353 lines
42 KiB
XML
353 lines
42 KiB
XML
<document id="BF3EEAAC10536C05C1423DE46D29FE71" ID-DOI="10.5281/zenodo.178134" ID-GBIF-Dataset="7aafe288-b0d6-42ab-9d18-50b06b21ca81" ID-ISSN="1175-5326" ID-Zenodo-Dep="178134" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459882195660" checkinUser="plazi" docAuthor="Walsh, Jonathan H. & Ebert, David A." docDate="2007" docId="03DA382CFFC6883676AAC857FC38F900" docLanguage="en" docName="zt01551p047.pdf" docOrigin="Zootaxa 1551" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Squatina formosa Shen & Ting 1972" docType="treatment" docVersion="8" lastPageNumber="38" masterDocId="FFE34054FFC28831763DCD4AFFCDFFC3" masterDocTitle="A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae)" masterLastPageNumber="47" masterPageNumber="31" pageNumber="35" updateTime="1698227577416" updateUser="plazi">
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<mods:title id="4273D1906C9CEB327353D88D689FF9C4">A review of the systematics of western North Pacific angel sharks, genus Squatina, with redescriptions of Squatina formosa, S. japonica, and S. nebulosa (Chondrichthyes: Squatiniformes, Squatinidae)</mods:title>
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<mods:namePart id="E2673FC3AC8772B612EB6569F3CB1ACE">Walsh, Jonathan H.</mods:namePart>
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<mods:namePart id="D7693ECA124ED0F9E30C6252A89C188D">Ebert, David A.</mods:namePart>
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<mods:typeOfResource id="50C930C63A9F487681BC5750BC389C99">text</mods:typeOfResource>
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<mods:title id="839DCB8A14053D8D578D49B7CF03D322">Zootaxa</mods:title>
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<mods:part id="FA62FFD35685C803683CC17C866056ED">
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<mods:date id="A42BE66FC0B904175743E4F09597C808">2007</mods:date>
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<mods:detail id="6D555027A572E6348FA6E4ACF595BF0D" type="volume">
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<mods:number id="BBDCA6D5DBA1FEC3E2DEAEAB651101DA">1551</mods:number>
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<mods:start id="33BE31B4DFCFD7A7189AB8245A9B016D">31</mods:start>
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<mods:end id="43193D23E9B014A3B5F7D56CD587214A">47</mods:end>
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<mods:identifier id="06E0BE6357140EC65B188288F9F0647E" type="DOI">10.5281/zenodo.178134</mods:identifier>
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<mods:identifier id="43F4E9B9FB06C8F9721CF8D0E37C0771" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0518F1501AB2FF72C1E3B4CD4DBDF2EE" type="Zenodo-Dep">178134</mods:identifier>
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<treatment id="03DA382CFFC6883676AAC857FC38F900" ID-DOI="http://doi.org/10.5281/zenodo.6240324" ID-GBIF-Taxon="119370972" ID-Zenodo-Dep="6240324" LSID="urn:lsid:plazi:treatment:03DA382CFFC6883676AAC857FC38F900" httpUri="http://treatment.plazi.org/id/03DA382CFFC6883676AAC857FC38F900" lastPageId="7" lastPageNumber="38" pageId="4" pageNumber="35">
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<subSubSection id="C369DAB1FFC6883576AAC857FDBFFAF4" box="[151,626,1309,1335]" pageId="4" pageNumber="35" type="nomenclature">
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<paragraph id="8BCC893AFFC6883576AAC857FDBFFAF4" blockId="4.[151,626,1309,1335]" box="[151,626,1309,1335]" pageId="4" pageNumber="35">
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<heading id="D0843E56FFC6883576AAC857FDBFFAF4" bold="true" box="[151,626,1309,1335]" fontSize="11" level="1" pageId="4" pageNumber="35" reason="1">
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<taxonomicName id="4C73F2B9FFC6883576AAC857FDBFFAF4" ID-CoL="6ZFHJ" authority="Shen & Ting, 1972" authorityName="Shen & Ting" authorityYear="1972" box="[151,626,1309,1335]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="4" pageNumber="35" phylum="Chordata" rank="species" species="formosa">
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<emphasis id="B9075528FFC6883576AAC857FDBFFAF4" bold="true" box="[151,626,1309,1335]" pageId="4" pageNumber="35">
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<emphasis id="B9075528FFC6883576AAC857FEA3FAF4" bold="true" box="[151,366,1309,1335]" italics="true" pageId="4" pageNumber="35">Squatina formosa</emphasis>
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(
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<bibRefCitation id="EFE2F4CBFFC688357743C857FDA4FAF4" author="Shen" box="[382,617,1309,1335]" pageId="4" pageNumber="47" refString="Shen S-C., & Ting, W-H. (1972) Ecological and morphological study on fish fauna from the waters around Taiwan and its adjacent islands. Notes on some continental shelf fishes and descriptions of two new species. Bulletin of the Institute of Zoologic Academy - Sinica, 11, 13 - 31." type="journal article" year="1972">Shen & Ting, 1972</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DAB1FFC6883576AAC828FD9DFA07" pageId="4" pageNumber="35" type="reference_group">
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<paragraph id="8BCC893AFFC6883576AAC828FBA2FABA" blockId="4.[151,1135,1378,1401]" box="[151,1135,1378,1401]" pageId="4" pageNumber="35">
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<treatmentCitationGroup id="AB63AE14FFC6883576AAC828FBA2FABA" box="[151,1135,1378,1401]" pageId="4" pageNumber="35">
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<taxonomicName id="4C73F2B9FFC6883576AAC828FD9CFABA" ID-CoL="6ZFHJ" authority="Shen & Ting 1972: 23" authorityName="Shen & Ting" authorityPageNumber="23" authorityYear="1972" box="[151,593,1378,1401]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="4" pageNumber="35" phylum="Chordata" rank="species" species="formosa">
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<emphasis id="B9075528FFC6883576AAC828FE9EFABA" box="[151,339,1378,1401]" italics="true" pageId="4" pageNumber="35">Squatina formosa</emphasis>
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,
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<treatmentCitation id="0AD2AF2BFFC68835775CC828FD9CFABA" author="Shen" box="[353,593,1378,1401]" page="23" pageId="4" pageNumber="35" year="1972">
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<bibRefCitation id="EFE2F4CBFFC68835775CC828FDE4FABA" author="Shen" box="[353,553,1378,1401]" pageId="4" pageNumber="47" refString="Shen S-C., & Ting, W-H. (1972) Ecological and morphological study on fish fauna from the waters around Taiwan and its adjacent islands. Notes on some continental shelf fishes and descriptions of two new species. Bulletin of the Institute of Zoologic Academy - Sinica, 11, 13 - 31." type="journal article" year="1972">Shen & Ting 1972</bibRefCitation>
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: 23
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</treatmentCitation>
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</taxonomicName>
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,
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<figureCitation id="134895BFFFC688357461C828FD7AFABA" box="[604,695,1378,1401]" captionStart="FIGURE 4" captionStartId="9.[151,255,1978,2002]" captionTargetBox="[257,1332,576,1934]" captionTargetId="figure@9.[235,1357,543,1959]" captionTargetPageId="9" captionText="FIGURE 4: A. Squatina japonica (HUMZ 105913), immature male, 441 mm, B. Close-up of snout." httpUri="https://zenodo.org/record/178137/files/figure.png" pageId="4" pageNumber="35">Figure 4</figureCitation>
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, valid,
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<typeStatus id="54C83798FFC688357538C828FCAAFABA" box="[773,871,1378,1401]" pageId="4" pageNumber="35" type="holotype">holotype</typeStatus>
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: NTT7213130.
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<figureCitation id="134895BFFFC688357232C828FBA6FABA" box="[1039,1131,1378,1401]" captionStart="FIGURE 2" captionStartId="6.[151,255,1977,2001]" captionTargetBox="[213,1375,309,1976]" captionTargetId="figure@6.[213,1397,287,1987]" captionTargetPageId="6" captionText="FIGURE 2: A. S. formosa holotype (NTUM 01329), immature female, 328 mm, B. Close-up of caudal fin, C. Close-up of snout." httpUri="https://zenodo.org/record/178135/files/figure.png" pageId="4" pageNumber="35">Figure 2</figureCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCC893AFFC6883576AAC8E0FD9DFA07" blockId="4.[151,1436,1450,2036]" box="[151,592,1450,1476]" pageId="4" pageNumber="35">
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<emphasis id="B9075528FFC6883576AAC8E0FE90FA07" bold="true" box="[151,349,1450,1476]" pageId="4" pageNumber="35">Common name.</emphasis>
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<collectingCountry id="F364C9AAFFC688357751C8E0FE0FFA07" box="[364,450,1450,1476]" name="Taiwan" pageId="4" pageNumber="35">Taiwan</collectingCountry>
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angelshark.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DAB1FFC6883576FBC898FB17F977" pageId="4" pageNumber="35" type="diagnosis">
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<paragraph id="8BCC893AFFC6883576FBC898FB17F977" blockId="4.[151,1436,1450,2036]" pageId="4" pageNumber="35">
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<emphasis id="B9075528FFC6883576FBC898FE8BFA2F" bold="true" box="[198,326,1490,1516]" pageId="4" pageNumber="35">Diagnosis.</emphasis>
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A squatinid with the following distinctive characters: upper lip arch semi-circular, height greater than other WNP squatinids (3.8–5.1% in width, 1.4–2.1% TL in height); pectoral fins broadly rounded, especially posterior free tip; pelvic girdle moderately broad, pelvic anterior margin slightly curved, angle of lateral apex considerably more obtuse than 120°, pelvic fin tips reaching first dorsal origin; dorsal fins lobe-like with slightly rounded anterior margin, first dorsal fin base slightly larger than second dorsal base; caudal fin lobed, especially dorsally, with a curvilinear caudal posterior ventral margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DAB1FFC6883476FBCB88FE4AF94D" lastPageId="5" lastPageNumber="36" pageId="4" pageNumber="35" type="description">
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<paragraph id="8BCC893AFFC6883576FBCB88FEE3F8EF" blockId="4.[151,1436,1450,2036]" pageId="4" pageNumber="35">
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<emphasis id="B9075528FFC6883576FBCB88FE9AF91F" bold="true" box="[198,343,1730,1756]" pageId="4" pageNumber="35">Description</emphasis>
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. Dorsal surface, except for posterior portion of caudal fin, covered with denticles of moderate roughness. Ventral surface smooth except for narrow bands of denticles on the pectoral and pelvic fins anterior margins.
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</paragraph>
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<paragraph id="8BCC893AFFC6883476FBCA70FA56FE61" blockId="4.[151,1436,1450,2036]" lastBlockId="5.[151,1437,152,2038]" lastPageId="5" lastPageNumber="36" pageId="4" pageNumber="35">Head rounded, length about 0.2 times total length, maximum width occurring just anterior of gill openings. Moderately rough tubercles interspersed above mouth and eye crests. Eyes almond-shaped, widely set, interorbital space 8.2 (7.7–8.9). Eye-spiracle space short. Spiracles are crescent shaped without large papillae. Interspiracle space (7.8–8.2) slightly less than interorbital space. Center of upper lip exposed at midpoint of upper jaw, exposure semi-circular in shape, extending dorsally approximately 0.6–0.7 of upper jaw space, upper lip height (1.2–2.1), upper lip arch width (3.8–5.1). Labial furrows conspicuous, roughly equal in length, extending from corners of mouth medially, with upper labial furrow partially covered with dermal folds. Distinct nasal flaps protruding from dermal folds above mouth, two barbels protruding from each flap. Inner nasal barbel rod-like with a spatulate tip, inner basal portion contains little if any fringe. Outer nasal barbel narrow. Nostrils large, slightly protruding. Dermal folds along exterior of head, one small lobe present at corners of mouth extending ventrally. Mouth length about 0.3 times as long as mouth width. Dentition consisting of small, dagger-like teeth, conical without cusplets on a broad base, in 3 orderly longitudinal rows, no</paragraph>
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<paragraph id="8BCC893AFFC788347424CCF8FD7BFE07" blockId="5.[151,1437,152,2038]" box="[537,694,434,454]" pageId="5" pageNumber="36">
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<date id="FFCDAFFAFFC788347424CCF8FD7BFE07" box="[537,694,434,454]" pageId="5" pageNumber="36" value="1910-09-10" valueMax="1910-09-09" valueMin="1910-09-10">9 − 10 − 9 − 10</date>
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</paragraph>
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<paragraph id="8BCC893AFFC7883476AACC8AFD02FE19" blockId="5.[151,1437,152,2038]" box="[151,719,448,483]" pageId="5" pageNumber="36">teeth at symphysis, teeth by row 10 − 10.</paragraph>
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<paragraph id="8BCC893AFFC7883476FBCCA6FB4FFDBD" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">Pectoral fins large, broadly rounded, originating just behind gills. Anterior margin of pectoral fin slightly convex and about three quarters as long as pectoral length, extending to a lateral apex. Angle of lateral apex slightly more obtuse than 120°. Margin from lateral apex to most posterior lobe slightly concave. Posterior lobe broadly convex. Pectoral inner margin convex, about one half of pectoral length.</paragraph>
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<paragraph id="8BCC893AFFC7883476FBCFC6FD98FC25" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">Overall pelvic fin shape somewhat triangular with rounded fintips. Pelvics originating anterior to pectoral fin free rear tips. Pelvic fin length approximately two thirds as long as pectoral fin length. Pelvic fin base approximately 1.2 times broader than pectoral base. Anterior margin slightly curvilinear, extending at roughly a 45° angle from trunk to rounded apex lateral of body, anterior margin 0.5 times as long as pelvic fin length. Pelvic girdle width (26.3–29.8) between pelvic fin apexes moderately broad, about 1.4 times head length. Posterior margin of pelvic fin, approximately 0.8 times pelvic fin length, straight to posterior free tip. Pelvic inner margin concave and short, only about 0.4 times as long as pectoral fin length. Pelvic insertion furrows on ventral extend in a narrow ellipse to anterior apogee of vent in most specimens, vent is within ellipse. Pelvic fin tips reach origin of first dorsal.</paragraph>
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<paragraph id="8BCC893AFFC7883476FBCEBEFAEEFB6D" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">Dorsal fins lobed and nearly equal in size, with denticles covering the whole of fins. Interdorsal space about 0.8 times as long as dorsal caudal finspace. Anterior margin of dorsals slightly convex, nearly equidistant. First dorsal base slightly shorter than second, first dorsal base 15.2 (10.0–11.8), second dorsal base 17.4 (13.0–15.0). Apex of dorsals lobed. Posterior margins slightly convex, about 0.7 times as long as anterior margins. Inner margins of dorsals slightly convex, approximately 0.7 times as long as anterior margins.</paragraph>
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<paragraph id="8BCC893AFFC7883476FBC9F6FE87FAB5" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">Caudal peduncle compressed dorso-ventrally with lateral longitudinal ridges, tapering posteriorly. Caudal fin lobe-like, markedly at dorsal apex. Caudal dorsal margin broadly rounded, about 0.8 times as long as preventral caudal fin margin. Subterminal caudal fin margin approximately half as long as caudal upper post ventral margin. Caudal lower postventral margin convex, approximately 0.8 times as long as caudal upper post ventral margin.</paragraph>
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<paragraph id="8BCC893AFFC7883476FBC8CEFCC8FA05" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">Total vertebrae 137–139; total precaudal vertebrae 107-110; monospondylous vertebrae 48–52; diplospondylous vertebrae 58–59; caudal vertebrae 29-30.</paragraph>
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<paragraph id="8BCC893AFFC7883476FBC89EFE4AF94D" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">
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<emphasis id="B9075528FFC7883476FBC89EFE80FA2D" bold="true" box="[198,333,1492,1518]" pageId="5" pageNumber="36">Coloration</emphasis>
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. Dorsal surface of specimens prior to preservation are light to dark brown throughout with numerous black and white spots of varying sizes. Black blotches laterally at origin of dorsals. Ventral surface pale white with some black mottling on abdomen, pectoral and pelvic fin ventral margins with denticles colored similar to dorsal. Color after preservation tends to fade to a lighter brown or pale yellow with spots becoming indistinct.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DAB1FFC7883476FBCBD6FBECF91D" pageId="5" pageNumber="36" type="distribution">
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<paragraph id="8BCC893AFFC7883476FBCBD6FBECF91D" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">
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<emphasis id="B9075528FFC7883476FBCBD6FE92F975" bold="true" box="[198,351,1692,1718]" pageId="5" pageNumber="36">Distribution</emphasis>
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. Endemic to western North Pacific including the East
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<collectingCountry id="F364C9AAFFC7883475D7CBD6FBFDF975" box="[1002,1072,1692,1718]" name="China" pageId="5" pageNumber="36">China</collectingCountry>
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Sea (
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<bibRefCitation id="EFE2F4CBFFC788347249CBD6FA43F975" author="Compagno" box="[1140,1422,1692,1718]" pageId="5" pageNumber="47" refString="Compagno, L. J. V., Dando, M & Fowler, S. (2005 a) Sharks of the world. Princeton University Press, Princeton and Oxford, 368 pp." type="book" year="2005" yearSuffix="a">Compagno et al, 2005a</bibRefCitation>
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), waters surrounding northern
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<collectingCountry id="F364C9AAFFC7883477D7CB8EFD8EF91D" box="[490,579,1732,1758]" name="Taiwan" pageId="5" pageNumber="36">Taiwan</collectingCountry>
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, and East
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<collectingCountry id="F364C9AAFFC788347487CB8EFCDDF91D" box="[698,784,1732,1758]" name="Taiwan" pageId="5" pageNumber="36">Taiwan</collectingCountry>
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Strait (
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<bibRefCitation id="EFE2F4CBFFC788347558CB8EFBD9F91D" author="Shuyuan" box="[869,1044,1732,1758]" pageId="5" pageNumber="47" refString="Shuyuan, C. (1994) On the chondrichthyian fauna of the Taiwan Strait. Fourth Indo-Pacific Fish Conference, Bangkok Thailand, 28 Nov- 4 Dec 1993, 127 - 137." type="journal article" year="1994">Shuyuan 1994</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DAB1FFC7883476FBCBA6FEBAF8ED" pageId="5" pageNumber="36" type="etymology">
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<paragraph id="8BCC893AFFC7883476FBCBA6FEBAF8ED" blockId="5.[151,1437,152,2038]" pageId="5" pageNumber="36">
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<emphasis id="B9075528FFC7883476FBCBA6FE86F8C5" bold="true" box="[198,331,1772,1798]" pageId="5" pageNumber="36">Etymology</emphasis>
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. Named in allusion to the known geographic range (
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<taxonomicName id="4C73F2B9FFC788347586CBA6FBE9F8C5" box="[955,1060,1772,1798]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="formosa">
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<collectingCountry id="F364C9AAFFC788347586CBA6FBE9F8C5" box="[955,1060,1772,1798]" name="Taiwan" pageId="5" pageNumber="36">Formosa</collectingCountry>
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</taxonomicName>
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Strait,
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<collectingCountry id="F364C9AAFFC788347247CBA6FB1FF8C5" box="[1146,1234,1772,1798]" name="Taiwan" pageId="5" pageNumber="36">Taiwan</collectingCountry>
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) where the
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<typeStatus id="54C83798FFC788347363CBA6FF07F8ED" pageId="5" pageNumber="36" type="holotype">holotype</typeStatus>
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was collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DAB1FFC7883676FBCA76FC7CFE38" lastPageId="7" lastPageNumber="38" pageId="5" pageNumber="36" type="discussion">
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<paragraph id="8BCC893AFFC7883776FBCA76FEEEFEC1" blockId="5.[151,1437,152,2038]" lastBlockId="6.[151,1436,152,258]" lastPageId="6" lastPageNumber="37" pageId="5" pageNumber="36">
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<emphasis id="B9075528FFC7883476FBCA76FEF5F895" bold="true" box="[198,312,1852,1878]" pageId="5" pageNumber="36">Remarks</emphasis>
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. Examination of the
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<typeStatus id="54C83798FFC788347412CA76FD58F895" box="[559,661,1852,1878]" pageId="5" pageNumber="36" type="holotype">holotype</typeStatus>
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(labeled NTT
|
||
<quantity id="4C8B24DFFFC788347506CA76FC0DF895" box="[827,960,1852,1878]" metricMagnitude="5" metricUnit="m" metricValue="1.83213502" pageId="5" pageNumber="36" unit="in" value="7213130.0">7213130 in</quantity>
|
||
<bibRefCitation id="EFE2F4CBFFC7883475F5CA76FB6EF895" author="Shen" box="[968,1187,1852,1878]" pageId="5" pageNumber="47" refString="Shen S-C., & Ting, W-H. (1972) Ecological and morphological study on fish fauna from the waters around Taiwan and its adjacent islands. Notes on some continental shelf fishes and descriptions of two new species. Bulletin of the Institute of Zoologic Academy - Sinica, 11, 13 - 31." type="journal article" year="1972">Shen & Ting 1972</bibRefCitation>
|
||
, now labeled NTUM 01329) and non-type comparison material revealed that
|
||
<taxonomicName id="4C73F2B9FFC78834757FCA2EFC0EF8BE" box="[834,963,1892,1917]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC78834757FCA2EFC0EF8BE" box="[834,963,1892,1917]" italics="true" pageId="5" pageNumber="36">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
is distinct from other WNP squatinids through four characters (
|
||
<figureCitation id="134895BFFFC788347780CAC6FDCDF865" box="[445,512,1932,1958]" captionStart="FIGURE 2" captionStartId="6.[151,255,1977,2001]" captionTargetBox="[213,1375,309,1976]" captionTargetId="figure@6.[213,1397,287,1987]" captionTargetPageId="6" captionText="FIGURE 2: A. S. formosa holotype (NTUM 01329), immature female, 328 mm, B. Close-up of caudal fin, C. Close-up of snout." httpUri="https://zenodo.org/record/178135/files/figure.png" pageId="5" pageNumber="36">Fig 2</figureCitation>
|
||
). First,
|
||
<taxonomicName id="4C73F2B9FFC788347463CAC6FD10F866" box="[606,733,1932,1957]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC788347463CAC6FD10F866" box="[606,733,1932,1957]" italics="true" pageId="5" pageNumber="36">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
possesses a lobed caudal fin, especially in the dorsal lobe, and has a more curved postventral caudal margin. Second,
|
||
<taxonomicName id="4C73F2B9FFC788347571CAFEFC07F80E" box="[844,970,1972,1997]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC788347571CAFEFC07F80E" box="[844,970,1972,1997]" italics="true" pageId="5" pageNumber="36">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
has lobed dorsal fins with a curvilinear anterior margin. Third, the pelvic girdle distance in
|
||
<taxonomicName id="4C73F2B9FFC7883474C4CA96FCBAF836" box="[761,887,2012,2037]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="5" pageNumber="36" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC7883474C4CA96FCBAF836" box="[761,887,2012,2037]" italics="true" pageId="5" pageNumber="36">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
is more narrow than other WNP squatinids, at 1.4 or less the head length (where both measurements are standardized by total length). Fourth,
|
||
<taxonomicName id="4C73F2B9FFC4883772CCCDD2FAA3FF72" box="[1265,1390,152,177]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC4883772CCCDD2FAA3FF72" box="[1265,1390,152,177]" italics="true" pageId="6" pageNumber="37">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
has an upper lip arch which is semicircular in shape, where the upper lip arch height is greater than 1.5% of the total length.
|
||
</paragraph>
|
||
<caption id="DF0CD9B2FFC4883776AACAF3FF38F830" httpUri="https://zenodo.org/record/178135/files/figure.png" pageId="6" pageNumber="37" targetBox="[213,1375,309,1976]" targetPageId="6">
|
||
<paragraph id="8BCC893AFFC4883776AACAF3FF38F830" blockId="6.[151,1436,1977,2035]" pageId="6" pageNumber="37">
|
||
<emphasis id="B9075528FFC4883776AACAF3FEDFF812" bold="true" box="[151,274,1977,2001]" pageId="6" pageNumber="37">FIGURE 2</emphasis>
|
||
: A.
|
||
<taxonomicName id="4C73F2B9FFC48837777ACAF0FE74F812" box="[327,441,1978,2001]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC48837777ACAF0FE74F812" box="[327,441,1978,2001]" italics="true" pageId="6" pageNumber="37">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
holotype (NTUM 01329), immature female, 328 mm, B. Close-up of caudal fin, C. Close-up of snout.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC893AFFC5883676FBCDD2FC97FE61" blockId="7.[151,1436,152,418]" pageId="7" pageNumber="38">
|
||
Comparison of the three
|
||
<taxonomicName id="4C73F2B9FFC588367436CDD2FD5CFF72" box="[523,657,152,177]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC588367436CDD2FD5CFF72" box="[523,657,152,177]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54C83798FFC58836749DCDD2FCDDFF71" box="[672,784,152,178]" pageId="7" pageNumber="38" type="paratype">paratypes</typeStatus>
|
||
(originally labeled as NTU7041631, NTU7041632, NTU
|
||
<quantity id="4C8B24DFFFC5883676EECD8AFE95FF19" box="[211,344,192,218]" metricMagnitude="5" metricUnit="m" metricValue="1.8344979819999998" pageId="7" pageNumber="38" unit="in" value="7222433.0">7222433 in</quantity>
|
||
<bibRefCitation id="EFE2F4CBFFC588367762CD8AFD80FF19" author="Shen" box="[351,589,192,218]" pageId="7" pageNumber="47" refString="Shen S-C., & Ting, W-H. (1972) Ecological and morphological study on fish fauna from the waters around Taiwan and its adjacent islands. Notes on some continental shelf fishes and descriptions of two new species. Bulletin of the Institute of Zoologic Academy - Sinica, 11, 13 - 31." type="journal article" year="1972">Shen and Ting 1972</bibRefCitation>
|
||
; now labeled as NTUM 01327(
|
||
<date id="FFCDAFFAFFC5883675FCCD8AFC13FF19" box="[961,990,192,218]" pageId="7" pageNumber="38">x2</date>
|
||
) and NTUM01328) with the
|
||
<typeStatus id="54C83798FFC58836730BCD8AFA51FF19" box="[1334,1436,192,218]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
reveals differences among these four characters (Table 2). The
|
||
<typeStatus id="54C83798FFC5883675BCCDA2FC3CFEC1" box="[897,1009,232,258]" pageId="7" pageNumber="38" type="paratype">paratypes</typeStatus>
|
||
possess angular caudal fins without curved postventral caudal margins, angular dorsal fins with straight anterior margins, a wider pelvic girdle distance, and upper lip arches which are more semi-oval in shape (
|
||
<figureCitation id="134895BFFFC5883675A1CC72FC15FE91" box="[924,984,312,338]" captionStart="FIGURE 3" captionStartId="8.[151,255,1930,1954]" captionTargetBox="[189,1382,196,1882]" captionTargetId="figure@8.[177,1410,180,1906]" captionTargetPageId="8" captionText="FIGURE 3: Comparisons of true S. formosa holotype (NTUM 01329) with mislabeled S. fomosa paratype (NTUM 01328). A. Caudal fins, B. Dorsal fins, C. Upper lip arches." httpUri="https://zenodo.org/record/178136/files/figure.png" pageId="7" pageNumber="38">Fig 3</figureCitation>
|
||
A-C). Therefore, our contention is that these
|
||
<typeStatus id="54C83798FFC5883676E1CC2AFE81FEB9" box="[220,332,352,378]" pageId="7" pageNumber="38" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C73F2B9FFC58836774BCC2AFE38FEBA" box="[374,501,352,377]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC58836774BCC2AFE38FEBA" box="[374,501,352,377]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
are, in fact, a different species. Furthermore, these characters are most consistent with the
|
||
<taxonomicName id="4C73F2B9FFC58836770CCCC2FE75FE62" box="[305,440,392,417]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="nebulosa">
|
||
<emphasis id="B9075528FFC58836770CCCC2FE75FE62" box="[305,440,392,417]" italics="true" pageId="7" pageNumber="38">S. nebulosa</emphasis>
|
||
</taxonomicName>
|
||
comparison material we examined.
|
||
</paragraph>
|
||
<paragraph id="8BCC893AFFC5883676AACCA8FC7CFE38" blockId="7.[151,945,479,507]" box="[151,945,479,507]" pageId="7" pageNumber="38">
|
||
<emphasis id="B9075528FFC5883676AACCA8FEC7FE39" bold="true" box="[151,266,482,506]" pageId="7" pageNumber="38">TABLE 2:</emphasis>
|
||
Distinguishing characters between
|
||
<taxonomicName id="4C73F2B9FFC5883674B5CCA8FD37FE3A" box="[648,762,482,505]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC5883674B5CCA8FD37FE3A" box="[648,762,482,505]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C73F2B9FFC588367513CCA8FC64FE3A" box="[814,937,482,505]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="nebulosa">
|
||
<emphasis id="B9075528FFC588367513CCA8FC64FE3A" box="[814,937,482,505]" italics="true" pageId="7" pageNumber="38">S. nebulosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369DAB1FFC5883676A1CF6BFD0FFA98" pageId="7" pageNumber="38" type="description">
|
||
<paragraph id="8BCC893AFFC5883676A1CF6BFC56FDFB" blockId="7.[156,1349,545,779]" box="[156,923,545,568]" pageId="7" pageNumber="38">
|
||
<emphasis id="B9075528FFC5883676A1CF6BFC56FDFB" box="[156,923,545,568]" italics="true" pageId="7" pageNumber="38">
|
||
<taxonomicName id="4C73F2B9FFC5883676A1CF6BFEC2FDFB" box="[156,271,545,568]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">S. formosa</taxonomicName>
|
||
<taxonomicName id="4C73F2B9FFC58836751DCF6BFC56FDFB" box="[800,923,545,568]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="nebulosa">S. nebulosa</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC893AFFC5883676A1CF04FE9AFD43" blockId="7.[156,1349,545,779]" pageId="7" pageNumber="38">1. Caudal fin lobed, especially dorsally; curvilinear caudal 1. Caudal fin angular; straight caudal postventral margin. postventral margin.</paragraph>
|
||
<paragraph id="8BCC893AFFC5883676A1CFD2FB92FCC8" blockId="7.[156,1349,545,779]" pageId="7" pageNumber="38">2. Dorsal fins lobed; curvilinear dorsal anterior margins. 2. Dorsal fins angular; straight dorsal anterior margins. 3. Pelvic Gridle Width / Head Length <1.4 3. Pelvic Gridle Width / Head Length>1.4 4. Upper Arch Height> 1.5% TL 4. Upper Arch Height <1.5% TL</paragraph>
|
||
<paragraph id="8BCC893AFFC5883676FBCE2BFEC8FBA8" blockId="7.[151,1437,865,1731]" pageId="7" pageNumber="38">
|
||
An apparent change in designation of one of the
|
||
<typeStatus id="54C83798FFC5883674C1CE2BFCA1FCB8" box="[764,876,865,891]" pageId="7" pageNumber="38" type="paratype">paratypes</typeStatus>
|
||
for unknown reasons has also added to the confusion in the
|
||
<typeStatus id="54C83798FFC588367712CEC3FEACFC60" box="[303,353,905,931]" pageId="7" pageNumber="38">type</typeStatus>
|
||
series. At present, one of the
|
||
<typeStatus id="54C83798FFC588367481CEC3FCE1FC60" box="[700,812,905,931]" pageId="7" pageNumber="38" type="paratype">paratypes</typeStatus>
|
||
(NTUM 01328) has been designated as the
|
||
<typeStatus id="54C83798FFC588367313CEC3FA55FC60" box="[1326,1432,905,931]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
, and the true
|
||
<typeStatus id="54C83798FFC588367714CEFBFE42FC08" box="[297,399,945,971]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
(NTUM 01329) now has a question mark on its catalogue card next to its
|
||
<typeStatus id="54C83798FFC5883672C0CEFBFAA9FC08" box="[1277,1380,945,971]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
designation, as documented by the junior author (DAE) who examined the
|
||
<typeStatus id="54C83798FFC5883675CACE93FBE4FC30" box="[1015,1065,985,1011]" pageId="7" pageNumber="38">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C73F2B9FFC588367280CE93FAF0FC31" box="[1213,1341,985,1010]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC588367280CE93FAF0FC31" box="[1213,1341,985,1010]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<date id="FFCDAFFAFFC58836735BCE93FF1FFBD8" pageId="7" pageNumber="38" value="1988-05">May 1988</date>
|
||
and again in
|
||
<date id="FFCDAFFAFFC58836774BC94BFE3EFBD8" box="[374,499,1025,1051]" pageId="7" pageNumber="38" value="2005-05">May 2005</date>
|
||
. However, comparison of the
|
||
<typeStatus id="54C83798FFC588367566C94BFC43FBD8" box="[859,910,1025,1051]" pageId="7" pageNumber="38">type</typeStatus>
|
||
material with photos of the
|
||
<typeStatus id="54C83798FFC5883672DCC94BFA85FBD8" box="[1249,1352,1025,1051]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
within the original species description (
|
||
<bibRefCitation id="EFE2F4CBFFC588367425C963FCCDFB80" author="Shen" box="[536,768,1065,1091]" pageId="7" pageNumber="47" refString="Shen S-C., & Ting, W-H. (1972) Ecological and morphological study on fish fauna from the waters around Taiwan and its adjacent islands. Notes on some continental shelf fishes and descriptions of two new species. Bulletin of the Institute of Zoologic Academy - Sinica, 11, 13 - 31." type="journal article" year="1972">Shen & Ting, 1972</bibRefCitation>
|
||
) confirms that the NTUM 0 1329 specimen is the actual
|
||
<typeStatus id="54C83798FFC5883676AAC91BFECFFBA8" box="[151,258,1105,1131]" pageId="7" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC893AFFC5883676FBC933FD0FFA98" blockId="7.[151,1437,865,1731]" pageId="7" pageNumber="38">
|
||
A squatinid reported from the
|
||
<collectingCountry id="F364C9AAFFC58836740FC933FD7BFB50" box="[562,694,1145,1171]" name="Philippines" pageId="7" pageNumber="38">Philippines</collectingCountry>
|
||
as
|
||
<taxonomicName id="4C73F2B9FFC5883674DDC933FC92FB51" box="[736,863,1145,1170]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC5883674DDC933FC92FB51" box="[736,863,1145,1170]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE2F4CBFFC58836754DC933FB5FFB50" author="Compagno" box="[880,1170,1145,1171]" pageId="7" pageNumber="47" refString="Compagno L. J. V., Last, PR, Stevens, JD & Alava, MNR. (2005 b) Checklist of Philippine Chondrichthyes, CSIRO Marine Laboratories, Report 243, Hobart, Australia, 109 pp." type="book" year="2005" yearSuffix="b">Compagno et al., 2005b</bibRefCitation>
|
||
) is also not likely this species. Examination of photographs of the
|
||
<collectingCountry id="F364C9AAFFC588367490C9EBFCFCFB78" box="[685,817,1185,1211]" name="Philippines" pageId="7" pageNumber="38">Philippines</collectingCountry>
|
||
specimen, provided by L.J.V. Compagno, revealed several characters inconsistent with
|
||
<taxonomicName id="4C73F2B9FFC588367478C983FD09FB21" box="[581,708,1225,1250]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC588367478C983FD09FB21" box="[581,708,1225,1250]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
. These characters include pelvic fins which do not reach the first dorsal base, a wider pelvic girdle, a more shallow upper lip arch and distinctly different coloration than observed in
|
||
<taxonomicName id="4C73F2B9FFC58836771BC853FE6EFAF1" box="[294,419,1305,1330]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC58836771BC853FE6EFAF1" box="[294,419,1305,1330]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, the
|
||
<collectingCountry id="F364C9AAFFC588367462C853FD2EFAF0" box="[607,739,1305,1331]" name="Philippines" pageId="7" pageNumber="38">Philippines</collectingCountry>
|
||
specimen most likely represents a different, possibly undescribed, squatinid species from true
|
||
<taxonomicName id="4C73F2B9FFC588367403C80BFD76FA99" box="[574,699,1345,1370]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC588367403C80BFD76FA99" box="[574,699,1345,1370]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369DAB1FFC5883676FBC822FC38F900" pageId="7" pageNumber="38" type="materials_examined">
|
||
<paragraph id="8BCC893AFFC5883676FBC822FB79F988" blockId="7.[151,1437,865,1731]" pageId="7" pageNumber="38">
|
||
<emphasis id="B9075528FFC5883676FBC822FE0CFA41" bold="true" box="[198,449,1384,1410]" pageId="7" pageNumber="38">Material Examined.</emphasis>
|
||
<typeStatus id="54C83798FFC5883677F4C823FDCEFA40" box="[457,515,1385,1411]" pageId="7" pageNumber="38">Type</typeStatus>
|
||
material:
|
||
<typeStatus id="54C83798FFC588367447C823FD2AFA40" box="[634,743,1385,1411]" pageId="7" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
<taxonomicName id="4C73F2B9FFC5883674D2C823FCA0FA41" box="[751,877,1385,1410]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">
|
||
<emphasis id="B9075528FFC5883674D2C823FCA0FA41" box="[751,877,1385,1410]" italics="true" pageId="7" pageNumber="38">S. formosa</emphasis>
|
||
</taxonomicName>
|
||
: NTT7213130 (now labeled as
|
||
<collectionCode id="ED6211FFFFC5883672DEC823FAF7FA40" box="[1251,1338,1385,1411]" pageId="7" pageNumber="38">NTUM</collectionCode>
|
||
01329), immature female, Tung-Kang, Pingtung,
|
||
<collectingCountry id="F364C9AAFFC588367444C8DBFD1FFA68" box="[633,722,1425,1451]" name="Taiwan" pageId="7" pageNumber="38">Taiwan</collectingCountry>
|
||
,
|
||
<date id="FFCDAFFAFFC5883674E0C8DBFCA1FA68" box="[733,876,1425,1451]" pageId="7" pageNumber="38" value="1972-01-31">31 Jan 1972</date>
|
||
, identified by S.C. Shen.
|
||
<typeStatus id="54C83798FFC5883672A1C8DBFAC3FA68" box="[1180,1294,1425,1451]" pageId="7" pageNumber="38" type="paratype">Paratypes</typeStatus>
|
||
<emphasis id="B9075528FFC588367328C8DBFA51FA69" box="[1301,1436,1425,1450]" italics="true" pageId="7" pageNumber="38">
|
||
<taxonomicName id="4C73F2B9FFC588367328C8DBFA5BFA69" box="[1301,1430,1425,1450]" class="Elasmobranchii" family="Squatinidae" genus="Squatina" kingdom="Animalia" order="Squatiniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="formosa">S. formosa</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
NTU7222433 (now labeled as
|
||
<collectionCode id="ED6211FFFFC58836743DC8F3FD9BFA10" box="[512,598,1465,1491]" pageId="7" pageNumber="38">NTUM</collectionCode>
|
||
01327), and NTU7041632 (now labeled as
|
||
<collectionCode id="ED6211FFFFC588367267C8F3FB7DFA10" box="[1114,1200,1465,1491]" pageId="7" pageNumber="38">NTUM</collectionCode>
|
||
1327), caught
|
||
<quantity id="4C8B24DFFFC58836735DC8F3FA51FA10" box="[1376,1436,1465,1491]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="38" unit="km" value="3.0">3 km</quantity>
|
||
off the coast of Tahsi,
|
||
<collectingCountry id="F364C9AAFFC588367791C8ABFDCFFA38" box="[428,514,1505,1531]" name="Taiwan" pageId="7" pageNumber="38">Taiwan</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE47EFFDFFC588367428C8ABFD59FA38" box="[533,660,1505,1531]" direction="north" orientation="latitude" pageId="7" pageNumber="38" precision="92" value="24.941668">24°56.5’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE47EFFDFFC58836749EC8ABFCE6FA38" box="[675,811,1505,1531]" direction="east" orientation="longitude" pageId="7" pageNumber="38" precision="92" value="121.88333">121°53.0’E</geoCoordinate>
|
||
) in single trawling net at 100-120 fathoms,
|
||
<date id="FFCDAFFAFFC58836737BC8ABFF19F9E0" pageId="7" pageNumber="38" value="1970-04-16">16 Apr 1970</date>
|
||
, collected by W.H. Ting, identified by S.C. Shen; NTU7041632 (now labeled as
|
||
<collectionCode id="ED6211FFFFC5883672B2CB43FB2BF9E0" box="[1167,1254,1545,1571]" pageId="7" pageNumber="38">NTUM</collectionCode>
|
||
01328), immature female, Tahsi,
|
||
<collectingCountry id="F364C9AAFFC58836774ACB7BFE1DF988" box="[375,464,1585,1611]" name="Taiwan" pageId="7" pageNumber="38">Taiwan</collectingCountry>
|
||
,
|
||
<date id="FFCDAFFAFFC5883677E1CB7BFDBCF988" box="[476,625,1585,1611]" pageId="7" pageNumber="38" value="1972-02-24">24 Feb 1972</date>
|
||
, collected by W.H. Ting, identified by S.C. Shen.
|
||
</paragraph>
|
||
<paragraph id="8BCC893AFFC5883676FBCB13FC38F900" blockId="7.[151,1437,865,1731]" pageId="7" pageNumber="38">
|
||
Comparative material: DAE 881805, immature male, Tahsi,
|
||
<collectingCountry id="F364C9AAFFC5883675A2CB13FC35F9B0" box="[927,1016,1625,1651]" name="Taiwan" pageId="7" pageNumber="38">Taiwan</collectingCountry>
|
||
,
|
||
<date id="FFCDAFFAFFC58836723BCB13FB4EF9B0" box="[1030,1155,1625,1651]" pageId="7" pageNumber="38" value="1988-05">May 1988</date>
|
||
, collected by David A. Ebert; DAE 0 52105, immature male, Tahsi,
|
||
<collectingCountry id="F364C9AAFFC58836749FCBCBFD36F958" box="[674,763,1665,1691]" name="Taiwan" pageId="7" pageNumber="38">Taiwan</collectingCountry>
|
||
,
|
||
<date id="FFCDAFFAFFC58836753BCBCBFC4DF958" box="[774,896,1665,1691]" pageId="7" pageNumber="38" value="2005-05">May 2005</date>
|
||
, collected by David A. Ebert; DAE
|
||
<date id="FFCDAFFAFFC588367318CBCBFA4FF958" box="[1317,1410,1665,1691]" pageId="7" pageNumber="38" value="05-05-23">052305</date>
|
||
-2, immature female, Tahsi,
|
||
<collectingCountry id="F364C9AAFFC588367787CBE3FDDEF900" box="[442,531,1705,1731]" name="Taiwan" pageId="7" pageNumber="38">Taiwan</collectingCountry>
|
||
,
|
||
<date id="FFCDAFFAFFC588367422CBE3FD54F900" box="[543,665,1705,1731]" pageId="7" pageNumber="38" value="2005-05">May 2005</date>
|
||
, collected by David A. Ebert.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |