: length including gnathosoma 382–407; width 172–191; length of gnathosoma 105–107; length of chelicerae 90–96; length of palp segments:
<date id="FFEB269DFFB4FFF2E1C2FDF9FC34FDAD" box="[903,940,517,541]" pageId="14" pageNumber="15">I 8</date>
; II +
<date id="FFEB269DFFB4FFF2E1BDFDF9FBA5FDAD" box="[1016,1085,517,541]" pageId="14" pageNumber="15" value="1984-03">III 84</date>
–86;
<date id="FFEB269DFFB4FFF2E63CFDF9FB75FDAD" box="[1145,1261,517,541]" pageId="14" pageNumber="15" value="2010-04-15">IV 10–15</date>
;
<date id="FFEB269DFFB4FFF2E6B8FDF9FAFEFDAD" box="[1277,1382,517,541]" pageId="14" pageNumber="15" value="1921-05-23">V 21–23</date>
;
<emphasis id="B921DC4FFFB4FFF2E733FDFAFA04FDAD" box="[1398,1436,517,541]" italics="true" pageId="14" pageNumber="15">des</emphasis>
118–122;
<emphasis id="B921DC4FFFB4FFF2E34DFDD5FEB4FDF1" box="[264,300,554,577]" italics="true" pageId="14" pageNumber="15">ves</emphasis>
75; length of legs: I 151–143; II 145–147; III 183–191; length of setae:
<date id="FFEB269DFFB4FFF2E622FDD5FB06FDF1" box="[1127,1182,553,577]" pageId="14" pageNumber="15" value="1961-06">
<emphasis id="B921DC4FFFB4FFF2E622FDD5FBE3FDF1" box="[1127,1147,554,577]" italics="true" pageId="14" pageNumber="15">vi</emphasis>
61
</date>
–76;
<emphasis id="B921DC4FFFB4FFF2E69FFDD5FB6BFDF1" box="[1242,1267,554,577]" italics="true" pageId="14" pageNumber="15">ve</emphasis>
19–21;
<emphasis id="B921DC4FFFB4FFF2E714FDD5FAE8FDF1" box="[1361,1392,554,577]" italics="true" pageId="14" pageNumber="15">sci</emphasis>
59;
<emphasis id="B921DC4FFFB4FFF2E2D2FDB0FF23FDD6" box="[151,187,591,614]" italics="true" pageId="14" pageNumber="15">sce</emphasis>
23–28;
<emphasis id="B921DC4FFFB4FFF2E352FDB0FEB3FDDA" box="[279,299,591,618]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
23;
<emphasis id="B921DC4FFFB4FFF2E318FDB0FEEAFDDA" box="[349,370,591,618]" italics="true" pageId="14" pageNumber="15">c2</emphasis>
38;
<emphasis id="B921DC4FFFB4FFF2E3E1FDB2FE2AFDD5" box="[420,434,589,613]" italics="true" pageId="14" pageNumber="15">d</emphasis>
21–25;
<emphasis id="B921DC4FFFB4FFF2E04BFDB0FD82FDD6" box="[526,538,591,614]" italics="true" pageId="14" pageNumber="15">e</emphasis>
22–24;
<emphasis id="B921DC4FFFB4FFF2E032FDB2FD16FDD5" box="[631,654,589,613]" italics="true" pageId="14" pageNumber="15">f1</emphasis>
46–48;
<emphasis id="B921DC4FFFB4FFF2E0ACFDB2FC98FDD5" box="[745,768,589,613]" italics="true" pageId="14" pageNumber="15">
f
<date id="FFEB269DFFB4FFF2E0B3FDB2FC98FDD5" box="[758,768,589,613]" pageId="14" pageNumber="15" value="1923-02-25">2</date>
</emphasis>
23–25;
<emphasis id="B921DC4FFFB4FFF2E119FDB2FCEFFDD5" box="[860,887,589,613]" italics="true" pageId="14" pageNumber="15">h1</emphasis>
60;
<emphasis id="B921DC4FFFB4FFF2E1EFFDB2FC5EFDD5" box="[938,966,589,613]" italics="true" pageId="14" pageNumber="15">h2</emphasis>
40–41; distance:
<emphasis id="B921DC4FFFB4FFF2E6C8FDB0FB5BFDD6" box="[1165,1219,590,614]" italics="true" pageId="14" pageNumber="15">
vi–
<date id="FFEB269DFFB4FFF2E6EAFDB0FB5BFDD6" box="[1199,1219,591,614]" pageId="14" pageNumber="15" value="1919-06">vi</date>
</emphasis>
19; first interspace (
<emphasis id="B921DC4FFFB4FFF2E2E5FD8AFF2CFD20" box="[160,180,629,656]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
to
<emphasis id="B921DC4FFFB4FFF2E29DFD8BFF7EFD3C" box="[216,230,628,652]" italics="true" pageId="14" pageNumber="15">d</emphasis>
) 38–46. Genital region not developed; three pairs of legs and basifemora not divided. Dorsally posterior annals have not yet developed. Ventrally three pairs of anal setae. Leg chaetotaxy differs from that of female as follows: coxae
<date id="FFEB269DFFB4FFF2E360FD3FFEF6FD67" box="[293,366,703,727]" pageId="14" pageNumber="15">I–III 2</date>
sts-1 sts-0; trochanters I–III 0-0-0; femora
<date id="FFEB269DFFB4FFF2E118FD3FFC26FD67" box="[861,958,703,727]" pageId="14" pageNumber="15">I–III 6-2</date>
-4 sts; genua
<date id="FFEB269DFFB4FFF2E613FD3FFB38FD67" box="[1110,1184,703,727]" pageId="14" pageNumber="15">I–III 5</date>
sts, 1σ -5-5 sts; tibiae
<date id="FFEB269DFFB4FFF2E2D2FD1BFF47FD4B" box="[151,223,739,763]" pageId="14" pageNumber="15">I–III 5</date>
sts, 2φ, 1ĸ -3 sts, 2φ -5
<emphasis id="B921DC4FFFB4FFF2E3ABFD1BFD93FD4B" box="[494,523,740,763]" italics="true" pageId="14" pageNumber="15">sts</emphasis>
, 1φ; tarsi
<date id="FFEB269DFFB4FFF2E03BFD1BFD4CFD4B" box="[638,724,739,763]" pageId="14" pageNumber="15" value="1913-01" valueMax="1913-03-31" valueMin="1913-01-13">I–III 13</date>
sts, 1ω -13 sts, 1ω -10 sts, 1tr.
</paragraph>
</subSubSection>
<subSubSection id="C34F53D6FFB4FFF2E28CFCF8FCDAFCD7" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BEA005DFFB4FFF2E28CFCF8FCDAFCD7" blockId="14.[151,1436,151,1055]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CFCF8FE1BFCAF" bold="true" box="[201,387,774,799]" pageId="14" pageNumber="15">
<typeStatus id="54EEBEFFFFB4FFF2E28CFCF8FE9CFCAF" box="[201,260,775,799]" pageId="14" pageNumber="15">Type</typeStatus>
material:
</emphasis>
<collectingCountry id="F34240CDFFB4FFF2E3D1FCF8FDF4FCAF" box="[404,620,775,799]" name="South Africa" pageId="14" pageNumber="15">SOUTH AFRICA</collectingCountry>
, KwaZulu/
<collectingCountry id="F34240CDFFB4FFF2E0B7FCF8FCB7FCAF" box="[754,815,775,799]" name="South Africa" pageId="14" pageNumber="15">Natal</collectingCountry>
Province:
<typeStatus id="54EEBEFFFFB4FFF2E1F9FCF8FBBCFCAF" box="[956,1060,775,799]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
female,
<specimenCount id="9D53CBD4FFB4FFF2E6DCFCF8FA9CFCAF" box="[1177,1284,775,799]" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
<typeStatus id="54EEBEFFFFB4FFF2E750FCF7FAE0FC90" box="[1301,1400,776,800]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
, 3 tritonymph
<typeStatus id="54EEBEFFFFB4FFF2E35FFCD3FE10FCF4" box="[282,392,812,836]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, 2 deutonymph
<typeStatus id="54EEBEFFFFB4FFF2E07EFCD3FD31FCF4" box="[571,681,812,836]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, 2 protonymph
<typeStatus id="54EEBEFFFFB4FFF2E11DFCD3FC5BFCF4" box="[856,963,812,836]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
and 3 larva
<typeStatus id="54EEBEFFFFB4FFF2E608FCD3FB23FCF4" box="[1101,1211,812,836]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
, Pongola, from soil,
<date id="FFEB269DFFB4FFF2E2D2FCB0FECEFCD7" box="[151,342,847,871]" pageId="14" pageNumber="15" value="1968-10-29">29 October 1968</date>
, Edenville, Free State, Coll. N. de L. Genis.
</paragraph>
</subSubSection>
<subSubSection id="C34F53D6FFB4FFF2E28CFC8CFA0DFBAB" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BEA005DFFB4FFF2E28CFC8CFCC2FC3B" blockId="14.[151,1436,151,1055]" box="[201,858,883,908]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CFC8CFED7FC3B" bold="true" box="[201,335,883,907]" pageId="14" pageNumber="15">Etymology.</emphasis>
The species was named after the
<typeStatus id="54EEBEFFFFB4FFF2E08DFC8BFD60FC3C" box="[712,760,884,908]" pageId="14" pageNumber="15">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BEA005DFFB4FFF2E28CFC69FA0DFBAB" blockId="14.[151,1436,151,1055]" pageId="14" pageNumber="15">
<emphasis id="B921DC4FFFB4FFF2E28CFC69FE4AFC1F" bold="true" box="[201,466,918,943]" pageId="14" pageNumber="15">Differential diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C557BDEFFB4FFF2E0AFFC68FBE9FC1F" authority="Meyer &amp; Ryke" authorityName="Meyer &amp; Ryke" box="[746,1137,919,943]" class="Arachnida" family="Bdellidae" genus="Spinibdella" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="thori">
<emphasis id="B921DC4FFFB4FFF2E0AFFC68FC31FC1F" box="[746,937,919,943]" italics="true" pageId="14" pageNumber="15">Spinibdella thori</emphasis>
(Meyer &amp; Ryke)
</taxonomicName>
but can be recognized on account of the transverse striations that curve anteromedially on the prodorsum, as seen in figure 31; tibiae I with two closely associated attenuate solenidia; tibiae II with one attenuate and one reduced blunt solenidia; tarsi II with one blunt solenidion in a distal position and genua I with an attenuate solenidion. Setae
<emphasis id="B921DC4FFFB4FFF2E621FBFBFBE1FBAF" box="[1124,1145,1028,1055]" italics="true" pageId="14" pageNumber="15">c2</emphasis>
about twice as long as
<emphasis id="B921DC4FFFB4FFF2E73FFBFBFA17FBAF" box="[1402,1423,1028,1055]" italics="true" pageId="14" pageNumber="15">c1</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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