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<document id="677BF23CA34B0453012A60EB5581BAEB" ID-DOI="10.5281/zenodo.189654" ID-GBIF-Dataset="320aec2f-ea3b-427e-a2a7-bdd4969cb030" ID-ISSN="1175-5326" ID-Zenodo-Dep="189654" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461103618203" checkinUser="plazi" docAuthor="Chiba, Hideyuki, Hsu, Yu-Feng, Tsukiyama, Hiroshi, Lo, Philip Yik-Fui, Chen, Chien-Ren &amp; Wang, Shou-Ming" docDate="2009" docId="03CC500DF301342DFF2CF6903B1EBE30" docLanguage="en" docName="zt02202p058.pdf" docOrigin="Zootaxa 2202" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Augiades bouddha subsp. niitakana Sonan 1936" docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="FFF52875F3043428FFBBF5023E1BB836" masterDocTitle="Jinhaku Sonans skipper type collection deposited at Taiwan Agricultural Research Institute (Lepidoptera: Hesperiidae)" masterLastPageNumber="58" masterPageNumber="48" pageNumber="53" updateTime="1698592197444" updateUser="plazi">
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<mods:title id="C76CF389C5D24FEF484D33D6780C9AEF">Jinhaku Sonans skipper type collection deposited at Taiwan Agricultural Research Institute (Lepidoptera: Hesperiidae)</mods:title>
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<mods:namePart id="3956DA1DDF8355E11397681AE52AC561">Chiba, Hideyuki</mods:namePart>
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<mods:namePart id="B07FC582167A6761D93CB1B4972DC8B6">Hsu, Yu-Feng</mods:namePart>
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<mods:namePart id="BD481C7AAACB379DF93C912215A7695E">Tsukiyama, Hiroshi</mods:namePart>
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<mods:namePart id="9FBC38AEF029B6CA7A005CB7A8B60B04">Lo, Philip Yik-Fui</mods:namePart>
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<mods:namePart id="4F3566151699A557A09D9421B36004E9">Chen, Chien-Ren</mods:namePart>
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<mods:namePart id="9F0A3B51D836A81F76271CE842ED8730">Wang, Shou-Ming</mods:namePart>
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<mods:date id="EE1919D513335063701D80F01B613314">2009</mods:date>
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<treatment id="03CC500DF301342DFF2CF6903B1EBE30" ID-DOI="http://doi.org/10.5281/zenodo.6225208" ID-GBIF-Taxon="119622494" ID-Zenodo-Dep="6225208" LSID="urn:lsid:plazi:treatment:03CC500DF301342DFF2CF6903B1EBE30" httpUri="http://treatment.plazi.org/id/03CC500DF301342DFF2CF6903B1EBE30" lastPageNumber="53" pageId="5" pageNumber="53">
<subSubSection id="C37FB290F301342DFF2CF6903F4CBBF9" pageId="5" pageNumber="53" type="nomenclature">
<paragraph id="8BDAE11BF301342DFF2CF6903C8ABB9A" blockId="5.[151,657,914,975]" box="[151,657,914,940]" pageId="5" pageNumber="53">
<heading id="D0925677F301342DFF2CF6903C8ABB9A" bold="true" box="[151,657,914,940]" fontSize="11" level="1" pageId="5" pageNumber="53" reason="1">
<taxonomicName id="4C659A98F301342DFF2CF6903C8ABB9A" authority="Sonan, 1936" authorityName="Sonan" authorityYear="1936" box="[151,657,914,940]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="bouddha" subSpecies="niitakana">
<emphasis id="B9113D09F301342DFF2CF6903C8ABB9A" bold="true" box="[151,657,914,940]" pageId="5" pageNumber="53">
<emphasis id="B9113D09F301342DFF2CF6903FE8BB9A" bold="true" box="[151,499,914,940]" italics="true" pageId="5" pageNumber="53">Augiades bouddha niitakana</emphasis>
Sonan, 1936
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDAE11BF301342DFF2CF6B73F4CBBF9" blockId="5.[151,657,914,975]" box="[151,343,949,975]" pageId="5" pageNumber="53">Zephyrus 6:213.</paragraph>
</subSubSection>
<subSubSection id="C37FB290F301342DFF2CF1003B1EBE30" pageId="5" pageNumber="53" type="description">
<paragraph id="8BDAE11BF301342DFF2CF1003FB1BC0F" blockId="5.[151,1436,1026,1081]" pageId="5" pageNumber="53">
Labels: “
<taxonomicName id="4C659A98F301342DFEBBF1003F7FBC2F" box="[256,356,1026,1049]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="genus">Augiades</taxonomicName>
/
<taxonomicName id="4C659A98F301342DFECFF1003FD4BC2F" box="[372,463,1026,1049]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="species" species="bouddha">bouddha</taxonomicName>
/ niitakana Sonan/ DET. J. SONAN,” “ 800- SUIGEN/
<date id="FFDBC7DBF301342DFB9FF1003ABCBC2F" box="[1060,1191,1026,1049]" pageId="5" pageNumber="53" value="1930-07-24">24 VII 1930</date>
/ Col. J. Sonan,” “ No. 60,” “
<typeStatus id="54DE5FB9F301342DFEB6F1203F59BC0F" box="[269,322,1058,1081]" pageId="5" pageNumber="53">Type</typeStatus>
[in red].”
</paragraph>
<paragraph id="8BDAE11BF301342DFF2CF16B3B1EBE30" blockId="5.[151,1437,1129,1542]" pageId="5" pageNumber="53">
A male specimen was designated as the
<typeStatus id="54DE5FB9F301342DFDCCF16B3CC5BCB5" box="[631,734,1129,1155]" pageId="5" pageNumber="53" type="holotype">holotype</typeStatus>
by Sonan (1936), and this
<typeStatus id="54DE5FB9F301342DFB98F16B3A91BCB5" box="[1059,1162,1129,1155]" pageId="5" pageNumber="53" type="holotype">holotype</typeStatus>
was retrieved in TARI (
<figureCitation id="135EFD9EF301342DFF1BF1923F2BBC9C" box="[160,304,1168,1194]" captionStart="FIGURES 1 18" captionStartId="2.[151,273,1746,1770]" captionTargetBox="[204,1392,187,1723]" captionTargetId="figure@2.[183,1411,179,1723]" captionTargetPageId="2" captionText="FIGURES 1 18. Types of skipper taxa described by Jinhaku Sonan: 1 3, holotype of Lobocla kodairai, upperside, underside, labels; 4 6, holotype of Coladenia sadakoe, upperside, underside, labels; 7 9, holotype of Tagiades menaka kotoshona, upperside, underside, labels; 10 12, holotype of Notocrypta arisana, upperside, underside, labels; 13 15, holotype of Ampittia maro matsumurai, upperside, underside, labels; 16 18. holotype of Augiades bouddha niitakana, upperside, underside, labels." httpUri="https://zenodo.org/record/189655/files/figure.png" pageId="5" pageNumber="53">Fig. 1618</figureCitation>
).
<bibRefCitation id="EFF49CEAF301342DFEFCF1923FF4BC9C" author="Evans" box="[327,495,1168,1194]" pageId="5" pageNumber="58" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History). The British Museum (Natural History), London, xix + 502 pp." type="book" year="1949">Evans (1949)</bibRefCitation>
treated this taxon as a subspecies of
<taxonomicName id="4C659A98F301342DFC71F1923B31BC9C" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[970,1322,1168,1194]" class="Insecta" family="Hesperiidae" genus="Ochlodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="B9113D09F301342DFC71F1923A67BC9F" box="[970,1148,1168,1193]" italics="true" pageId="5" pageNumber="53">Ochlodes siva</emphasis>
Moore, 1878
</taxonomicName>
, and this treatment was followed by
<bibRefCitation id="EFF49CEAF301342DFE5FF1B43C84BCE6" author="Bridges" box="[484,671,1206,1232]" pageId="5" pageNumber="57" refString="Bridges, C. A. (1983) Lepidoptera: Hesperiidae, Notes on Species-Group Names. Published by author, Urbana, 274 pp." type="book" year="1983">Bridges (1983)</bibRefCitation>
.
<bibRefCitation id="EFF49CEAF301342DFD10F1B43D7DBCE6" author="Shirozu" box="[683,870,1206,1232]" pageId="5" pageNumber="58" refString="Shirozu, T. (1960) Butterflies of Formosa in Colour. Hoikusha, Osaka, 481 pp. [In Japanese]" type="book" year="1960">Shirôzu (1960)</bibRefCitation>
, however, considered this taxon a synonym of
<taxonomicName id="4C659A98F301342DFF2CF1DF3F6BBCC0" box="[151,368,1245,1270]" class="Insecta" family="Hesperiidae" genus="Augiades" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="species" species="sylvanus">
<emphasis id="B9113D09F301342DFF2CF1DF3F6BBCC0" box="[151,368,1245,1270]" italics="true" pageId="5" pageNumber="53">Augiades sylvanus</emphasis>
</taxonomicName>
<taxonomicName id="4C659A98F301342DFECCF1DF3CE4BCC1" authority="Matsumura, 1919" authorityName="Matsumura" authorityYear="1919" box="[375,767,1245,1271]" pageId="5" pageNumber="53" rank="variety" variety="formosana">
var.
<emphasis id="B9113D09F301342DFE11F1DF3C3CBCC0" box="[426,551,1245,1270]" italics="true" pageId="5" pageNumber="53">formosana</emphasis>
Matsumura, 1919
</taxonomicName>
, which was recognized to represent a valid subspecies of
<taxonomicName id="4C659A98F301342DFF2CF0063CB3BD28" authority="Bremer &amp; Grey, 1853" authorityName="Bremer &amp; Grey" authorityYear="1853" box="[151,680,1284,1310]" class="Insecta" family="Hesperiidae" genus="Ochlodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="species" species="subhyalina">
<emphasis id="B9113D09F301342DFF2CF0063F95BD2B" box="[151,398,1284,1309]" italics="true" pageId="5" pageNumber="53">Ochlodes subhyalina</emphasis>
(Bremer &amp; Grey, 1853)
</taxonomicName>
by
<bibRefCitation id="EFF49CEAF301342DFD6EF0063D6DBD28" author="Evans" box="[725,886,1284,1310]" pageId="5" pageNumber="58" refString="Evans, W. H. (1949) A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum (Natural History). The British Museum (Natural History), London, xix + 502 pp." type="book" year="1949">Evans (1949)</bibRefCitation>
. Based on the characteristics of male genitalia,
<bibRefCitation id="EFF49CEAF301342DFF2CF0283FE3BD72" author="Chiba" box="[151,504,1322,1348]" lastPageId="5" lastPageNumber="58" pageId="5" pageNumber="57" refString="Chiba, H &amp; Tsukiyama, H. (1996) A review of the genus Ochlodes Scudder, 1872, with special reference to the Eurasian species (Lepidoptera: Hesperiidae). Butterflies, 14, 3 - 16. [In Japanese with English abstract]" type="journal article" year="1996">Chiba and Tsukiyama (1996)</bibRefCitation>
subsequently raised
<emphasis id="B9113D09F301342DFD4CF0293D63BD72" box="[759,888,1323,1348]" italics="true" pageId="5" pageNumber="53">formosana</emphasis>
to species status, and considered it a species endemic to
<collectingCountry id="F372A18BF301342DFE98F0533F65BD5D" box="[291,382,1361,1387]" name="Taiwan" pageId="5" pageNumber="53">Taiwan</collectingCountry>
.
<bibRefCitation id="EFF49CEAF301342DFE32F0533C43BD5D" box="[393,600,1361,1387]" pageId="5" pageNumber="58" refString="Hsu, Y. F., Chiba, H., Yang, Y. C., Tsukiyama, H., Yang, P. S. &amp; Wang S. M. (2006) A Taxonomic review of the Taiwanese skippers described by Shonen Matsumura (Lepidoptera: Hesperiidae). Zoological Studies, 45, 322 - 338." type="journal article">
Hsu
<emphasis id="B9113D09F301342DFE7AF0533FE5BD5C" box="[449,510,1361,1386]" italics="true" pageId="5" pageNumber="53">et al.</emphasis>
(2006)
</bibRefCitation>
, however, found that
<emphasis id="B9113D09F301342DFCE7F0533DC0BD5C" box="[860,987,1361,1386]" italics="true" pageId="5" pageNumber="53">formosana</emphasis>
Matsumura, 1919 is actually a junior subjective synonym of
<taxonomicName id="4C659A98F301342DFE04F07A3DDCBDA4" authority="Bremer &amp; Grey, 1853" authorityName="Bremer &amp; Grey" authorityYear="1853" box="[447,967,1400,1426]" class="Insecta" family="Hesperiidae" genus="Ochlodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="species" species="venata">
<emphasis id="B9113D09F301342DFE04F07A3C89BDA7" box="[447,658,1400,1425]" italics="true" pageId="5" pageNumber="53">Ochlodes venata</emphasis>
(Bremer &amp; Grey, 1853)
</taxonomicName>
, so
<emphasis id="B9113D09F301342DFC40F07A3A69BDA7" box="[1019,1138,1400,1425]" italics="true" pageId="5" pageNumber="53">niitakana</emphasis>
Sonan, 1936 should be resurrected in the combination
<taxonomicName id="4C659A98F301342DFDA1F09D3D0CBD8E" box="[538,791,1439,1464]" class="Insecta" family="Hesperiidae" genus="Ochlodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="53" phylum="Arthropoda" rank="species" species="niitakanus">
<emphasis id="B9113D09F301342DFDA1F09D3D0CBD8E" box="[538,791,1439,1464]" italics="true" pageId="5" pageNumber="53">Ochlodes niitakanus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF49CEAF301342DFC93F09C3DF9BD8E" author="Shirozu" box="[808,994,1438,1464]" pageId="5" pageNumber="58" refString="Shirozu, T. (1960) Butterflies of Formosa in Colour. Hoikusha, Osaka, 481 pp. [In Japanese]" type="book" year="1960">Shirôzu (1960)</bibRefCitation>
claimed that the allotype female of
<emphasis id="B9113D09F301342DFF2CF0C73F13BDE8" box="[151,264,1477,1502]" italics="true" pageId="5" pageNumber="53">niitakana</emphasis>
was probably a different species. His statement seems to be groundless. We examined the female
<typeStatus id="54DE5FB9F301342DFF2CF0EE3EE7BE30" box="[151,252,1516,1542]" pageId="5" pageNumber="53" type="paratype">paratype</typeStatus>
of
<emphasis id="B9113D09F301342DFE98F0EE3F89BE33" box="[291,402,1516,1541]" italics="true" pageId="5" pageNumber="53">niitakana</emphasis>
(
<figureCitation id="135EFD9EF301342DFE1AF0EE3C38BE30" box="[417,547,1516,1542]" captionStart="FIGURES 19 39" captionStartId="4.[151,270,1872,1896]" captionTargetBox="[192,1389,202,1833]" captionTargetId="figure@4.[189,1401,197,1835]" captionTargetPageId="4" captionText="FIGURES 19 39. Types of skipper taxa described by Jinhaku Sonan: 19 21, paratype of Augiades bouddha niitakana, upperside, underside, labels; 22 24, holotype of Parnara kotoshona, upperside, underside, labels; 25 27, holotype of Parnara ranrunna, upperside, underside, labels; 28 30, holotype of Pamara kiraizana, upperside, underside, labels; 31 33, holotype of Gangara thyrsis hainana, upperside, underside, labels; 34 36, holotype of Telicota palmarum hainanum, upperside, upperside, labels; 37 39, holotype of Tagiades menaka hainana, upperside, underside, labels." httpUri="https://zenodo.org/record/189656/files/figure.png" pageId="5" pageNumber="53">Fig. 1921</figureCitation>
) and confirmed that it was conspecific with the male
<typeStatus id="54DE5FB9F301342DFB2CF0EE3B1ABE30" box="[1175,1281,1516,1542]" pageId="5" pageNumber="53" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
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