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<document id="CBB720FDFAF77B2C38B410314199A162" ID-DOI="10.37828/em.2023.68.14" ID-ISSN="2336-9744" ID-Zenodo-Dep="13247254" ID-ZooBank="urn:lsid:zoobank.org:pub:AAC1A7FA-3259-454E-A278-EE4F64426DD1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722972682661" checkinUser="felipe" docAuthor="Volynkin, Anton V. &amp; Saldaitis, Aidas" docDate="2023" docId="03CCE50E4716FFDD899551A39207FB0B" docLanguage="en" docName="EcolMontenegrina.68.140-154.pdf" docOrigin="Ecologica Montenegrina 68" docSource="http://dx.doi.org/10.37828/em.2023.68.14" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Kuatunittia Volynkin &amp; Saldaitis, 2023, gen. n." docType="treatment" docUuid="D00B6BA1-EF3E-4613-82F6-ABC78B1B0CDE" docUuidSource="ZooBank" docVersion="3" lastPageNumber="149" masterDocId="FFF59D76471FFFD489025167965CFFA1" masterDocTitle="Two new genera, two new subgenera, and four new species of Lithosiina from China and northern Indochina (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="154" masterPageNumber="140" pageNumber="149" updateTime="1722981492806" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="904A5F7CB6E1FB90D69DDF652C586790">Two new genera, two new subgenera, and four new species of Lithosiina from China and northern Indochina (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="02AB846E0EB64E39517ABA8B1B89BFAF">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="31C7800D725D5BDADF86F596FBE2A2BE">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia.</mods:affiliation>
<mods:nameIdentifier id="58A4951D26B59375CAC1E4A99177FE6C" type="email">monstruncusarctia@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E7684B5EF90B965FA0FA64C2B17D3DD7">Saldaitis, Aidas</mods:namePart>
<mods:nameIdentifier id="21332ACE390C8BC95682160C625ED11B" type="ORCID">0000-0003-0999-3996</mods:nameIdentifier>
<mods:affiliation id="3C3C387DB841E797CA14D00BD11B422E">Nature Research Centre, Akademijos str., 2, LT- 08412, Vilnius- 21, Lithuania. E-mail: saldrasa @ gmail. com; https: // orcid. org / 0000 - 0003 - 0999 - 3996 * Corresponding author. E-mail: monstruncusarctia @ gmail. com; https: // orcid. org / 0000 - 0001 - 9447 - 4925</mods:affiliation>
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<mods:title id="E9636FEE59281A2062FBDF8189F26C79">Ecologica Montenegrina</mods:title>
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<mods:date id="E6B6E1F0A8F2206B18606E9B4EE9AC74">2023</mods:date>
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<mods:number id="28197D7CEB79C60B75930EAEAB7B75A0">2023-11-13</mods:number>
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<mods:number id="E4837BF4F190EFAFE8C13FB4538E9C54">68</mods:number>
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<treatment id="03CCE50E4716FFDD899551A39207FB0B" ID-DOI="http://doi.org/10.5281/zenodo.13247795" ID-Zenodo-Dep="13247795" LSID="urn:lsid:zoobank.org:act:D00B6BA1-EF3E-4613-82F6-ABC78B1B0CDE" httpUri="http://treatment.plazi.org/id/03CCE50E4716FFDD899551A39207FB0B" lastPageNumber="149" pageId="9" pageNumber="149">
<subSubSection id="C37F07934716FFDD899551A39224FF5D" pageId="9" pageNumber="149" type="nomenclature">
<paragraph id="8BDA54184716FFDD899551A39792FF7F" blockId="9.[151,1144,196,283]" box="[151,462,196,222]" pageId="9" pageNumber="149">
<heading id="D092E3744716FFDD899551A39792FF7F" box="[151,462,196,222]" fontSize="11" level="2" pageId="9" pageNumber="149" reason="7">
Genus
<emphasis id="B911880A4716FFDD89E851A39792FF7F" bold="true" box="[234,462,196,222]" pageId="9" pageNumber="149">
<taxonomicName id="4C652F9B4716FFDD89E851A39728FF7F" ID-CoL="BXN3F" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[234,372,196,222]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD89E851A39728FF7F" bold="true" box="[234,372,196,222]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
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<taxonomicNameLabel id="A22235714716FFDD887E51A39792FF7F" box="[380,462,196,222]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
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<paragraph id="8BDA54184716FFDD899551829224FF5D" blockId="9.[151,1144,196,283]" box="[151,1144,229,252]" pageId="9" pageNumber="149">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C37F07934716FFDD8995506395ECFEBA" box="[151,944,260,283]" pageId="9" pageNumber="149" type="materials_examined">
<paragraph id="8BDA54184716FFDD8995506395ECFEBA" blockId="9.[151,1144,196,283]" box="[151,944,260,283]" pageId="9" pageNumber="149">
<typeStatus id="54DEEABA4716FFDD899550639692FEBA" box="[151,206,260,283]" pageId="9" pageNumber="149">Type</typeStatus>
species:
<taxonomicName id="4C652F9B4716FFDD882D506394F1FEBA" authority="Daniel, 1954" authorityName=", Daniel" authorityYear="1954" box="[303,685,260,283]" class="Insecta" family="Halictidae" genus="Systropha" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="B911880A4716FFDD882D50639446FEBA" box="[303,538,260,283]" italics="true" pageId="9" pageNumber="149">Systropha klapperichi</emphasis>
<bibRefCitation id="EFF429E94716FFDD8B23506394F1FEBA" author="Daniel, F." box="[545,685,260,283]" pageId="9" pageNumber="149" pagination="89 - 138" refId="ref5916" refString="Daniel, F. (1954) Beitrage zur Kenntnis der Arctiidae Ostasiens unter besonderer Berucksichtigung der Ausbeuten von Dr. h. c. H. Hone aus diesem Gebiet (Lep. Het.). Bonner zoologische Beitrage, 5, 89 - 138. [in German]" type="journal article" year="1954">Daniel, 1954</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C37F07934716FFDD899550229566FBC6" pageId="9" pageNumber="149" type="diagnosis">
<paragraph id="8BDA54184716FFDD89955022921FFE47" blockId="9.[151,1437,325,1194]" pageId="9" pageNumber="149">
<emphasis id="B911880A4716FFDD89955022974CFEFE" bold="true" box="[151,272,325,351]" pageId="9" pageNumber="149">Diagnosis</emphasis>
. Species of the new genus (
<figureCitation id="135E489D4716FFDD8B65502294BFFEFE" box="[615,739,325,351]" captionStart-0="Figures 1–8" captionStart-1="Figures 9–18" captionStartId-0="4.[151,237,1947,1970]" captionStartId-1="5.[151,237,1934,1957]" captionTargetBox-0="[196,1424,190,1916]" captionTargetBox-1="[160,1430,193,1905]" captionTargetId-0="figure-14@4.[151,1436,189,1928]" captionTargetId-1="figure-10@5.[156,1431,189,1916]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Figures 1–8. Fangeilema (1–6) and Kuatunittia (7, 8), adults. Depositories of the specimens: 1 and 2 in NHMUK (©The Trustees of NHMUK); 3–6, and 8 in MWM/ZSM; 7 in ZFMK (photo by V. Zolotuhin, by courtesy of V. Dubatolov)." captionText-1="Figures 9–18. Adults of Kuatunittia (9, 10), and the type species of the corresponding genera: Wittia (11, 12), Pseudoscaptia (13, 14), Dolgoma (15, 16), and Teuloma (17, 18). Depositories of the specimens: 9 in WIGJ; 10 in MWM/ZSM; 11 and 12 in ANHRT; 13 and 14 in NHMUK (©The Trustees of NHMUK); 15 and 16 in CKC; 17 and 18 in CAV." figureDoi-0="http://doi.org/10.5281/zenodo.13247256" figureDoi-1="http://doi.org/10.5281/zenodo.13247258" httpUri-0="https://zenodo.org/record/13247256/files/figure.png" httpUri-1="https://zenodo.org/record/13247258/files/figure.png" pageId="9" pageNumber="149">Figs 7–10</figureCitation>
,
<figureCitation id="135E489D4716FFDD8BF350229560FEFE" box="[753,828,325,351]" captionStart="Figures 23–26" captionStartId="7.[151,237,1947,1970]" captionTargetBox="[171,1428,200,1911]" captionTargetId="figure-10@7.[151,1436,189,1928]" captionTargetPageId="7" captionText="Figures 23–26. Kuatunittia, male genitalia. Depositories of the specimens dissected: 23, 25 and 26 in MWM/ZSM; 24 in WIGJ." figureDoi="http://doi.org/10.5281/zenodo.13247262" httpUri="https://zenodo.org/record/13247262/files/figure.png" pageId="9" pageNumber="149">23–26</figureCitation>
) are externally reminiscent of
<taxonomicName id="4C652F9B4716FFDD8DB350229361FEFE" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[1201,1341,325,351]" class="Insecta" family="Erebidae" genus="Fangeilema" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8DB350229361FEFE" box="[1201,1341,325,351]" italics="true" pageId="9" pageNumber="149">Fangeilema</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8C45502293C0FEFE" bold="true" box="[1351,1436,325,351]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8C45502293C0FEFE" box="[1351,1436,325,351]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135E489D4716FFDD899D50009755FE20" box="[159,265,359,385]" captionStart="Figures 1–8" captionStartId="4.[151,237,1947,1970]" captionTargetBox="[196,1424,190,1916]" captionTargetId="figure-14@4.[151,1436,189,1928]" captionTargetPageId="4" captionText="Figures 1–8. Fangeilema (1–6) and Kuatunittia (7, 8), adults. Depositories of the specimens: 1 and 2 in NHMUK (©The Trustees of NHMUK); 3–6, and 8 in MWM/ZSM; 7 in ZFMK (photo by V. Zolotuhin, by courtesy of V. Dubatolov)." figureDoi="http://doi.org/10.5281/zenodo.13247256" httpUri="https://zenodo.org/record/13247256/files/figure.png" pageId="9" pageNumber="149">Figs 1–6</figureCitation>
,
<figureCitation id="135E489D4716FFDD88145000973EFE20" box="[278,354,359,385]" captionStart="Figures 19–22" captionStartId="6.[151,237,1947,1970]" captionTargetBox="[158,1420,197,1906]" captionTargetId="figure-14@6.[151,1436,189,1928]" captionTargetPageId="6" captionText="Figures 19–22. Fangeilema, male genitalia. Depositories of the specimens dissected: 19 in NHMUK (©The Trustees of NHMUK); 20–22 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13247260" httpUri="https://zenodo.org/record/13247260/files/figure.png" pageId="9" pageNumber="149">19–22</figureCitation>
) but differ in the distally bilobate saccular process. In the female genitalia,
<taxonomicName id="4C652F9B4716FFDD8DDE5000933EFE20" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[1244,1378,359,385]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8DDE5000933EFE20" box="[1244,1378,359,385]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8C6B500096F3FE03" bold="true" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8C6B500096F3FE03" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135E489D4716FFDD89C250EF9744FE03" box="[192,280,392,418]" captionStart="Figures 31–34" captionStartId="11.[151,237,1947,1970]" captionTargetBox="[218,1390,199,1911]" captionTargetId="figure-10@11.[151,1436,189,1928]" captionTargetPageId="11" captionText="Figures 31–34. Fangeilema (31–33) and Kuatunittia, female genitalia. Depositories of the specimens dissected: 31 in NHMUK (©The Trustees of NHMUK); 32–34 in MWM/ZSM" figureDoi="http://doi.org/10.5281/zenodo.13247268" httpUri="https://zenodo.org/record/13247268/files/figure.png" pageId="9" pageNumber="149">Fig. 34</figureCitation>
) is distinguished from
<taxonomicName id="4C652F9B4716FFDD8B2F50EF94E6FE03" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[557,698,392,418]" class="Insecta" family="Erebidae" genus="Fangeilema" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8B2F50EF94E6FE03" box="[557,698,392,418]" italics="true" pageId="9" pageNumber="149">Fangeilema</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8BC150EF954BFE03" bold="true" box="[707,791,392,418]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8BC150EF954BFE03" box="[707,791,392,418]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135E489D4716FFDD8A2550EF95EDFE03" box="[807,945,392,418]" captionStart="Figures 31–34" captionStartId="11.[151,237,1947,1970]" captionTargetBox="[218,1390,199,1911]" captionTargetId="figure-10@11.[151,1436,189,1928]" captionTargetPageId="11" captionText="Figures 31–34. Fangeilema (31–33) and Kuatunittia, female genitalia. Depositories of the specimens dissected: 31 in NHMUK (©The Trustees of NHMUK); 32–34 in MWM/ZSM" figureDoi="http://doi.org/10.5281/zenodo.13247268" httpUri="https://zenodo.org/record/13247268/files/figure.png" pageId="9" pageNumber="149">Figs 31, 32</figureCitation>
) by the conical appendix bursae directed posteriorly (it is sack-like and directed laterally in
<taxonomicName id="4C652F9B4716FFDD8A0350CD95D1FE65" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[769,909,426,452]" class="Insecta" family="Erebidae" genus="Fangeilema" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8A0350CD95D1FE65" box="[769,909,426,452]" italics="true" pageId="9" pageNumber="149">Fangeilema</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8A9550CD95B1FE65" bold="true" box="[919,1005,426,452]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8A9550CD95B1FE65" box="[919,1005,426,452]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
), and the presence of the band-like signum (whereas species of
<taxonomicName id="4C652F9B4716FFDD88E050AB9433FE47" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[482,623,460,486]" class="Insecta" family="Erebidae" genus="Fangeilema" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD88E050AB9433FE47" box="[482,623,460,486]" italics="true" pageId="9" pageNumber="149">Fangeilema</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8B7450AB9494FE47" bold="true" box="[630,712,460,486]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8B7450AB9494FE47" box="[630,712,460,486]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
have round signa or lack them).
</paragraph>
<paragraph id="8BDA54184716FFDD89F7508A9566FBC6" blockId="9.[151,1437,325,1194]" pageId="9" pageNumber="149">
The similar bilobate sacculus is found in two other Asiatic Lithosiina genera,
<taxonomicName id="4C652F9B4716FFDD8DF3508A970FFD88" authority="Hampson, 1914" authorityName="Hampson" authorityYear="1914" class="Insecta" family="Erebidae" genus="Pseudoscaptia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8DF3508A93C0FDA6" box="[1265,1436,493,519]" italics="true" pageId="9" pageNumber="149">Pseudoscaptia</emphasis>
Hampson, 1914
</taxonomicName>
(
<figureCitation id="135E489D4716FFDD8866536897B1FD88" box="[356,493,527,553]" captionStart="Figures 9–18" captionStartId="5.[151,237,1934,1957]" captionTargetBox="[160,1430,193,1905]" captionTargetId="figure-10@5.[156,1431,189,1916]" captionTargetPageId="5" captionText="Figures 9–18. Adults of Kuatunittia (9, 10), and the type species of the corresponding genera: Wittia (11, 12), Pseudoscaptia (13, 14), Dolgoma (15, 16), and Teuloma (17, 18). Depositories of the specimens: 9 in WIGJ; 10 in MWM/ZSM; 11 and 12 in ANHRT; 13 and 14 in NHMUK (©The Trustees of NHMUK); 15 and 16 in CKC; 17 and 18 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13247258" httpUri="https://zenodo.org/record/13247258/files/figure.png" pageId="9" pageNumber="149">Figs 13, 14</figureCitation>
) and
<taxonomicName id="4C652F9B4716FFDD8B31536895ACFD88" authority="Volynkin &amp; N. Singh, 2019" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[563,1008,527,553]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8B31536894C6FD88" box="[563,666,527,553]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
Volynkin &amp; N.
<bibRefCitation id="EFF429E94716FFDD8A5D536895ACFD88" author="Singh, N. &amp; Kirti, J. S. &amp; Datta H. S. &amp; Joshi, R. &amp; Volynkin, A. V." box="[863,1008,527,553]" pageId="9" pageNumber="149" pagination="33 - 54" refId="ref6207" refString="Singh, N., Kirti, J. S., Datta H. S., Joshi, R. &amp; Volynkin, A. V. (2019) A review of the genus Dolgoma Moore from India, with notes on the genus taxonomy and descriptions of a new genus, four new species and a new subspecies (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa, 4683 (1), 33 - 54. https: // doi. org / 10.11646 / zootaxa. 4683.1.2" type="journal article" year="2019">Singh, 2019</bibRefCitation>
</taxonomicName>
(
<figureCitation id="135E489D4716FFDD8D02536892D0FD88" box="[1024,1164,527,553]" captionStart="Figures 9–18" captionStartId="5.[151,237,1934,1957]" captionTargetBox="[160,1430,193,1905]" captionTargetId="figure-10@5.[156,1431,189,1916]" captionTargetPageId="5" captionText="Figures 9–18. Adults of Kuatunittia (9, 10), and the type species of the corresponding genera: Wittia (11, 12), Pseudoscaptia (13, 14), Dolgoma (15, 16), and Teuloma (17, 18). Depositories of the specimens: 9 in WIGJ; 10 in MWM/ZSM; 11 and 12 in ANHRT; 13 and 14 in NHMUK (©The Trustees of NHMUK); 15 and 16 in CKC; 17 and 18 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13247258" httpUri="https://zenodo.org/record/13247258/files/figure.png" pageId="9" pageNumber="149">Figs 17, 18</figureCitation>
), but the former genus (
<figureCitation id="135E489D4716FFDD89A2535696AAFDEA" box="[160,246,561,587]" captionStart="Figures 27–30" captionStartId="8.[151,237,1947,1970]" captionTargetBox="[170,1436,200,1911]" captionTargetId="figure-14@8.[151,1436,189,1928]" captionTargetPageId="8" captionText="Figures 27–30. Lithosiina spp., male genitalia of the type species of the corresponding genera: Wittia (27), Pseudoscaptia (28), Dolgoma (29), and Teuloma (30). Depositories of the specimens dissected: 27 in ANHRT; 28 in NHMUK (©The Trustees of NHMUK); 29 in CKC; 30 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13247266" httpUri="https://zenodo.org/record/13247266/files/figure.png" pageId="9" pageNumber="149">Fig. 28</figureCitation>
) differs from
<taxonomicName id="4C652F9B4716FFDD889E5356947EFDEA" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[412,546,561,587]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD889E5356947EFDEA" box="[412,546,561,587]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8B2853569422FDEA" bold="true" box="[554,638,561,587]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8B2853569422FDEA" box="[554,638,561,587]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
in the asymmetrical sacculi sparsely covered with tiny spikes (they are symmetrical and densely covered with setae in
<taxonomicName id="4C652F9B4716FFDD8BF35334952BFDCC" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[753,887,595,621]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8BF35334952BFDCC" box="[753,887,595,621]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8A7D5334958EFDCC" bold="true" box="[895,978,595,621]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8A7D5334958EFDCC" box="[895,978,595,621]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
), the weakly sclerotised phallus carina (it is heavily sclerotised and bearing a cluster of setae in the new genus), and the vesica having a sclerotised and spinulose subbasal diverticulum. The female genitalia of
<taxonomicName id="4C652F9B4716FFDD8AB253F1926AFD11" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[944,1078,662,688]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8AB253F1926AFD11" box="[944,1078,662,688]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8D4453F192FDFD11" bold="true" box="[1094,1185,662,688]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8D4453F192FDFD11" box="[1094,1185,662,688]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
differ clearly from
<taxonomicName id="4C652F9B4716FFDD899553DF971EFD73" authorityName="Hampson" authorityYear="1914" box="[151,322,696,722]" class="Insecta" family="Erebidae" genus="Pseudoscaptia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD899553DF971EFD73" box="[151,322,696,722]" italics="true" pageId="9" pageNumber="149">Pseudoscaptia</emphasis>
</taxonomicName>
(
<figureCitation id="135E489D4716FFDD885753DF97F0FD73" box="[341,428,696,722]" captionStart="Figures 35–38" captionStartId="12.[151,237,1947,1970]" captionTargetBox="[158,1372,217,1911]" captionTargetId="figure-14@12.[151,1436,189,1928]" captionTargetPageId="12" captionText="Figures 35–38. Lithosiina spp., female genitalia of the type species of the corresponding genera: Wittia (35), Pseudoscaptia (36), Dolgoma (37), and Teuloma (38). Depositories of the specimens dissected: 35 in ANHRT; 36 in NHMUK (©The Trustees of NHMUK); 37 in CKC; 38 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13247272" httpUri="https://zenodo.org/record/13247272/files/figure.png" pageId="9" pageNumber="149">Fig. 36</figureCitation>
) in the symmetrical and smooth 7
<superScript id="7C10F9504716FFDD8A4A53D2950AFD64" attach="left" box="[840,854,693,709]" fontSize="7" pageId="9" pageNumber="149">th</superScript>
abdominal sternite lacking the antevaginal plate (it is asymmetrical and laterally rugose in
<taxonomicName id="4C652F9B4716FFDD8B9953BD951AFD55" authorityName="Hampson" authorityYear="1914" box="[667,838,730,756]" class="Insecta" family="Erebidae" genus="Pseudoscaptia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8B9953BD951AFD55" box="[667,838,730,756]" italics="true" pageId="9" pageNumber="149">Pseudoscaptia</emphasis>
</taxonomicName>
), and the narrow and symmetrical ductus bursae, which is broad and strongly asymmetrical in
<taxonomicName id="4C652F9B4716FFDD8BD0539B9521FCB7" authorityName="Hampson" authorityYear="1914" box="[722,893,764,790]" class="Insecta" family="Erebidae" genus="Pseudoscaptia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8BD0539B9521FCB7" box="[722,893,764,790]" italics="true" pageId="9" pageNumber="149">Pseudoscaptia</emphasis>
</taxonomicName>
. Compared to
<taxonomicName id="4C652F9B4716FFDD8D3F539B92F8FCB7" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[1085,1188,764,790]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8D3F539B92F8FCB7" box="[1085,1188,764,790]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
(
<figureCitation id="135E489D4716FFDD8DB8539B9344FCB7" box="[1210,1304,764,790]" captionStart="Figures 27–30" captionStartId="8.[151,237,1947,1970]" captionTargetBox="[170,1436,200,1911]" captionTargetId="figure-14@8.[151,1436,189,1928]" captionTargetPageId="8" captionText="Figures 27–30. Lithosiina spp., male genitalia of the type species of the corresponding genera: Wittia (27), Pseudoscaptia (28), Dolgoma (29), and Teuloma (30). Depositories of the specimens dissected: 27 in ANHRT; 28 in NHMUK (©The Trustees of NHMUK); 29 in CKC; 30 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13247266" httpUri="https://zenodo.org/record/13247266/files/figure.png" pageId="9" pageNumber="149">Fig. 30</figureCitation>
), the male genital capsule of
<taxonomicName id="4C652F9B4716FFDD887B527A97A3FC96" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[377,511,797,823]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD887B527A97A3FC96" box="[377,511,797,823]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8B08527A943CFC96" bold="true" box="[522,608,797,823]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8B08527A943CFC96" box="[522,608,797,823]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
has a markedly shorter and narrower vinculum, a juxta lacking the tubular and rugose dorsal extension (a feature characteristic of
<taxonomicName id="4C652F9B4716FFDD8AA552589251FCF8" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[935,1037,831,857]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8AA552589251FCF8" box="[935,1037,831,857]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
), and a markedly broader distal saccular process having a larger, setose and more proximally situated proximal lobe. The phallus of
<taxonomicName id="4C652F9B4716FFDD899552E59741FC3D" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[151,285,898,924]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD899552E59741FC3D" box="[151,285,898,924]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD882B52E597DCFC3D" bold="true" box="[297,384,898,924]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD882B52E597DCFC3D" box="[297,384,898,924]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
is shorter and narrower than in
<taxonomicName id="4C652F9B4716FFDD8A1652E59527FC3D" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[788,891,898,924]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8A1652E59527FC3D" box="[788,891,898,924]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
and has a considerably longer coecum. The vesica of
<taxonomicName id="4C652F9B4716FFDD880E52C397CEFC1F" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[268,402,932,958]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD880E52C397CEFC1F" box="[268,402,932,958]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD889952C397ACFC1F" bold="true" box="[411,496,932,958]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD889952C397ACFC1F" box="[411,496,932,958]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
is substantially shorter than in
<taxonomicName id="4C652F9B4716FFDD8A6E52C3958FFC1F" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[876,979,932,958]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8A6E52C3958FFC1F" box="[876,979,932,958]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
and bears a sclerotised serrulate plate distally whereas it is granulose without plates and cornuti in
<taxonomicName id="4C652F9B4716FFDD8A7E52A195BFFC41" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[892,995,966,992]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8A7E52A195BFFC41" box="[892,995,966,992]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
. The female genitalia of
<taxonomicName id="4C652F9B4716FFDD8C1552A193C0FC41" authorityName="Volynkin &amp; Saldaitis" authorityYear="2023" box="[1303,1436,966,992]" class="Insecta" family="Erebidae" genus="Kuatunittia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911880A4716FFDD8C1552A193C0FC41" box="[1303,1436,966,992]" italics="true" pageId="9" pageNumber="149">Kuatunittia</emphasis>
</taxonomicName>
<emphasis id="B911880A4716FFDD8995528096B2FBA0" bold="true" box="[151,238,999,1025]" pageId="9" pageNumber="149">
<taxonomicNameLabel id="A22235714716FFDD8995528096B2FBA0" box="[151,238,999,1025]" pageId="9" pageNumber="149" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
are distinguished from
<taxonomicName id="4C652F9B4716FFDD8B1552809422FBA0" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[535,638,999,1025]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8B1552809422FBA0" box="[535,638,999,1025]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
(
<figureCitation id="135E489D4716FFDD8B90528094B1FBA0" box="[658,749,999,1025]" captionStart="Figures 35–38" captionStartId="12.[151,237,1947,1970]" captionTargetBox="[158,1372,217,1911]" captionTargetId="figure-14@12.[151,1436,189,1928]" captionTargetPageId="12" captionText="Figures 35–38. Lithosiina spp., female genitalia of the type species of the corresponding genera: Wittia (35), Pseudoscaptia (36), Dolgoma (37), and Teuloma (38). Depositories of the specimens dissected: 35 in ANHRT; 36 in NHMUK (©The Trustees of NHMUK); 37 in CKC; 38 in CAV." figureDoi="http://doi.org/10.5281/zenodo.13247272" httpUri="https://zenodo.org/record/13247272/files/figure.png" pageId="9" pageNumber="149">Fig. 38</figureCitation>
) by the lack of the antrum, the markedly narrower and straight ductus bursae (it is curved in
<taxonomicName id="4C652F9B4716FFDD8B6B556E9493FB82" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[617,719,1033,1059]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8B6B556E9493FB82" box="[617,719,1033,1059]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
), the larger and sack-like corpus bursae (it is more or less globular in
<taxonomicName id="4C652F9B4716FFDD8822554C97DAFBE4" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[288,390,1067,1093]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8822554C97DAFBE4" box="[288,390,1067,1093]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
), and the shorter and conical appendix bursae directed posteriorly whereas it is medially helicoid, distally dilated and directed distally in
<taxonomicName id="4C652F9B4716FFDD8BCF552A956FFBC6" authorityName="Volynkin &amp; N. Singh" authorityYear="2019" box="[717,819,1101,1127]" class="Insecta" family="Erebidae" genus="Teuloma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B911880A4716FFDD8BCF552A956FFBC6" box="[717,819,1101,1127]" italics="true" pageId="9" pageNumber="149">Teuloma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F07934716FFDD89F755099207FB0B" pageId="9" pageNumber="149" type="etymology">
<paragraph id="8BDA54184716FFDD89F755099207FB0B" blockId="9.[151,1437,325,1194]" pageId="9" pageNumber="149">
<emphasis id="B911880A4716FFDD89F755099721FB29" bold="true" box="[245,381,1134,1160]" pageId="9" pageNumber="149">Etymology</emphasis>
. The genus name is an aggregate of the historical toponym Kuatun which is the
<typeStatus id="54DEEABA4716FFDD8C68550993C0FB29" box="[1386,1436,1134,1160]" pageId="9" pageNumber="149">type</typeStatus>
locality of the
<typeStatus id="54DEEABA4716FFDD884055F79729FB0B" box="[322,373,1168,1194]" pageId="9" pageNumber="149">type</typeStatus>
species, and the genus-group name
<taxonomicName id="4C652F9B4716FFDD8A1E55F79538FB0B" authorityName="de Freina" authorityYear="1980" box="[796,868,1168,1194]" class="Magnoliopsida" family="Cactaceae" genus="Wittia" higherTaxonomySource="IPNI" kingdom="Pantae" order="Caryophyllales" pageId="9" pageNumber="149" phylum="Magnoliophyta" rank="genus">
<emphasis id="B911880A4716FFDD8A1E55F79538FB0B" box="[796,868,1168,1194]" italics="true" pageId="9" pageNumber="149">Wittia</emphasis>
</taxonomicName>
. Gender is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>
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