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<document id="ACC53AA0C8BBA48BA5B5724EF8EBB355" ID-DOI="10.11646/zootaxa.3688.1.1" ID-GBIF-Dataset="151481a0-71e1-4402-ab7e-8262caddf539" ID-ISSN="1175-5326" ID-Zenodo-Dep="284201" ID-ZooBank="DB84C679-C2AA-41CC-816E-83E2F6DCA391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460649331738" checkinUser="plazi" docAuthor="Haddad, Charles R." docDate="2013" docId="03DB8445FFA9FF84FF2AF9873560FC54" docLanguage="en" docName="zt03688p079.pdf" docOrigin="Zootaxa 3688 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Copuetta maputa Haddad, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="55" masterDocId="FFE2FC3DFF9AFFB2FFBDFFE6345DFFF0" masterDocTitle="Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae" masterLastPageNumber="79" masterPageNumber="1" pageNumber="52" updateTime="1698337697679" updateUser="plazi">
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<mods:title id="4AAD156B29B06BCAA0218D227093F8D1">Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae</mods:title>
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<mods:namePart id="E030B912B8C303FCC59D05922792109B">Haddad, Charles R.</mods:namePart>
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<mods:date id="EA6D06A173CC65D08C5C0BF865DFEEDB">2013</mods:date>
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<treatment id="03DB8445FFA9FF84FF2AF9873560FC54" ID-DOI="http://doi.org/10.5281/zenodo.6153951" ID-GBIF-Taxon="127664918" ID-Zenodo-Dep="6153951" LSID="urn:lsid:plazi:treatment:03DB8445FFA9FF84FF2AF9873560FC54" httpUri="http://treatment.plazi.org/id/03DB8445FFA9FF84FF2AF9873560FC54" lastPageId="54" lastPageNumber="55" pageId="51" pageNumber="52">
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<paragraph id="8BCD3553FFA9FF81FF2AF9873597F98B" blockId="51.[151,779,1633,1692]" box="[151,458,1633,1659]" pageId="51" pageNumber="52">
<heading id="D085823FFFA9FF81FF2AF9873597F98B" bold="true" box="[151,458,1633,1659]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="B906E941FFA9FF81FF2AF9873597F98B" bold="true" box="[151,458,1633,1659]" pageId="51" pageNumber="52">
<taxonomicName id="4C724ED0FFA9FF81FF2AF987353BF98B" ID-CoL="Y8YL" box="[151,358,1633,1659]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="maputa" status="sp. nov.">
<emphasis id="B906E941FFA9FF81FF2AF987353BF98B" bold="true" box="[151,358,1633,1659]" italics="true" pageId="51" pageNumber="52">Copuetta maputa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235543AFFA9FF81FED0F9873597F98B" box="[365,458,1633,1659]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD3553FFA9FF81FF2AF9623756F96C" blockId="51.[151,779,1633,1692]" box="[151,779,1668,1692]" pageId="51" pageNumber="52">
<figureCitation id="134929D6FFA9FF81FF2AF962354BF96C" box="[151,278,1668,1692]" captionStart="FIGURES 7 13" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[206,1374,772,1904]" captionTargetId="figure@5.[206,1374,772,1904]" captionTargetPageId="5" captionText="FIGURES 7 13. General habitus of Afrotropical Copuetta gen. nov. species: 7. C. erecta sp. nov. male, Chidenguele, Mozambique. 8. C. erecta sp. nov. female, Xai-Xai, Mozambique. 9. C. lacustris (Strand, 1916) female, Bloemfontein, South Africa. 10. C. lotzi sp. nov. male, Bloemfontein. 11. C. magna sp. nov. male, Ndumo Game Reserve, South Africa. 12. C. maputa sp. nov. male, Bilene, Mozambique. 13. C. maputa sp. nov. female, Chidenguele." httpUri="https://zenodo.org/record/284203/files/figure.png" pageId="51" pageNumber="52">Figs 12, 13</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FE9FF962351CF96C" box="[290,321,1668,1692]" captionStart="FIGURES 14 25" captionStartId="9.[151,264,1609,1632]" captionTargetBox="[156,1436,193,1586]" captionTargetId="figure@9.[156,1441,193,1588]" captionTargetPageId="9" captionText="FIGURES 14 25. Scanning electron microscope photos of Cambalida dippenaarae Haddad, 2012 (14, 21, 23), Copa flavoplumosa Simon, 1885 (15, 17, 24), Apochinomma formicaeforme Pavesi, 1881 (16, 18, 19), Copuetta maputa sp. nov. (20) and Merenius alberti Lessert, 1923 (22, 25): 14 16. Carapace, anterior view. 17. Male, promarginal cheliceral curved setae. 18. Female, promarginal cheliceral curved setae. 19. Anterior of female abdomen, indicating epigastric sclerite and petiolate pedicel. 20, 21. Male emboli, stippled lines indicating width (W) and length (L) of embolus from base to tip, arrow indicating point from which basal width is measured (retrolateral bend of distal end of tegulum). 22. Rod-like setae on male palpal cymbium, indicated by arrows. 23. Distal thickened spiniform setae arranged in three rows on male palpal cymbium. 24. Female epigyne, ventral view, indicating copulatory opening (CO) and coiled sclerotised epigynal ridge (ER). 25. Female epigyne, dorsal view, indicating spermathecae I and II (ST I and ST II, respectively) and copulatory duct (CD)." httpUri="https://zenodo.org/record/284204/files/figure.png" pageId="51" pageNumber="52">20</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FEF1F96235C9F96C" box="[332,404,1668,1692]" captionStart="FIGURES 45 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="51" pageNumber="52">56, 57</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FE1DF96235E3F96C" box="[416,446,1668,1692]" captionStart="FIGURES 62 67" captionStartId="25.[151,264,906,929]" captionTargetBox="[156,1436,191,880]" captionTargetId="figure@25.[156,1441,191,881]" captionTargetPageId="25" captionText="FIGURES 62 67. Scanning electron microscope photographs of Copuetta lacustris (Strand, 1916) female (62, 64, 66) and male (63, 65) and C. maputa sp. nov. female (67): 62. Dorsal carapace setae. 63, 64. Eye region and clypeus in dorsal (63) and anterior (64) views. 65, 66. Cheliceral promarginal curved setae, anterior view. 67. endite, ventral view." httpUri="https://zenodo.org/record/284211/files/figure.png" pageId="51" pageNumber="52">67</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FE77F962364EF96C" box="[458,531,1668,1692]" captionStart="FIGURES 86 91" captionStartId="28.[151,264,1782,1805]" captionTargetBox="[151,1436,193,1760]" captionTargetId="figure@28.[151,1436,193,1761]" captionTargetPageId="28" captionText="FIGURES 86 91. Scanning electron microscope photographs of Copuetta magna sp. nov. male (86 88) and C. maputa sp. nov. female (89 91) spinneret morphology: 86, 89. Anterior lateral spinneret. 87, 90. Posterior median spinneret. 88, 91. Posterior lateral spinneret. Abbreviations: Ac aciniform gland spigot (s); Cy cylindrical gland spigot (s); MAmp major ampullate gland spigot (s); mAmp minor ampullate gland spigot (s); n nubbin (s); Pi piriform gland spigot; ta tartipore." httpUri="https://zenodo.org/record/284214/files/figure.png" pageId="51" pageNumber="52">8991</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FDA3F962363BF96C" box="[542,614,1668,1692]" captionStart="FIGURES 92 98" captionStartId="29.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1582]" captionTargetId="figure@29.[151,1436,193,1584]" captionTargetPageId="29" captionText="FIGURES 92 98. Scanning electron microscope photographs of Copuetta gen. nov. genitalia: 92 96. Male palpal emboli of C. erecta sp. nov. (92), C. lacustris (Strand, 1916) (93), C. lotzi sp. nov. (94), C. magna sp. nov. (95) and C. maputa sp. nov. (96). 97, 98. Female epigynes of C. lacustris (97) and C. maputa sp. nov. (98)." httpUri="https://zenodo.org/record/284215/files/figure.png" pageId="51" pageNumber="52">96, 98</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FDCFF96236C3F96C" box="[626,670,1668,1692]" captionStart="FIGURES 99 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="51" pageNumber="52">106</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FD17F9623756F96C" box="[682,779,1668,1692]" captionStart="FIGURES 141 144" captionStartId="53.[151,264,1734,1757]" captionTargetBox="[165,1424,789,1699]" captionTargetId="figure@53.[151,1436,774,1712]" captionTargetPageId="53" captionText="FIGURES 141 144. Genitalic morphology of Copuetta maputa sp. nov.: 141. Male palp, ventral view. 142. Same, retrolateral view. 143. Female epigyne, ventral view. 144. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284226/files/figure.png" pageId="51" pageNumber="52">141144</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA9FF81FF2AF92B3539F8F0" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="8BCD3553FFA9FF81FF2AF92B3539F8F0" blockId="51.[151,1437,1740,1793]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="AB62127DFFA9FF81FF2AF92B3539F8F0" pageId="51" pageNumber="52">
<taxonomicName id="4C724ED0FFA9FF81FF2AF92B3649F911" ID-CoL="427TN" authority="Simon, 1898: 214" authorityName="Simon" authorityPageNumber="214" authorityYear="1898" box="[151,532,1740,1762]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="B906E941FFA9FF81FF2AF92B3512F912" box="[151,335,1741,1762]" italics="true" pageId="51" pageNumber="52">Messapus martini</emphasis>
<treatmentCitation id="0AD31342FFA9FF81FEEAF92A3649F911" author="Simon" box="[343,532,1740,1762]" page="214" pageId="51" pageNumber="52" year="1898">
<bibRefCitation id="EFE348A2FFA9FF81FEEAF92A3582F911" author="Simon" box="[343,479,1740,1762]" pageId="51" pageNumber="52" refString="Simon, E. (1898) Histoire naturelle des araignees. Tome 2, Librairie Encyclopedique de Roret, Paris, pp. 193 - 380." type="journal article" year="1898">Simon, 1898</bibRefCitation>
: 214
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AD31342FFA9FF81FD9FF92A37DDF911" author="Bosselaers" box="[546,896,1740,1762]" page="307" pageId="51" pageNumber="52" year="2000">
<bibRefCitation id="EFE348A2FFA9FF81FD9FF92A3716F911" author="Bosselaers" box="[546,843,1740,1762]" pageId="51" pageNumber="52" refString="Bosselaers, J. &amp; Jocque, R. (2000 a) Studies in Corinnidae: transfer of four genera and descriptions of the female of Lessertina mutica Lawrence 1942. Tropical Zoology, 13, 305 - 325. http: // dx. doi. org / 10.1080 / 03946975.2000.10531138" type="journal article" year="2000" yearSuffix="a">Bosselaers &amp; Jocqué, 2000a</bibRefCitation>
: 307
</treatmentCitation>
, fig. 1AH (3 misidentified, see remarks under
<taxonomicName id="4C724ED0FFA9FF81FA3DF92B354FF8F1" ID-CoL="427TN" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="B906E941FFA9FF81FA3DF92B354FF8F1" italics="true" pageId="51" pageNumber="52">M. martini</emphasis>
</taxonomicName>
above).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA9FF81FF2AF8C8378AF899" pageId="51" pageNumber="52" type="etymology">
<paragraph id="8BCD3553FFA9FF81FF2AF8C8378AF899" blockId="51.[151,1436,1837,2006]" pageId="51" pageNumber="52">
<emphasis id="B906E941FFA9FF81FF2AF8C8357CF8B6" bold="true" box="[151,289,1838,1862]" pageId="51" pageNumber="52">Etymology:</emphasis>
The species name is derived from the Maputaland ecoregion in southern
<collectingCountry id="F36575C3FFA9FF81FBE3F8CB30ACF8B5" box="[1118,1265,1837,1861]" name="Mozambique" pageId="51" pageNumber="52">Mozambique</collectingCountry>
and KwaZulu-
<collectingCountry id="F36575C3FFA9FF81FF2AF8B73489F899" box="[151,212,1873,1897]" name="South Africa" pageId="51" pageNumber="52">Natal</collectingCountry>
Province,
<collectingCountry id="F36575C3FFA9FF81FEF1F8B73582F899" box="[332,479,1873,1897]" name="South Africa" pageId="51" pageNumber="52">South Africa</collectingCountry>
, to which this species is apparently endemic.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA9FF86FF7AF893311EFF07" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BCD3553FFA9FF86FF7AF893311EFF07" blockId="51.[151,1436,1837,2006]" lastBlockId="52.[151,1437,151,2034]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis id="B906E941FFA9FF81FF7AF893351EF87E" bold="true" box="[199,323,1909,1934]" pageId="51" pageNumber="52">Diagnosis:</emphasis>
Males of this species are closely related to
<taxonomicName id="4C724ED0FFA9FF81FCFEF890378CF87E" box="[835,977,1910,1934]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="comorica" status="sp. nov.">
<emphasis id="B906E941FFA9FF81FCFEF890378CF87E" box="[835,977,1910,1934]" italics="true" pageId="51" pageNumber="52">C. comorica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235543AFFA9FF81FC67F890306BF87E" box="[986,1078,1910,1934]" pageId="51" pageNumber="52" rank="species">
<emphasis id="B906E941FFA9FF81FC67F890306BF87E" bold="true" box="[986,1078,1910,1934]" pageId="51" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C724ED0FFA9FF81FBCFF89030AFF87E" box="[1138,1266,1910,1934]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="wagneri" status="sp. nov.">
<emphasis id="B906E941FFA9FF81FBCFF89030AFF87E" box="[1138,1266,1910,1934]" italics="true" pageId="51" pageNumber="52">C. wagneri</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA9FF81FB46F890310AF87E" bold="true" box="[1275,1367,1910,1934]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A235543AFFA9FF81FB46F890310AF87E" box="[1275,1367,1910,1934]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
In
<taxonomicName id="4C724ED0FFA9FF81FA3FF89034B6F841" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="maputa" status="sp. nov.">
<emphasis id="B906E941FFA9FF81FA3FF89034B6F841" italics="true" pageId="51" pageNumber="52">C. maputa</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA9FF81FF4EF87C3513F842" bold="true" box="[243,334,1946,1970]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A235543AFFA9FF81FF4EF87C3513F842" box="[243,334,1946,1970]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the embolus base is quite long and upright and the embolus tip is short and directed prolaterally (
<figureCitation id="134929D6FFA9FF81FF22F85B34ABF825" box="[159,246,1981,2005]" captionStart="FIGURES 14 25" captionStartId="9.[151,264,1609,1632]" captionTargetBox="[156,1436,193,1586]" captionTargetId="figure@9.[156,1441,193,1588]" captionTargetPageId="9" captionText="FIGURES 14 25. Scanning electron microscope photos of Cambalida dippenaarae Haddad, 2012 (14, 21, 23), Copa flavoplumosa Simon, 1885 (15, 17, 24), Apochinomma formicaeforme Pavesi, 1881 (16, 18, 19), Copuetta maputa sp. nov. (20) and Merenius alberti Lessert, 1923 (22, 25): 14 16. Carapace, anterior view. 17. Male, promarginal cheliceral curved setae. 18. Female, promarginal cheliceral curved setae. 19. Anterior of female abdomen, indicating epigastric sclerite and petiolate pedicel. 20, 21. Male emboli, stippled lines indicating width (W) and length (L) of embolus from base to tip, arrow indicating point from which basal width is measured (retrolateral bend of distal end of tegulum). 22. Rod-like setae on male palpal cymbium, indicated by arrows. 23. Distal thickened spiniform setae arranged in three rows on male palpal cymbium. 24. Female epigyne, ventral view, indicating copulatory opening (CO) and coiled sclerotised epigynal ridge (ER). 25. Female epigyne, dorsal view, indicating spermathecae I and II (ST I and ST II, respectively) and copulatory duct (CD)." httpUri="https://zenodo.org/record/284204/files/figure.png" pageId="51" pageNumber="52">Figs 20</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FEBFF85B357CF825" box="[258,289,1981,2005]" captionStart="FIGURES 92 98" captionStartId="29.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1582]" captionTargetId="figure@29.[151,1436,193,1584]" captionTargetPageId="29" captionText="FIGURES 92 98. Scanning electron microscope photographs of Copuetta gen. nov. genitalia: 92 96. Male palpal emboli of C. erecta sp. nov. (92), C. lacustris (Strand, 1916) (93), C. lotzi sp. nov. (94), C. magna sp. nov. (95) and C. maputa sp. nov. (96). 97, 98. Female epigynes of C. lacustris (97) and C. maputa sp. nov. (98)." httpUri="https://zenodo.org/record/284215/files/figure.png" pageId="51" pageNumber="52">96</figureCitation>
,
<figureCitation id="134929D6FFA9FF81FE90F85B3506F825" box="[301,347,1981,2005]" captionStart="FIGURES 99 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="51" pageNumber="52">106</figureCitation>
). In
<taxonomicName id="4C724ED0FFA9FF81FE30F8583644F826" box="[397,537,1982,2006]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="comorica" status="sp. nov.">
<emphasis id="B906E941FFA9FF81FE30F8583644F826" box="[397,537,1982,2006]" italics="true" pageId="51" pageNumber="52">C. comorica</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA9FF81FD9DF8583627F826" bold="true" box="[544,634,1982,2006]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A235543AFFA9FF81FD9DF8583627F826" box="[544,634,1982,2006]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the base is very short and the embolus tip is directed distally (
<figureCitation id="134929D6FFA9FF81FA86F85B31D2F825" box="[1339,1423,1981,2005]" captionStart="FIGURES 99 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="51" pageNumber="52">Fig. 99</figureCitation>
), while in
<taxonomicName id="4C724ED0FFAEFF86FF44FF7E3528FF40" box="[249,373,152,176]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="wagneri" status="sp. nov.">
<emphasis id="B906E941FFAEFF86FF44FF7E3528FF40" box="[249,373,152,176]" italics="true" pageId="52" pageNumber="53">C. wagneri</emphasis>
</taxonomicName>
<emphasis id="B906E941FFAEFF86FEC1FF7E3588FF40" bold="true" box="[380,469,152,176]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="A235543AFFAEFF86FEC1FF7E3588FF40" box="[380,469,152,176]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the embolus has an oblique base and a narrow elongate tip that is directed at the distal end of the cymbium (
<figureCitation id="134929D6FFAEFF86FE3BFF5A35B5FF24" box="[390,488,188,212]" captionStart="FIGURES 99 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="52" pageNumber="53">Fig. 109</figureCitation>
). Females have an epigyne with opposing lateral semi-circular ridges that are clearly separated (
<figureCitation id="134929D6FFAEFF86FEADFF39352CFF07" box="[272,369,223,247]" captionStart="FIGURES 141 144" captionStartId="53.[151,264,1734,1757]" captionTargetBox="[165,1424,789,1699]" captionTargetId="figure@53.[151,1436,774,1712]" captionTargetPageId="53" captionText="FIGURES 141 144. Genitalic morphology of Copuetta maputa sp. nov.: 141. Male palp, ventral view. 142. Same, retrolateral view. 143. Female epigyne, ventral view. 144. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284226/files/figure.png" pageId="52" pageNumber="53">Fig. 143</figureCitation>
), while those of
<taxonomicName id="4C724ED0FFAEFF86FD96FF0636E8FF08" box="[555,693,224,248]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="comorica" status="sp. nov.">
<emphasis id="B906E941FFAEFF86FD96FF0636E8FF08" box="[555,693,224,248]" italics="true" pageId="52" pageNumber="53">C. comorica</emphasis>
</taxonomicName>
<emphasis id="B906E941FFAEFF86FD01FF06374BFF08" bold="true" box="[700,790,224,248]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="A235543AFFAEFF86FD01FF06374BFF08" box="[700,790,224,248]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
meet along the midline of the epigyne (
<figureCitation id="134929D6FFAEFF86FB68FF39316BFF07" box="[1237,1334,223,247]" captionStart="FIGURES 110 113" captionStartId="32.[151,264,1213,1236]" captionTargetBox="[169,1421,205,1162]" captionTargetId="figure@32.[151,1436,195,1187]" captionTargetPageId="32" captionText="FIGURES 110 113. Genitalic morphology of Copuetta comorica sp. nov.: 110. Male palp, ventral view. 111. Same, retrolateral view. 112. Female epigyne, ventral view. 113. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284217/files/figure.png" pageId="52" pageNumber="53">Fig. 112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFAEFF87FF7AFEE23663FD21" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="description">
<paragraph id="8BCD3553FFAEFF86FF7AFEE230CAFE94" blockId="52.[151,1437,151,2034]" pageId="52" pageNumber="53">
<emphasis id="B906E941FFAEFF86FF7AFEE2372EFEEC" bold="true" box="[199,883,259,284]" pageId="52" pageNumber="53">
Male (
<typeStatus id="54C98BF1FFAEFF86FEAEFEE53520FEEC" box="[275,381,259,284]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
, Ndumo, NCA 2006/425). Measurements:
</emphasis>
CL 3.75, CW 2.80, AL 3.90, AW 2.43, TL 7.75 (6.158.90), FL 0.46, SL 1.62, SW 1.57, AMEAME 0.05, AMEALE 0.01, ALEALE 0.53, PMEPME 0.16, PMEPLE 0.08, PLEPLE 0.63,
<collectingCountry id="F36575C3FFAEFF86FDB0FEAB3604FE94" box="[525,601,333,356]" name="Peru" pageId="52" pageNumber="53">PERW</collectingCountry>
0.97, MOQAW 0.59, MOQPW 0.57, MOQL 0.70.
</paragraph>
<paragraph id="8BCD3553FFAEFF86FF7AFE8930DBFE5C" blockId="52.[151,1437,151,2034]" pageId="52" pageNumber="53">Length of leg segments: I 3.23 + 1.40 + 2.60 + 2.73 + 1.50 = 11.46; II 3.20 + 1.35 + 2.48 + 2.60 + 1.40 = 11.03; III 3.10 + 1.35 + 2.30 + 2.90 + 1.27 = 10.92; IV 3.90 + 1.44 + 3.05 + 4.25 + 1.45 = 14.09.</paragraph>
<paragraph id="8BCD3553FFAEFF86FF7AFE5136E2F9C1" blockId="52.[151,1437,151,2034]" pageId="52" pageNumber="53">
General appearance as in
<figureCitation id="134929D6FFAEFF86FE4EFE513614FE3F" box="[499,585,439,463]" captionStart="FIGURES 7 13" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[206,1374,772,1904]" captionTargetId="figure@5.[206,1374,772,1904]" captionTargetPageId="5" captionText="FIGURES 7 13. General habitus of Afrotropical Copuetta gen. nov. species: 7. C. erecta sp. nov. male, Chidenguele, Mozambique. 8. C. erecta sp. nov. female, Xai-Xai, Mozambique. 9. C. lacustris (Strand, 1916) female, Bloemfontein, South Africa. 10. C. lotzi sp. nov. male, Bloemfontein. 11. C. magna sp. nov. male, Ndumo Game Reserve, South Africa. 12. C. maputa sp. nov. male, Bilene, Mozambique. 13. C. maputa sp. nov. female, Chidenguele." httpUri="https://zenodo.org/record/284203/files/figure.png" pageId="52" pageNumber="53">Figs 12</figureCitation>
and
<figureCitation id="134929D6FFAEFF86FD38FE5136F9FE3F" box="[645,676,439,463]" captionStart="FIGURES 45 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="52" pageNumber="53">56</figureCitation>
. Carapace bright yellow, eye region black, entire clypeus mottled black; paired black mediolateral markings from palpal coxae to posterior slope; pale along midline from between PME to posterior of carapace, with faint black mottling between mediolateral markings from PER to midpoint of carapace and with narrow mottled black marking in front of fovea; striae orange, radiating from fovea, directed between coxae, covered in dense black mottling within mediolateral markings; lateral margins with faint black fringe, with indistinct black spots between leg coxae; markings covered with black feathery setae, areas between them covered in white feathery and short straight setae. AME separated by distance slightly less than 15 their diameter; AME touching ALE; clypeus height equal to AME diameter; PME slightly larger than PLE; PME separated by distance equal to ½ their diameter; PME separated from PLE by distance equal to ½ PLE diameter; CW:
<collectingCountry id="F36575C3FFAEFF86FF7AFCE7354AFCE9" box="[199,279,769,793]" name="Peru" pageId="52" pageNumber="53">PERW</collectingCountry>
= 2.89:1. Chelicerae dark brown with black mottling, yellow prolaterally distally; promargin with two teeth separated by the basal width of proximal tooth, distal tooth much larger; retromargin with two teeth separated by ½ their basal width, distal tooth slightly larger than proximal tooth, close to fang base. Endites dark yellowbrown with faint black mottling, white prolaterally distally; labium orange-brown, creamy-orange distally; sternum bright yellow-orange, slightly darker around borders. Legs with faint lateral mottling and faint black spots at spine bases; markings covered in black feathery setae, with yellowish feathery setae between them; femora IIV yellow, III and IV with faint distal band; patellae IIV yellow with lateral mottling; tibiae IIII yellow and IV yelloworange, all with very faint broad median band; metatarsi IIV pale orange-brown with very faint proximal, median and distal bands; tarsi IIV yellow-orange. Leg spination: femora: I pl 3 do 3 rl 1, II pl 2 do 3 rl 1, III pl 2 do 3 rl 2, IV pl 2 do 3 rl 2; all femora with scattered erect ventral setae; patellae: I and II with fine proximal and distal do setae, III and IV with proximal seta and distal spine dorsally; tibiae: I pl 2 plv 2 rlv 2, II pl 2 plv 2 rlv 2, III pl 2 do 1 rl 2 plv 2 rlv 2, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I plv 1-2 rlv 2, II plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 5 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 1 plv 1; tarsi: pl 1 plv 3. Abdomen with yellow anterior dorsal scutum extending to ½ abdomen length; dorsum mottled black anteriorly, laterally and posteriorly, with small white blotches and streaks laterally; dorsum pale along midline, with broadened cream patch behind scutum, with scattered small black blotches; markings covered in black feathery and short straight setae, areas between them with white feathery setae and brown scattered short straight setae; spinnerets surrounded by narrow black ring dorsally and laterally, absent ventrally; venter cream, densely covered in white feathery setae and scattered short straight setae; epigastric scutum weakly sclerotised, creamy-yellow; ventral sclerite absent; inframamillary sclerite yellow-orange. Male palp yellow, cymbium yellow-brown; tegulum dark orange-brown, with dark red-brown ducts; embolus with narrow base and two compressed coils, of which the distal coil is much broader than the proximal; embolus tip sharp and triangular, directed prolaterally (
<figureCitation id="134929D6FFAEFF86FE35F9FF3582F9C1" box="[392,479,1561,1585]" captionStart="FIGURES 14 25" captionStartId="9.[151,264,1609,1632]" captionTargetBox="[156,1436,193,1586]" captionTargetId="figure@9.[156,1441,193,1588]" captionTargetPageId="9" captionText="FIGURES 14 25. Scanning electron microscope photos of Cambalida dippenaarae Haddad, 2012 (14, 21, 23), Copa flavoplumosa Simon, 1885 (15, 17, 24), Apochinomma formicaeforme Pavesi, 1881 (16, 18, 19), Copuetta maputa sp. nov. (20) and Merenius alberti Lessert, 1923 (22, 25): 14 16. Carapace, anterior view. 17. Male, promarginal cheliceral curved setae. 18. Female, promarginal cheliceral curved setae. 19. Anterior of female abdomen, indicating epigastric sclerite and petiolate pedicel. 20, 21. Male emboli, stippled lines indicating width (W) and length (L) of embolus from base to tip, arrow indicating point from which basal width is measured (retrolateral bend of distal end of tegulum). 22. Rod-like setae on male palpal cymbium, indicated by arrows. 23. Distal thickened spiniform setae arranged in three rows on male palpal cymbium. 24. Female epigyne, ventral view, indicating copulatory opening (CO) and coiled sclerotised epigynal ridge (ER). 25. Female epigyne, dorsal view, indicating spermathecae I and II (ST I and ST II, respectively) and copulatory duct (CD)." httpUri="https://zenodo.org/record/284204/files/figure.png" pageId="52" pageNumber="53">Figs 20</figureCitation>
,
<figureCitation id="134929D6FFAEFF86FE56F9FF3654F9C1" box="[491,521,1561,1585]" captionStart="FIGURES 92 98" captionStartId="29.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1582]" captionTargetId="figure@29.[151,1436,193,1584]" captionTargetPageId="29" captionText="FIGURES 92 98. Scanning electron microscope photographs of Copuetta gen. nov. genitalia: 92 96. Male palpal emboli of C. erecta sp. nov. (92), C. lacustris (Strand, 1916) (93), C. lotzi sp. nov. (94), C. magna sp. nov. (95) and C. maputa sp. nov. (96). 97, 98. Female epigynes of C. lacustris (97) and C. maputa sp. nov. (98)." httpUri="https://zenodo.org/record/284215/files/figure.png" pageId="52" pageNumber="53">96</figureCitation>
,
<figureCitation id="134929D6FFAEFF86FDA9F9FF361DF9C1" box="[532,576,1561,1585]" captionStart="FIGURES 99 109" captionStartId="30.[151,264,1595,1618]" captionTargetBox="[153,1436,183,1577]" captionTargetId="figure@30.[153,1438,183,1578]" captionTargetPageId="30" captionText="FIGURES 99 109. Digital microscope photographs of left male emboli of Copuetta gen. nov. species in ventral view: 99. C. comorica sp. nov.. 100. C. erecta sp. nov.. 101. C. kwamgumi sp. nov.. 102. C. lacustris (Strand, 1916). 103. C. lesnei sp. nov. (mirror image of right embolus). 104. C. lotzi sp. nov.. 105. C. magna sp. nov.. 106. C. maputa sp. nov.. 107. C. naja sp. nov.. 108. C. uzungwa sp. nov.. 109. C. wagneri sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284216/files/figure.png" pageId="52" pageNumber="53">106</figureCitation>
,
<figureCitation id="134929D6FFAEFF86FDF1F9FF36EFF9C1" box="[588,690,1561,1585]" captionStart="FIGURES 141 144" captionStartId="53.[151,264,1734,1757]" captionTargetBox="[165,1424,789,1699]" captionTargetId="figure@53.[151,1436,774,1712]" captionTargetPageId="53" captionText="FIGURES 141 144. Genitalic morphology of Copuetta maputa sp. nov.: 141. Male palp, ventral view. 142. Same, retrolateral view. 143. Female epigyne, ventral view. 144. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284226/files/figure.png" pageId="52" pageNumber="53">141, 142</figureCitation>
).
</paragraph>
<paragraph id="8BCD3553FFAEFF86FF7AF9D8308BF96D" blockId="52.[151,1437,151,2034]" pageId="52" pageNumber="53">
<emphasis id="B906E941FFAEFF86FF7AF9D8378FF9A6" bold="true" box="[199,978,1597,1622]" pageId="52" pageNumber="53">
Female (
<typeStatus id="54C98BF1FFAEFF86FE93F9DB35C0F9A6" box="[302,413,1597,1622]" pageId="52" pageNumber="53" type="paratype">paratype</typeStatus>
, Chidenguele, NCA 2008/200). Measurements:
</emphasis>
CL 3.35, CW 2.63, AL 3.70, AW 2.20, TL 7.20 (7.208.75), FL 0.37, SL 1.59, SW 1.51, AMEAME 0.06, AMEALE 0.01, ALEALE 0.54, PMEPME 0.12, PMEPLE 0.06, PLEPLE 0.59,
<collectingCountry id="F36575C3FFAEFF86FDF0F96036C4F96D" box="[589,665,1670,1693]" name="Peru" pageId="52" pageNumber="53">PERW</collectingCountry>
0.90, MOQAW 0.53, MOQPW 0.52, MOQL 0.62.
</paragraph>
<paragraph id="8BCD3553FFAEFF86FF7AF94F30D8F915" blockId="52.[151,1437,151,2034]" pageId="52" pageNumber="53">Length of leg segments: I 3.00 + 1.35 + 2.40 + 2.55 + 1.40 = 10.70; II 2.95 + 1.34 + 2.30 + 2.43 + 1.35 = 10.37; III 2.87 + 1.30 + 2.11 + 2.65 + 1.23 = 10.16; IV 3.60 + 1.42 + 2.88 + 3.95 + 1.25 = 13.10.</paragraph>
<paragraph id="8BCD3553FFAEFF87FF7AF9173663FD21" blockId="52.[151,1437,151,2034]" lastBlockId="53.[151,1437,151,721]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
General appearance as in
<figureCitation id="134929D6FFAEFF86FE4BF9173610F8F9" box="[502,589,1777,1801]" captionStart="FIGURES 7 13" captionStartId="5.[151,264,1921,1944]" captionTargetBox="[206,1374,772,1904]" captionTargetId="figure@5.[206,1374,772,1904]" captionTargetPageId="5" captionText="FIGURES 7 13. General habitus of Afrotropical Copuetta gen. nov. species: 7. C. erecta sp. nov. male, Chidenguele, Mozambique. 8. C. erecta sp. nov. female, Xai-Xai, Mozambique. 9. C. lacustris (Strand, 1916) female, Bloemfontein, South Africa. 10. C. lotzi sp. nov. male, Bloemfontein. 11. C. magna sp. nov. male, Ndumo Game Reserve, South Africa. 12. C. maputa sp. nov. male, Bilene, Mozambique. 13. C. maputa sp. nov. female, Chidenguele." httpUri="https://zenodo.org/record/284203/files/figure.png" pageId="52" pageNumber="53">Figs 13</figureCitation>
and
<figureCitation id="134929D6FFAEFF86FD37F91736F4F8F9" box="[650,681,1777,1801]" captionStart="FIGURES 45 61" captionStartId="24.[151,264,1801,1824]" captionTargetBox="[151,1436,193,1778]" captionTargetId="figure@24.[151,1436,193,1780]" captionTargetPageId="24" captionText="FIGURES 45 61. Digital microscope photographs of Copuetta kakamega sp. nov. (45), C. kwamgumi sp. nov. (46, 47), C. lacustris (Strand, 1907) (48, 49), C. lesnei sp. nov. (50), C. litipo sp. nov. (51), C. lotzi sp. nov. (52, 53), C. magna sp. nov. (54, 55), C. maputa sp. nov. (56, 57), C. naja sp. nov. (58, 59), C. uzungwa sp. nov. (60) and C. wagneri sp. nov. (61): 45, 47, 49, 51, 53, 55, 57, 59. Female habitus, dorsal view. 46, 48, 50, 52, 54, 56, 58, 60, 61. Male habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284210/files/figure.png" pageId="52" pageNumber="53">57</figureCitation>
, female more robustly built than male. Carapace creamy-yellow, markings and setae as for male, except with black mottling from PER to mediolateral markings. AME separated by distance slightly larger than 15 their diameter; AME nearly touching ALE, separated by distance less than 110 ALE diameter; clypeus height slightly larger than 45 AME diameter; PLE very slightly larger than PME; PME separated by distance slightly less than 35 their diameter; PME separated from PLE by distance slightly larger than ½ PME diameter; CW:
<collectingCountry id="F36575C3FFAEFF86FE81F85035D6F83E" box="[316,395,1974,1998]" name="Peru" pageId="52" pageNumber="53">PERW</collectingCountry>
= 2.92:1. Chelicerae dark brown with dense black mottling, yellow prolaterally distally; promargin with two teeth separated by distance equal to the basal width of proximal tooth, distal tooth much larger; retromargin with two teeth separated by ½ their basal width, distal tooth slightly smaller than proximal tooth, close to fang base. Endites yellow-brown with faint grey mottling, white prolaterally distally; labium pale orange-brown with grey mottling, cream distally; sternum yellow, yellow-brown around borders. Legs creamy-yellow, metatarsi yellow-brown, markings and setae as for male. Leg spination: femora: I pl 2 do 3 rl 1, II pl 2 do 3 rl 1, III pl 2 do 3 rl 2, IV pl 2 do 3 rl 1; all femora with scattered erect ventral setae; patellae: I and II with fine proximal and distal do setae, III and IV with proximal seta and distal spine dorsally; tibiae: I pl 1 plv 2 rlv 2, II pl 1-2 plv 2 rlv 2, III pl 2 do 1 rl 2 plv 2 rlv 2, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I plv 2 rlv 2, II plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 5 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 1 plv 1; tarsi: pl 1 plv 3 rlv 1. Abdomen with pale yellow dorsal scutum, extending to 15 abdomen length; dorsum with black marking anteriorly, laterally and posteriorly, with several small white blotches; dorsum cream along midline with narrow chevrons, broadened in posterior half, with small black spots along midline; venter cream, densely covered in white feathery setae; epigastric scutum creamy-yellow; inframamillary sclerite orange. Epigyne with opposing semi-circular ridges, with copulatory openings situated anteriorly within depression, entering directly into ST II (
<figureCitation id="134929D6FFAFFF87FE6BFD973664FD79" box="[470,569,625,649]" captionStart="FIGURES 141 144" captionStartId="53.[151,264,1734,1757]" captionTargetBox="[165,1424,789,1699]" captionTargetId="figure@53.[151,1436,774,1712]" captionTargetPageId="53" captionText="FIGURES 141 144. Genitalic morphology of Copuetta maputa sp. nov.: 141. Male palp, ventral view. 142. Same, retrolateral view. 143. Female epigyne, ventral view. 144. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284226/files/figure.png" pageId="53" pageNumber="54">Fig. 143</figureCitation>
); copulatory openings often plugged (
<figureCitation id="134929D6FFAFFF87FC58FD973067FD79" box="[997,1082,625,649]" captionStart="FIGURES 92 98" captionStartId="29.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1582]" captionTargetId="figure@29.[151,1436,193,1584]" captionTargetPageId="29" captionText="FIGURES 92 98. Scanning electron microscope photographs of Copuetta gen. nov. genitalia: 92 96. Male palpal emboli of C. erecta sp. nov. (92), C. lacustris (Strand, 1916) (93), C. lotzi sp. nov. (94), C. magna sp. nov. (95) and C. maputa sp. nov. (96). 97, 98. Female epigynes of C. lacustris (97) and C. maputa sp. nov. (98)." httpUri="https://zenodo.org/record/284215/files/figure.png" pageId="53" pageNumber="54">Fig. 98</figureCitation>
); ST II oval with several lobes, connected to kidney-shaped posterior ST I by long narrow ducts; both ST with a few folds on their surface; ST I only half as broad as ST II (
<figureCitation id="134929D6FFAFFF87FE72FD5F366CFD21" box="[463,561,697,721]" captionStart="FIGURES 141 144" captionStartId="53.[151,264,1734,1757]" captionTargetBox="[165,1424,789,1699]" captionTargetId="figure@53.[151,1436,774,1712]" captionTargetPageId="53" captionText="FIGURES 141 144. Genitalic morphology of Copuetta maputa sp. nov.: 141. Male palp, ventral view. 142. Same, retrolateral view. 143. Female epigyne, ventral view. 144. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284226/files/figure.png" pageId="53" pageNumber="54">Fig. 144</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0D65DBFFAFFF87FF2AF9203045F90B" httpUri="https://zenodo.org/record/284226/files/figure.png" pageId="53" pageNumber="54" targetBox="[165,1424,789,1699]" targetPageId="53">
<paragraph id="8BCD3553FFAFFF87FF2AF9203045F90B" blockId="53.[151,1436,1734,1787]" pageId="53" pageNumber="54">
<emphasis id="B906E941FFAFFF87FF2AF9203530F92B" bold="true" box="[151,365,1734,1757]" pageId="53" pageNumber="54">FIGURES 141144.</emphasis>
Genitalic morphology of
<taxonomicName id="4C724ED0FFAFFF87FDC9F921377EF92C" box="[628,803,1735,1756]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="maputa" status="sp. nov.">
<emphasis id="B906E941FFAFFF87FDC9F921377EF92C" box="[628,803,1735,1756]" italics="true" pageId="53" pageNumber="54">Copuetta maputa</emphasis>
</taxonomicName>
<emphasis id="B906E941FFAFFF87FC94F9213724F92C" bold="true" box="[809,889,1735,1756]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A235543AFFAFFF87FC94F9213724F92C" box="[809,889,1735,1756]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 141. Male palp, ventral view. 142. Same, retrolateral view. 143. Female epigyne, ventral view. 144. Same, dorsal view. Scale bars = 0.25 mm.
</paragraph>
</caption>
<subSubSection id="C36866D8FFAFFF84FF7AF8C3350EFCE4" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BCD3553FFAFFF87FF7AF8C330BEF891" blockId="53.[151,1436,1829,2033]" pageId="53" pageNumber="54">
<emphasis id="B906E941FFAFFF87FF7AF8C33527F8CE" bold="true" box="[199,378,1829,1854]" pageId="53" pageNumber="54">
<typeStatus id="54C98BF1FFAFFF87FF7AF8C3355FF8CD" box="[199,258,1829,1853]" pageId="53" pageNumber="54">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54C98BF1FFAFFF87FE39F8C335B1F8CD" box="[388,492,1829,1853]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
3:
<emphasis id="B906E941FFAFFF87FDA0F8C3375DF8CE" bold="true" box="[541,768,1829,1854]" pageId="53" pageNumber="54">
<collectingCountry id="F36575C3FFAFFF87FDA0F8C336A6F8CE" box="[541,763,1829,1854]" name="South Africa" pageId="53" pageNumber="54">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="B906E941FFAFFF87FCB7F8C337E5F8CD" box="[778,952,1829,1853]" italics="true" pageId="53" pageNumber="54">KwaZulu-Natal</emphasis>
: Ndumo Game Reserve, Crocodile Farm,
<geoCoordinate id="EE465394FFAFFF87FF2AF8AF357FF891" box="[151,290,1865,1889]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-26.9071">26°54.426'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFAFFF87FE90F8AF35E7F891" box="[301,442,1865,1889]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="32.31975">32°19.185'E</geoCoordinate>
, leg. C. Haddad,
<date id="FFCC1393FFAFFF87FDC1F8AF36BAF891" box="[636,743,1865,1889]" pageId="53" pageNumber="54" value="2006-01-17">17.I.2006</date>
(on wall of house at night) (NCA 2006/425).
</paragraph>
<paragraph id="8BCD3553FFAFFF87FF7AF88831C1F801" blockId="53.[151,1436,1829,2033]" pageId="53" pageNumber="54">
<typeStatus id="54C98BF1FFAFFF87FF7AF8883565F876" box="[199,312,1902,1926]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B906E941FFAFFF87FEF9F88B3644F876" bold="true" box="[324,537,1901,1926]" pageId="53" pageNumber="54">
<collectingCountry id="F36575C3FFAFFF87FEF9F88B3649F876" box="[324,532,1901,1926]" name="Mozambique" pageId="53" pageNumber="54">MOZAMBIQUE</collectingCountry>
:
</emphasis>
Bilene, Praia do Bilene,
<geoCoordinate id="EE465394FFAFFF87FC87F88B3799F875" box="[826,964,1901,1925]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-25.260817">25°15.649'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFAFFF87FC6FF88B3002F875" box="[978,1119,1901,1925]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="33.294315">33°17.659'E</geoCoordinate>
,
<quantity id="4C8A98B6FFAFFF87FBD1F88B30C0F875" box="[1132,1181,1901,1925]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="53" pageNumber="54" unit="m" value="27.0">27m</quantity>
a.s.l., leg. C. Haddad, R. Lyle &amp; R. Fourie,
<date id="FFCC1393FFAFFF87FE1FF8773676F859" box="[418,555,1937,1961]" pageId="53" pageNumber="54" value="2007-12-20">20.XII.2007</date>
(leaf litter, coastal forest), 13 (NCA 2008/209); Chidenguele, Paraiso de Chidenguele,
<geoCoordinate id="EE465394FFAFFF87FE8AF853359EF83D" box="[311,451,1973,1997]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-24.9546">24°57.276'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFAFFF87FE6EF8533602F83D" box="[467,607,1973,1997]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="34.197666">34°11.860'E</geoCoordinate>
,
<quantity id="4C8A98B6FFAFFF87FDCDF85336FFF83D" box="[624,674,1973,1997]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="53" pageNumber="54" unit="m" value="38.0">38m</quantity>
a.s.l., leg. C. Haddad, R. Lyle &amp; R. Fourie,
<date id="FFCC1393FFAFFF87FB79F8533111F83D" box="[1220,1356,1973,1997]" pageId="53" pageNumber="54" value="2007-12-16">16.XII.2007</date>
(night collecting, dune forest), 1Ƥ (NCA 2008/200).
<emphasis id="B906E941FFAFFF87FD7CF83E37F8F801" bold="true" box="[705,933,2008,2033]" pageId="53" pageNumber="54">
<collectingCountry id="F36575C3FFAFFF87FD7CF83E37FDF801" box="[705,928,2008,2033]" name="South Africa" pageId="53" pageNumber="54">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="B906E941FFAFFF87FC0FF83F303DF801" box="[946,1120,2009,2033]" italics="true" pageId="53" pageNumber="54">KwaZulu-Natal</emphasis>
: iSimangaliso [Greater St.
</paragraph>
<paragraph id="8BCD3553FFACFF84FF2AFF713736FE3F" blockId="54.[151,1436,151,932]" pageId="54" pageNumber="55">
Lucia] Wetlands Park, False Bay Park [
<geoCoordinate id="EE465394FFACFF84FDF0FF7136F4FF5F" box="[589,681,151,175]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-27.916666">27°55'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FD09FF71374FFF5F" box="[692,786,151,175]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="32.266666">32°16'E</geoCoordinate>
], leg. J. Esterhuizen,
<date id="FFCC1393FFACFF84FC42FF713038FF5F" box="[1023,1125,151,175]" pageId="54" pageNumber="55" value="2004-02-04">4.II.2004</date>
(tsetse fly traps), 13 (NCA 2004/761); iSimangaliso Wetlands Park, Hells Gate [
<geoCoordinate id="EE465394FFACFF84FD4EFF5A3713FF24" box="[755,846,188,212]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-28.0">28°00'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FCE1FF5A37E4FF24" box="[860,953,188,212]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="32.483334">32°29'E</geoCoordinate>
], leg. J. Esterhuizen,
<date id="FFCC1393FFACFF84FB16FF5A317BFF24" box="[1195,1318,188,212]" pageId="54" pageNumber="55" value="2003-12-08">8.XII.2003</date>
(tsetse fly traps), 13 (NCA 2004/785); same data,
<date id="FFCC1393FFACFF84FDE1FF3936B5FF07" box="[604,744,223,247]" pageId="54" pageNumber="55" value="2003-12-14">14.XII.2003</date>
, 13 (NCA 2004/799); same data,
<date id="FFCC1393FFACFF84FBD1FF393084FF07" box="[1132,1241,223,247]" pageId="54" pageNumber="55" value="2004-01-19">19.I.2004</date>
, 33 (NCA 2004/ 812); same data,
<date id="FFCC1393FFACFF84FEDFFEE23596FEEC" box="[354,459,260,284]" pageId="54" pageNumber="55" value="2004-02-09">9.II.2004</date>
, 13 (NCA 2004/804); Lake Sibaya,
<geoCoordinate id="EE465394FFACFF84FCC2FEE23784FEEC" box="[895,985,260,284]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-27.333334">27°20'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FC54FEE23018FEEC" box="[1001,1093,260,284]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="32.7">32°42'E</geoCoordinate>
, leg. B. Lamoral,
<date id="FFCC1393FFACFF84FAA3FEE231C1FEEB" box="[1310,1436,259,284]" pageId="54" pageNumber="55" value="1968-01-13">I.1968, 13</date>
(
<collectionCode id="ED63AD96FFACFF84FF23FEC134ACFECF" LSID="urn:lsid:biocol.org:col:34985" box="[158,241,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="54" pageNumber="55">NMSA</collectionCode>
12337); Ndumo Game Reserve, Southern shore of Hotwe Pan,
<geoCoordinate id="EE465394FFACFF84FC00FEC13015FECF" box="[957,1096,295,319]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-26.878834">26°52.730'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FBE9FEC130BDFECF" box="[1108,1248,295,319]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="32.307533">32°18.452'E</geoCoordinate>
, leg. C. Haddad,
<date id="FFCC1393FFACFF84FF2AFEAA34A3FE94" box="[151,254,332,356]" pageId="54" pageNumber="55" value="2005-02-07">7.II.2005</date>
(
<taxonomicName id="4C724ED0FFACFF84FEABFEAA3651FE94" box="[278,524,332,356]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B906E941FFACFF84FEABFEAA3651FE94" box="[278,524,332,356]" italics="true" pageId="54" pageNumber="55">Acacia xanthophloea</emphasis>
</taxonomicName>
bark), 1Ƥ (NCA 2008/2910); same locality, Crocodile Farm,
<geoCoordinate id="EE465394FFACFF84FAB1FEAA31C5FE94" box="[1292,1432,332,356]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-26.9071">26°54.426'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FF2AFE89357EFE77" box="[151,291,367,391]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="32.31975">32°19.185'E</geoCoordinate>
, leg. C. Haddad,
<date id="FFCC1393FFACFF84FE50FE893605FE77" box="[493,600,367,391]" pageId="54" pageNumber="55" value="2006-01-17">17.I.2006</date>
(on wall of house at night), 13 (NCA 2008/271); Sodwana Bay to Lake Sibaya,
<geoCoordinate id="EE465394FFACFF84FF48FE723521FE5C" box="[245,380,404,428]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-27.41563">27.41563°S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FE36FE72364FFE5C" box="[395,530,404,428]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="32.70982">32.70982°E</geoCoordinate>
, leg. P. van Niekerk, G. van Niekerk, X. Combrinck &amp; J. Warner,
<date id="FFCC1393FFACFF84FA9AFE7231C1FE5C" box="[1319,1436,404,428]" pageId="54" pageNumber="55" value="2007-02-28">28.II.2007</date>
(beats, undergrowth, coastal dune forest), 13 (NCA 2007/3829).
</paragraph>
<paragraph id="8BCD3553FFACFF84FF7AFE3D350EFCE4" blockId="54.[151,1436,151,932]" pageId="54" pageNumber="55">
<emphasis id="B906E941FFACFF84FF7AFE3D371DFE04" bold="true" box="[199,832,475,500]" pageId="54" pageNumber="55">
Additional material examined:
<collectingCountry id="F36575C3FFACFF84FDEAFE3D3767FE04" box="[599,826,475,500]" name="South Africa" pageId="54" pageNumber="55">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="B906E941FFACFF84FCEDFE3A37A3FE04" box="[848,1022,476,500]" italics="true" pageId="54" pageNumber="55">KwaZulu-Natal</emphasis>
: iSimangaliso [Greater
<collectingCountry id="F36575C3FFACFF84FA98FE3A31CBFE04" box="[1317,1430,476,500]" name="Saint Lucia" pageId="54" pageNumber="55">St. Lucia</collectingCountry>
] Wetlands Park, False Bay Park [
<geoCoordinate id="EE465394FFACFF84FDACFE193630FDE7" box="[529,621,511,535]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-27.916666">27°55'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FDC1FE193687FDE7" box="[636,730,511,535]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="32.266666">32°16'E</geoCoordinate>
], leg. J. Esterhuizen,
<date id="FFCC1393FFACFF84FC6BFE19300CFDE7" box="[982,1105,511,535]" pageId="54" pageNumber="55" value="2004-12-01">1.XII.2004</date>
(tsetse fly traps), 13 (NCA 2005/196); iSimangaliso Wetlands Park, Hells Gate [
<geoCoordinate id="EE465394FFACFF84FD4BFDC2370CFDCC" box="[758,849,548,572]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-28.0">28°00'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FCDDFDC237E0FDCC" box="[864,957,548,572]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="32.483334">32°29'E</geoCoordinate>
], Block A, leg. J. Esterhuizen,
<date id="FFCC1393FFACFF84FA9DFDC231C1FDCC" box="[1312,1436,548,572]" pageId="54" pageNumber="55" value="2004-04-26">26.IV.2004</date>
(tsetse fly traps), 13 (NCA 2005/192); same data,
<date id="FFCC1393FFACFF84FCADFDA137CEFDAF" box="[784,915,583,607]" pageId="54" pageNumber="55" value="2004-06-14">14.VI.2004</date>
, 1 imm. 1Ƥ (NCA 2005/332); same data,
<date id="FFCC1393FFACFF84FF2AFD8A3544FD74" box="[151,281,620,644]" pageId="54" pageNumber="55" value="2004-11-29">29.XI.2004</date>
, 13 (NCA 2005/195); same data,
<date id="FFCC1393FFACFF84FD29FD8A374FFD74" box="[660,786,620,644]" pageId="54" pageNumber="55" value="2004-12-06">6.XII.2004</date>
, 13 (NCA 2005/197); iSimangaliso Wetlands Park, Hells Gate [
<geoCoordinate id="EE465394FFACFF84FF61FD693565FD57" box="[220,312,655,679]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-28.0">28°00'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFACFF84FEFBFD6935FEFD57" box="[326,419,655,679]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="32.483334">32°29'E</geoCoordinate>
], Block B, leg. J. Esterhuizen,
<date id="FFCC1393FFACFF84FCB4FD693727FD57" box="[777,890,655,679]" pageId="54" pageNumber="55" value="2004-06-07">7.VI.2004</date>
(tsetse fly traps), 1 imm. 1Ƥ (NCA 2005/194); same data,
<date id="FFCC1393FFACFF84FE9AFD5235E1FD3C" box="[295,444,692,716]" pageId="54" pageNumber="55" value="2004-08-16">16.VIII.2004</date>
, 1Ƥ (NCA 2005/334); same data,
<date id="FFCC1393FFACFF84FCC8FD5237AEFD3C" box="[885,1011,692,716]" pageId="54" pageNumber="55" value="2004-12-05">5.XII.2004</date>
, 1Ƥ (NCA 2005/336); same data,
<date id="FFCC1393FFACFF84FF2AFD313549FD1F" box="[151,276,727,751]" pageId="54" pageNumber="55" value="2004-12-06">6.XII.2004</date>
, 13 (NCA 2005/198);
<collectingCountry id="F36575C3FFACFF84FD8CFD31362DFD1F" box="[561,624,727,751]" name="South Africa" pageId="54" pageNumber="55">Natal</collectingCountry>
, leg. C. Martin, misidentified 3
<typeStatus id="54C98BF1FFACFF84FBB9FD3130C6FD1F" box="[1028,1179,727,751]" pageId="54" pageNumber="55" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C724ED0FFACFF84FB70FD3F31C1FD00" box="[1229,1436,729,752]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="B906E941FFACFF84FB70FD3F31C1FD00" box="[1229,1436,729,752]" italics="true" pageId="54" pageNumber="55">Messapus martini</emphasis>
</taxonomicName>
(
<collectionCode id="ED63AD96FFACFF84FF23FD1A34ABFCE4" LSID="urn:lsid:biocol.org:col:34988" box="[158,246,764,788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
19680).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFACFF84FF7AFCF93690FCAC" pageId="54" pageNumber="55" type="distribution">
<paragraph id="8BCD3553FFACFF84FF7AFCF93690FCAC" blockId="54.[151,1436,151,932]" pageId="54" pageNumber="55">
<emphasis id="B906E941FFACFF84FF7AFCF9353FFCC8" bold="true" box="[199,354,799,824]" pageId="54" pageNumber="55">Distribution:</emphasis>
Apparently endemic to the coastal regions and adjacent interior of southern
<collectingCountry id="F36575C3FFACFF84FB6AFCF93136FCC7" box="[1239,1387,799,823]" name="Mozambique" pageId="54" pageNumber="55">Mozambique</collectingCountry>
and KwaZulu-Natal Province,
<collectingCountry id="F36575C3FFACFF84FE02FCA23613FCAC" box="[447,590,836,860]" name="South Africa" pageId="54" pageNumber="55">South Africa</collectingCountry>
(
<figureCitation id="134929D6FFACFF84FDE3FCA2369CFCAC" box="[606,705,836,860]" captionStart="FIGURE 153" captionStartId="59.[151,250,1619,1642]" captionTargetBox="[151,1426,339,1576]" captionTargetId="figure@59.[151,1436,339,1596]" captionTargetPageId="59" captionText="FIGURE 153. Distribution of Copuetta lotzi sp. nov. (black circles), C. magna sp. nov. (white circles), C. maputa sp. nov. (black triangles), C. naja sp. nov. (white triangle), C. uzungwa sp. nov. (black star) and C. wagneri sp. nov. (white star) in the Afrotropical Region." httpUri="https://zenodo.org/record/284229/files/figure.png" pageId="54" pageNumber="55">Fig. 153</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFACFF84FF7AFC813560FC54" pageId="54" pageNumber="55" type="biology_ecology">
<paragraph id="8BCD3553FFACFF84FF7AFC813560FC54" blockId="54.[151,1436,151,932]" pageId="54" pageNumber="55">
<emphasis id="B906E941FFACFF84FF7AFC813577FC70" bold="true" box="[199,298,871,896]" pageId="54" pageNumber="55">Biology:</emphasis>
A species primarily collected in the lower strata of forest and woodland habitats (shrubs, tree bark and leaf litter).
</paragraph>
</subSubSection>
</treatment>
</document>