587 lines
107 KiB
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587 lines
107 KiB
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<document id="BF8021C060013E1EDC54C5D31BBAB00E" ID-CLB-Dataset="305526" ID-DOI="10.5852/cr-palevol2023v22a34" ID-GBIF-Dataset="f8720f20-84eb-40c0-9d37-ae1c065df2bd" ID-ISSN="1777-571X" ID-Zenodo-Dep="14224371" ID-ZooBank="urn:lsid:zoobank.org:pub:69E324CE-1990-46CA-92A5-FFD83FF41DD8" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1732633430085" checkinUser="felipe" docAuthor="Cadena, Edwin-Alberto, Link, Andrés, Vanegas, Andrés, Avellaneda-Otero, Carlos A., Perdomo, César, Urueña-Carrillo, Diego A., Sánchez, Rodolfo & Vanegas, Rubén" docDate="2023" docId="03CAF60EFFB7B34AFEE736BF41BA1C76" docLanguage="en" docName="CRPalevol.22.34.689-709.pdf" docOrigin="Comptes Rendus Palevol 22 (34)" docSource="http://dx.doi.org/10.5852/cr-palevol2023v22a34" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Chelus colombiana Wood 1976" docType="treatment" docVersion="2" lastPageNumber="702" masterDocId="FFF38E76FFBCB345FFF1354B402B1A5B" masterDocTitle="New insights into the fossil record of the turtle genus Chelus Duméril, 1806 including new specimens with information on cervicals and limb bones" masterLastPageNumber="709" masterPageNumber="689" pageNumber="698" updateTime="1732751168290" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="16F35D14687A3FF9D9D8AB14F3814FAF">New insights into the fossil record of the turtle genus Chelus Duméril, 1806 including new specimens with information on cervicals and limb bones</mods:title>
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<mods:namePart id="B2A8187478080FEEA60DF1F0A0B97DB5">Cadena, Edwin-Alberto</mods:namePart>
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<mods:affiliation id="BFF24034B2F7283D6DB5B1DF8F08B022">Universidad del Rosario, Facultad de Ciencias Naturales, Grupo de Investigación Paleontología Neotropical Tradicional y Molecular (PaleoNeo), Cra 26 63 C- 69, 111211, Bogotá (Colombia) and Smithsonian Tropical Research Institute, Box 0843 - 03092, Balboa-Ancón, Panama City (Panama) and Field Museum of Natural History, 1400 S DuSable Lake Shore Dr, 60605, Chicago (United States of America) edwin. cadena @ urosario. edu. co (corresponding author)</mods:affiliation>
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<mods:namePart id="FD5251F29C9CA756A33781DC34A52396">Link, Andrés</mods:namePart>
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<mods:affiliation id="B5F1E841F0D53147E0F4D4246D0640D2">Departamento de Ciencias Biológicas, Universidad de Los Andes, Cra 1 18 A- 12, 111711, Bogotá (Colombia) a. link 74 @ uniandes. edu. co</mods:affiliation>
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<mods:affiliation id="D9ECD40F378E556B88707BFA1477BFE4">Museo de Historia Natural La Tatacoa, 411028, La Victoria (Colombia) andresvanegasv @ gmail. com</mods:affiliation>
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<mods:namePart id="2075CBC0085A11C71E186D2DA45B5D3F">Avellaneda-Otero, Carlos A.</mods:namePart>
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<mods:affiliation id="97922D326164B5B88BF13CA69DD84002">Universidad del Rosario, Facultad de Ciencias Naturales, Grupo de Investigación Paleontología Neotropical Tradicional y Molecular (PaleoNeo), Cra 26 63 C- 69, 111211, Bogotá (Colombia) caavellanedao @ unal. edu. co</mods:affiliation>
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<mods:affiliation id="7CD772C34C84DC6F8E00A750BB8E71E1">Museo de Paleontología La Tormenta, Desierto Tatacoa, 411028, Villavieja (Colombia) mezondelatatacoa @ gmail. com</mods:affiliation>
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<mods:namePart id="E1EFCE7C41B614EB131EF58016001397">Urueña-Carrillo, Diego A.</mods:namePart>
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<mods:affiliation id="E4CCD8C41BE1EE01142972F5D9ACADED">Museo de Historia Natural La Tatacoa, 411028, La Victoria (Colombia) dauac 2003 @ gmail. com</mods:affiliation>
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<mods:namePart id="3E98C9E6EC647F2DB72E61B2E1F40583">Sánchez, Rodolfo</mods:namePart>
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<mods:affiliation id="4AD3321F711D78C739D9C43642FB53E3">Museo Paleontológico de Urumaco, Alcaldía de Urumaco, 5 PXW + 98, 4141, Urumaco (Venezuela) rodolfosanchez 128 @ gmail. com</mods:affiliation>
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<mods:namePart id="F490CB8A10D01D01516ED900DB151EEF">Vanegas, Rubén</mods:namePart>
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<mods:affiliation id="A9368C343A71E26940E32BF396476F61">Museo de Historia Natural La Tatacoa, 411028, La Victoria (Colombia) rubendario 2496 @ gmail. com</mods:affiliation>
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<mods:title id="9044DCE92660C7D5A59A7B9896DD282A">Comptes Rendus Palevol</mods:title>
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<treatment id="03CAF60EFFB7B34AFEE736BF41BA1C76" ID-DOI="http://doi.org/10.5281/zenodo.14232279" ID-GBIF-Taxon="247601468" ID-Zenodo-Dep="14232279" LSID="urn:lsid:plazi:treatment:03CAF60EFFB7B34AFEE736BF41BA1C76" httpUri="http://treatment.plazi.org/id/03CAF60EFFB7B34AFEE736BF41BA1C76" lastPageId="15" lastPageNumber="702" pageId="11" pageNumber="698" scope_class="Testudines" scope_family="Chelidae" scope_order="Testudines">
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<taxonomicName id="4C633C9BFFB7B34EFEE736BF425E1E54" authority="Wood, 1976" authorityName="Wood" authorityYear="1976" box="[278,629,1012,1039]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="colombiana">
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<emphasis id="B9179B0AFFB7B34EFEE736BF41CF1E54" bold="true" box="[278,484,1012,1039]" italics="true" pageId="11" pageNumber="698">Chelus colombiana</emphasis>
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<bibRefCitation id="EFF23AE9FFB7B34EFE1B36BE425E1E54" author="WOOD R. C." box="[490,629,1013,1039]" pageId="11" pageNumber="698" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood, 1976</bibRefCitation>
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<paragraph id="8BDC4718FFB7B34EFE60315E41D11E74" blockId="11.[401,506,1045,1072]" box="[401,506,1045,1072]" pageId="11" pageNumber="698">
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(
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<figureCitation id="13585B9DFFB7B34EFE6B315E41DA1E74" box="[410,497,1045,1072]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="12.[132,143,1372,1389]" captionStartId-1="13.[134,143,1374,1391]" captionStartId-2="14.[131,142,1602,1619]" captionStartId-3="16.[132,143,1825,1842]" captionTargetBox-0="[135,1455,217,1324]" captionTargetBox-1="[153,1457,219,1327]" captionTargetBox-2="[151,1441,215,1560]" captionTargetBox-3="[132,1445,217,1796]" captionTargetId-0="figure-498@12.[698,888,218,1331]" captionTargetId-1="figure-500@13.[697,1266,223,787]" captionTargetId-2="figure-288@14.[687,1256,789,1366]" captionTargetId-3="figure-294@16.[698,1267,702,1159]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="16" captionText-0="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." captionText-1="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." captionText-2="FIG. 7. — Complete plastra of Chelus colombiana Wood, 1976 VPPLT-380 and MT-22 from the Tatacoa of Colombia: A, B, VPPLT-380 specimen in dorsal view; C, D, VPPLT-380 specimen in ventral view; E, F, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Ext II, extragular scute II; Fem, femoral scute; Gul, gular scute; isc, ischium scar; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; Pec, pectoral scute; psc, pubis scar; xip, xiphiplastron. Scale bar: 20 cm." captionText-3="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="11" pageNumber="698">Figs 5-8</figureCitation>
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)
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<treatmentCitation id="0AC26109FFB7B34EFF75311941F91E31" author="WOOD R. C." box="[132,466,1106,1130]" page="3" pageId="11" pageNumber="698" year="1976">
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<taxonomicName id="4C633C9BFFB7B34EFF75311941F91E31" authority="Wood, 1976: 3" authorityName="Wood" authorityPageNumber="3" authorityYear="1976" box="[132,466,1106,1130]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="colombiana">
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<emphasis id="B9179B0AFFB7B34EFF753119411E1E31" box="[132,309,1106,1130]" italics="true" pageId="11" pageNumber="698">Chelus colombiana</emphasis>
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<bibRefCitation id="EFF23AE9FFB7B34EFECA311941F91E31" author="WOOD R. C." box="[315,466,1106,1130]" pageId="11" pageNumber="698" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood, 1976: 3</bibRefCitation>
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</subSubSection>
|
|
<subSubSection id="C3791493FFB7B34EFF7531CC42951FD7" pageId="11" pageNumber="698" type="materials_examined">
|
|
<paragraph id="8BDC4718FFB7B34EFF7531CC41121EE2" blockId="11.[132,774,1158,1209]" pageId="11" pageNumber="698">
|
|
<materialsCitation id="3B0B4D45FFB7B34EFF7531CC41121EE2" ID-GBIF-Occurrence="4985334323" collectionCode="UCMP" pageId="11" pageNumber="698" specimenCode="UCMP-78762" specimenCount="1" typeStatus="type">
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFF7531CC409D1EC7" baselines="1176,1176" box="[132,182,1159,1182]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="11" pageNumber="698">
|
|
<typeStatus id="54D8F9BAFFB7B34EFF7531CC409D1EC7" box="[132,182,1159,1182]" pageId="11" pageNumber="698">TYPE</typeStatus>
|
|
</smallCapsWord>
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFF4E31C7410B1EC6" baselines="1177" box="[191,288,1164,1181]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="specimen" pageId="11" pageNumber="698">SPECIMEN</smallCapsWord>
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|
. —
|
|
<specimenCode id="DBC5EF63FFB7B34EFEA331CD41C11EC5" box="[338,490,1158,1182]" pageId="11" pageNumber="698">UCMP-78762</specimenCode>
|
|
, an articulated shell (
|
|
<bibRefCitation id="EFF23AE9FFB7B34EFD3831CD40931EE2" author="WOOD R. C." pageId="11" pageNumber="698" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood 1976</bibRefCitation>
|
|
) (
|
|
<figureCitation id="13585B9DFFB7B34EFF3C31EA41191EE2" box="[205,306,1185,1209]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5A, B</figureCitation>
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)
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB7B34EFF75319D42951FD7" blockId="11.[132,777,1237,1420]" pageId="11" pageNumber="698">
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFF75319D40C01EB0" baselines="1255,1255" box="[132,235,1238,1261]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Referred" pageId="11" pageNumber="698">REFERRED</smallCapsWord>
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<smallCapsWord id="8D3AD1C4FFB7B34EFF053190417D1EB7" baselines="1256" box="[244,342,1243,1260]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="11" pageNumber="698">MATERIAL</smallCapsWord>
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. —
|
|
<materialsCitation id="3B0B4D45FFB7B34EFE7B319E41D31F66" ID-GBIF-Occurrence="4985334333" pageId="11" pageNumber="698" specimenCode="IGMp-002045" specimenCount="1">
|
|
Specimens referred by
|
|
<bibRefCitation id="EFF23AE9FFB7B34EFD87319E432B1EB6" author="WOOD R. C." box="[630,768,1237,1261]" pageId="11" pageNumber="698" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood (1976)</bibRefCitation>
|
|
:
|
|
<specimenCode id="DBC5EF63FFB7B34EFF7531A441311F5C" box="[132,282,1263,1287]" pageId="11" pageNumber="698">IGMp-002045</specimenCode>
|
|
, incomplete shell lacking part of the right side of the carapace and anterior plastral lobe, possibly a male (
|
|
<bibRefCitation id="EFF23AE9FFB7B34EFD41304141641F67" author="SANCHEZ-VILLAGRA M. R. & LINARES O. J. & PAOLILLO A." pageId="11" pageNumber="698" pagination="159 - 167" refId="ref13381" refString="SANCHEZ-VILLAGRA M. R., LINARES O. J. & PAOLILLO A. 1995 a. - Consideraciones sobre la sistematica de las tortugas del genero Chelus (Pleurodira: Chelidae) y nuevas evidencias fosiles del Mioceno de Colombia y Venezuela. Ameghiniana 32: 159 - 167." type="journal article" year="1995">
|
|
Sánchez-Villagra
|
|
<emphasis id="B9179B0AFFB7B34EFF28306E41211F67" box="[217,266,1316,1340]" italics="true" pageId="11" pageNumber="698">et al.</emphasis>
|
|
1995a
|
|
</bibRefCitation>
|
|
: fig. 1A;
|
|
<figureCitation id="13585B9DFFB7B34EFE40306F41C11F67" box="[433,490,1316,1340]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="11" pageNumber="698">Fig. 6</figureCitation>
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|
);
|
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</materialsCitation>
|
|
<materialsCitation id="3B0B4D45FFB7B34EFDF1306E41681F0C" ID-GBIF-Occurrence="4985334343" pageId="11" pageNumber="698" specimenCode="IGMp-002049" specimenCount="1">
|
|
<specimenCode id="DBC5EF63FFB7B34EFDF1306E42B31F66" box="[512,664,1317,1341]" pageId="11" pageNumber="698">IGMp-002049</specimenCode>
|
|
, a partially disarticulated shell;
|
|
</materialsCitation>
|
|
<materialsCitation id="3B0B4D45FFB7B34EFEB8307442BD1F0C" ID-GBIF-Occurrence="4985334315" box="[329,662,1343,1367]" pageId="11" pageNumber="698" specimenCode="IGMp-002085" specimenCount="1">
|
|
<specimenCode id="DBC5EF63FFB7B34EFEB8307441CB1F0C" box="[329,480,1343,1367]" pageId="11" pageNumber="698">IGMp-002085</specimenCode>
|
|
, a left epiplastron.
|
|
</materialsCitation>
|
|
<materialsCitation id="3B0B4D45FFB7B34EFD6D307442AD1F29" ID-GBIF-Occurrence="4985334341" collectionCode="VPPLT" pageId="11" pageNumber="698" specimenCode="VPPLT-1748" specimenCount="1">
|
|
This study:
|
|
<specimenCode id="DBC5EF63FFB7B34EFF74301141221F29" box="[133,265,1370,1394]" pageId="11" pageNumber="698">VPPLT-1748</specimenCode>
|
|
, nearly complete carapace (
|
|
<figureCitation id="13585B9DFFB7B34EFDE2301242601F2A" box="[531,587,1369,1393]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5</figureCitation>
|
|
C-G),
|
|
</materialsCitation>
|
|
<materialsCitation id="3B0B4D45FFB7B34EFD7D301141761FD7" ID-GBIF-Occurrence="4985334329" collectionCode="VPPLT" pageId="11" pageNumber="698" specimenCode="VPPLT-380" specimenCount="1">
|
|
<specimenCode id="DBC5EF63FFB7B34EFD7D301143281F29" box="[652,771,1370,1394]" pageId="11" pageNumber="698">VPPLT-380</specimenCode>
|
|
, complete plastron and
|
|
</materialsCitation>
|
|
<materialsCitation id="3B0B4D45FFB7B34EFE95303F42921FD7" ID-GBIF-Occurrence="4985334337" box="[356,697,1396,1420]" collectionCode="MT" pageId="11" pageNumber="698" specimenCode="MT-22" specimenCount="1">
|
|
<specimenCode id="DBC5EF63FFB7B34EFE95303F41861FD7" box="[356,429,1396,1420]" pageId="11" pageNumber="698">MT-22</specimenCode>
|
|
, complete shell (
|
|
<figureCitation id="13585B9DFFB7B34EFDA0303F42A11FD7" box="[593,650,1396,1420]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5</figureCitation>
|
|
H-J)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3791493FFB7B34EFF7530E341CA1FAF" pageId="11" pageNumber="698" type="distribution">
|
|
<paragraph id="8BDC4718FFB7B34EFF7530E341CA1FAF" blockId="11.[132,775,1448,1525]" pageId="11" pageNumber="698">
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFF7530E3413E1FE4" baselines="1466,1467" box="[132,277,1448,1472]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Occurrences" pageId="11" pageNumber="698">OCCURRENCES</smallCapsWord>
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|
<smallCapsWord id="8D3AD1C4FFB7B34EFEEA30E5416E1FE4" baselines="1467" box="[283,325,1454,1471]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="11" pageNumber="698">AND</smallCapsWord>
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFEBB30E541441FE4" baselines="1467" box="[330,367,1454,1471]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="age" pageId="11" pageNumber="698">AGE</smallCapsWord>
|
|
. — Villavieja and La Victoria formations,
|
|
<collectingRegion id="49A789FAFFB7B34EFF753089416D1F81" box="[132,326,1474,1498]" country="Colombia" name="Huila" pageId="11" pageNumber="698">Huila Department</collectingRegion>
|
|
,
|
|
<collectingCountry id="F3740788FFB7B34EFEA2308941961F81" box="[339,445,1474,1498]" name="Colombia" pageId="11" pageNumber="698">Colombia</collectingCountry>
|
|
. Middle to Late Miocene 16 to 10.52 Ma (
|
|
<bibRefCitation id="EFF23AE9FFB7B34EFF02309641FE1FAE" author="MORA-ROJAS L. & CARDENAS A. & JARAMILLO C. & SILVESTRO D. & BAYONA G. & ZAPATA S. & MORENO F. & SILVA C. & MORENO-BERNAL J. & JARAMILLO J. S. & VALENCIA V. & IBANEZ M." box="[243,469,1501,1525]" pageId="11" pageNumber="698" pagination="197 - 221" refId="ref13006" refString="MORA-ROJAS L., CARDENAS A., JARAMILLO C., SILVESTRO D., BAYONA G., ZAPATA S., MORENO F., SILVA C., MORENO-BERNAL J., JARAMILLO J. S., VALENCIA V. & IBANEZ M. 2023. - Stratigraphy of a middle Miocene Neotropical Lagerstatte (La Venta Site). Geodiversitas 45 (6): 197 - 221. https: // doi. org / 10.5252 / geodiversitas 2023 v 45 a 6" type="journal article" year="2023">
|
|
Mora-Rojas
|
|
<emphasis id="B9179B0AFFB7B34EFE9D309541B01FAE" box="[364,411,1501,1525]" italics="true" pageId="11" pageNumber="698">et al.</emphasis>
|
|
2023
|
|
</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3791493FFB7B34EFF753359417D1DB0" pageId="11" pageNumber="698" type="diagnosis">
|
|
<paragraph id="8BDC4718FFB7B34EFF753359417D1DB0" blockId="11.[131,777,1553,2027]" pageId="11" pageNumber="698">
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFF75335940FC1C73" baselines="1571,1572" box="[132,215,1554,1577]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Revised" pageId="11" pageNumber="698">REVISED</smallCapsWord>
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFF2A335C416A1C73" baselines="1572" box="[219,321,1559,1576]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="11" pageNumber="698">DIAGNOSIS</smallCapsWord>
|
|
. —
|
|
<taxonomicName id="4C633C9BFFB7B34EFE9B335A42321C72" authorityName="Wood" authorityYear="1976" box="[362,537,1553,1577]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="colombiana">
|
|
<emphasis id="B9179B0AFFB7B34EFE9B335A42321C72" box="[362,537,1553,1577]" italics="true" pageId="11" pageNumber="698">Chelus colombiana</emphasis>
|
|
</taxonomicName>
|
|
differs from all other extant and fossil
|
|
<taxonomicName id="4C633C9BFFB7B34EFEFD336741611C1F" authorityName="Dumeril" authorityYear="1806" box="[268,330,1580,1604]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="genus">
|
|
<emphasis id="B9179B0AFFB7B34EFEFD336741611C1F" box="[268,330,1580,1604]" italics="true" pageId="11" pageNumber="698">Chelus</emphasis>
|
|
</taxonomicName>
|
|
by having narrow knobs that form continuous dorsal ridges on the carapace that do not expand laterally, a feature that is present independently of size. It shares with
|
|
<taxonomicName id="4C633C9BFFB7B34EFD85332A42C21C23" authorityName="Schneider" authorityYear="1783" box="[628,745,1632,1656]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="fimbriata">
|
|
<emphasis id="B9179B0AFFB7B34EFD85332A42AF1C23" box="[628,644,1633,1656]" italics="true" pageId="11" pageNumber="698">C</emphasis>
|
|
.
|
|
<emphasis id="B9179B0AFFB7B34EFD60332B42C21C23" box="[657,745,1632,1656]" italics="true" pageId="11" pageNumber="698">fimbriata</emphasis>
|
|
</taxonomicName>
|
|
an almost rectangular carapace, with parallel lateral margins.
|
|
<taxonomicName id="4C633C9BFFB7B34EFD38333040DB1CF6" authorityName="Wood" authorityYear="1976" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="colombiana">
|
|
<emphasis id="B9179B0AFFB7B34EFD38333040DB1CF6" italics="true" pageId="11" pageNumber="698">Chelus colombiana</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C633C9BFFB7B34EFED333DE41B01CF6" authorityName="Wood" authorityYear="1976" box="[290,411,1685,1709]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="lewisi">
|
|
<emphasis id="B9179B0AFFB7B34EFED333DE41B01CF6" box="[290,411,1685,1709]" italics="true" pageId="11" pageNumber="698">Chelus lewisi</emphasis>
|
|
</taxonomicName>
|
|
share and differ from the extant taxa by the presence of an axillary buttress that extends onto costal 2, instead of being restricted to costal 1 (extant taxa); inguinal buttress onto costal 5, instead of costal 4 (extant taxa); two or three pairs of extragular scutes that encapsulate the gular, instead of gular reaching the anterior margin of the plastron and a single pair of extragular scutes (extant taxa); vertebral scute 1 almost rectangular, instead of pentagonal (extant); and vertebral scute 1 almost same width of vertebral scute 2, instead vertebral 1 wider than 2 (extant). It shares with the two extant taxa a wider anterior plastral lobe than
|
|
<taxonomicName id="4C633C9BFFB7B34EFD4132CF432A1DC0" authorityName="Wood" authorityYear="1976" box="[688,769,1923,1947]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="lewisi">
|
|
<emphasis id="B9179B0AFFB7B34EFD4132CF42EB1DC0" box="[688,704,1924,1947]" italics="true" pageId="11" pageNumber="698">C</emphasis>
|
|
.
|
|
<emphasis id="B9179B0AFFB7B34EFD3C32C8432A1DC0" box="[717,769,1923,1947]" italics="true" pageId="11" pageNumber="698">lewisi</emphasis>
|
|
</taxonomicName>
|
|
. It differs from
|
|
<taxonomicName id="4C633C9BFFB7B34EFEE532D541A21DED" authorityName="Schneider" authorityYear="1783" box="[276,393,1950,1974]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="698" phylum="Chordata" rank="species" species="fimbriata">
|
|
<emphasis id="B9179B0AFFB7B34EFEE532D541A21DED" box="[276,393,1950,1974]" italics="true" pageId="11" pageNumber="698">C. fimbriata</emphasis>
|
|
</taxonomicName>
|
|
by cervicals 7 and 8 having longer and more dorsoventrally projected neural spines, and broader internal scapular angle of 85°.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3791493FFB7B34AFCDC359341BA1C76" lastPageId="15" lastPageNumber="702" pageId="11" pageNumber="698" type="description">
|
|
<paragraph id="8BDC4718FFB7B34EFCDC359343E31AAB" blockId="11.[811,1458,216,2028]" box="[813,968,216,242]" pageId="11" pageNumber="698">
|
|
<smallCapsWord id="8D3AD1C4FFB7B34EFCDC359343E31AAB" baselines="235,235" box="[813,968,216,242]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Descriptions" pageId="11" pageNumber="698">DESCRIPTIONS</smallCapsWord>
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB7B34EFCDC35B3448B1B69" blockId="11.[811,1458,216,2028]" pageId="11" pageNumber="698">
|
|
Measurements for the new and all other previously described specimens are provided in
|
|
<tableCitation id="C6E172A3FFB7B34EFBBB345C44B21B69" box="[1098,1177,279,306]" captionStart="TABLE" captionStartId="7.[131,142,219,236]" captionTargetPageId="7" captionText="TABLE 1. — Measurements of the new fossil specimens of Chelus colombiana Wood,1976 and C. lewisi Wood,1976 describe here,as well as some extant individuals of C. fimbriata (Schneider,1783) and C. orinocensis Vargas-Ramírez,Caballero,Morales-Betancourt,Lasso,Amaya,Martînez,Silva-Viana,Vogt,Pires-Farias, Hrbek, Campbell & Fritz, 2020 figured in Figure 9, in millimeters (mm). Abbreviations: CL, carapace length at midline as preserved; CML, carapace maximum length as preserved; CW, carapace width at midline as preserved; PL, plastron length at midline as preserved; PW, plastron width at midline as preserved." httpUri="http://table.plazi.org/id/DF1C1790FFBBB342FF72359045A71B6B" pageId="11" pageNumber="698" tableUuid="DF1C1790FFBBB342FF72359045A71B6B">Table 1</tableCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB7B34EFCB2347345BA1F75" blockId="11.[811,1458,216,2028]" pageId="11" pageNumber="698">
|
|
VPPLT-1748 is a nearly complete carapace, missing most of its left and posterior margins (
|
|
<figureCitation id="13585B9DFFB7B34EFB64341C45251B2A" box="[1173,1294,343,370]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5C, D</figureCitation>
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|
). The carapace is almost rectangular in shape, exhibits three dorsal ridges formed by knobs, one on the left costals, one along midline of the neurals, and one on the right costals. The knobs are narrow and do not extend radially or laterally. The surface of the bone is well preserved, allowing clear identification of sutural contacts between the bones and the sulci left by the scutes.The nuchal bone is hexagonal in shape.The neural series is composed of six bones, most of which are wider than long (except neural 4) and have a hexagonal shape. VPPLT-1748 has eight pairs of costal bones. The peripherals have a marked lateral indentation at the contact between the marginals. The sulci indicate that the cervical scute was almost rectangular in shape. Five vertebral scutes were present, of which vertebrals 1 to 4 were wider than long and rectangular to slightly hexagonal in shape, while vertebral 5 was trapezoidal. Pleural scute 1 was large, covering most of costals 1 and 2, and peripherals 1 to 3. The other pleurals were slightly smaller than pleural 1 and rectangular to trapezoidal in shape, with pleural 3 being the smallest of the four pairs. Some of the sulci between the marginals are preserved. Marginals 1 to 3 were restricted to the anterior peripherals and marginals 8 to 10 were potentially restricted to the posterior peripherals, without reaching the costals. On the ventral surface of the carapace (
|
|
<figureCitation id="13585B9DFFB7B34EFC6D311E443B1E34" box="[924,1040,1108,1135]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5E, F</figureCitation>
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|
), most of both axillary buttresses are preserved reaching costals 2. The inguinal buttress scar left is restricted to costal 5, while the iliac scar covered most of costal 8 and potentially the suprapygal. In right lateral view (
|
|
<figureCitation id="13585B9DFFB7B34EFCC7319F43A01EB5" box="[822,907,1236,1263]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5G</figureCitation>
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), the carapace exhibits a low dome shape showing the left and right costal series of knobs forming the ridges, which are narrow without extending radially or laterally.
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</paragraph>
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|
<paragraph id="8BDC4718FFB7B34EFCB2307F451B1CD6" blockId="11.[811,1458,216,2028]" pageId="11" pageNumber="698">
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MT-22 is an almost complete shell (
|
|
<figureCitation id="13585B9DFFB7B34EFB48307F44DF1F14" box="[1209,1268,1332,1359]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5</figureCitation>
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H-J). It has a rectangular shape, with the left and right lateral margins almost parallel to each other. The three ridges of knobs are narrow without expanding radially or laterally on the dorsal surface of the carapace. Some of the sutural contacts between contacts are visible. The nuchal bone is hexagonal, slightly wider than long, and its anterior margin a bit eroded. Bioerosion marks are well preserved on one of the left peripherals and some costals (
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|
<figureCitation id="13585B9DFFB7B34EFC30337844211C16" box="[961,1034,1587,1614]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5I</figureCitation>
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). In left lateral view (
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|
<figureCitation id="13585B9DFFB7B34EFB02337845161C16" box="[1267,1341,1587,1614]" captionStart="FIG" captionStartId="12.[132,143,1372,1389]" captionTargetBox="[135,1455,217,1324]" captionTargetId="figure-498@12.[698,888,218,1331]" captionTargetPageId="12" captionText="FIG. 5. — Shell and carapaces of Chelus colombiana Wood, 1976 from the Tatacoa of Colombia: A, B, UCMP-78762 holotype, articulated shell: A, shell in dorsal view; B, shell in ventral view; C-G, VPPLT-1748: C, D, carapace in dorsal view; E, F, carapace in ventral view; G, carapace in right lateral view; H-J, MT-22: H, I, carapace in dorsal view; J, carapace in left lateral view. Abbreviations: axb, axillary buttress; bim, bioerosion marks; Cer, cervical scute; co, costal bone; ilsc, iliac scar; ing, inguinal scar; lcr, left costal ridge; M, marginal scute; ne, neural bone; ner, neural ridge; P, pleural scute; rcr, right costal ridge; V, vertebral scute. Scale bar: 20 cm." pageId="11" pageNumber="698">Fig. 5J</figureCitation>
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), the carapace shows a low dome shape and the narrow and low knobs without developing peaks or extending laterally.
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</paragraph>
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<paragraph id="8BDC4718FFB7B34EFCB233D944461D37" blockId="11.[811,1458,216,2028]" pageId="11" pageNumber="698">
|
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IGMp-002045 is an articulated shell (
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|
<figureCitation id="13585B9DFFB7B34EFB1933D845031CF6" box="[1256,1320,1683,1709]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="11" pageNumber="698">Fig. 6</figureCitation>
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), preserving most of the left half of the carapace and a portion of the medial region of the right half. Most of the anterior plastral lobe and right bridge with the carapace region are missing. The missing parts of the carapace and right bridge region were reconstructed with plaster originally during the preparation of the specimen in the 1970s.
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</paragraph>
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|
<paragraph id="8BDC4718FFB7B349FCB2323945161C75" blockId="11.[811,1458,216,2028]" lastBlockId="12.[811,1457,1523,2027]" lastPageId="12" lastPageNumber="699" pageId="11" pageNumber="698">
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The carapace of IGMp-002045 (
|
|
<figureCitation id="13585B9DFFB7B34EFB51323945261DD7" box="[1184,1293,1906,1932]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="11" pageNumber="698">Fig. 6A, B</figureCitation>
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) exhibits on its dorsal surface three ridges of narrow knobs, without developing lateral or radial expansion. The sutural contacts between bones are poorly identifiable. However, the sulci left by the scutes are well-preserved. There were five vertebral scutes. Vertebral 1 was almost rectangular in shape and vertebrals 2 and 3 hexagonal and wider than long. Vertebral scute 4 was almost equally wide than long and vertebral scute 5 trapezoidal in shape. There were eleven marginal scutes and four pleural scutes, of which pleural 1 was the largest. On its ventral surface, the carapace of IGMp-002045 exhibits an axillary buttress that reaches costal 2, and a very large costovertebral tunnel (
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|
<figureCitation id="13585B9DFFB0B349FF2C33B941181D57" box="[221,307,1778,1804]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="12" pageNumber="699">Fig. 6C</figureCitation>
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). The plastron is complete and articulated with the carapace (
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|
<figureCitation id="13585B9DFFB0B349FEA4325941831D77" box="[341,424,1810,1837]" captionStart="FIG" captionStartId="14.[131,142,1602,1619]" captionTargetBox="[151,1441,215,1560]" captionTargetId="figure-288@14.[687,1256,789,1366]" captionTargetPageId="14" captionText="FIG. 7. — Complete plastra of Chelus colombiana Wood, 1976 VPPLT-380 and MT-22 from the Tatacoa of Colombia: A, B, VPPLT-380 specimen in dorsal view; C, D, VPPLT-380 specimen in ventral view; E, F, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Ext II, extragular scute II; Fem, femoral scute; Gul, gular scute; isc, ischium scar; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; Pec, pectoral scute; psc, pubis scar; xip, xiphiplastron. Scale bar: 20 cm." pageId="12" pageNumber="699">Fig. 7E</figureCitation>
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). The sutures between bones, as well as the sulci left by the scutes are clearly identifiable. The preserved sulci indicate that the femoral was longer than the abdominal and pectoral, and the anal was the shortest one of these scutes (
|
|
<figureCitation id="13585B9DFFB0B349FEDD32D941501DF7" box="[300,379,1938,1964]" captionStart="FIG" captionStartId="14.[131,142,1602,1619]" captionTargetBox="[151,1441,215,1560]" captionTargetId="figure-288@14.[687,1256,789,1366]" captionTargetPageId="14" captionText="FIG. 7. — Complete plastra of Chelus colombiana Wood, 1976 VPPLT-380 and MT-22 from the Tatacoa of Colombia: A, B, VPPLT-380 specimen in dorsal view; C, D, VPPLT-380 specimen in ventral view; E, F, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Ext II, extragular scute II; Fem, femoral scute; Gul, gular scute; isc, ischium scar; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; Pec, pectoral scute; psc, pubis scar; xip, xiphiplastron. Scale bar: 20 cm." pageId="12" pageNumber="699">Fig. 7F</figureCitation>
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). The xiphiplastra exhibit a large and deep U-shaped anal notch and long and narrow posterior tips, as well as they are characterized by a deep medial concavity. The anterior plastral lobe is wide and has evidence of two pairs of extragulars and an elongated gular scute.
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</paragraph>
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|
<caption id="DF1C1790FFB0B349FF753017417B1F93" pageId="12" pageNumber="699" startId="12.[132,143,1372,1389]" targetBox="[135,1455,217,1324]" targetPageId="12" targetType="figure">
|
|
<paragraph id="8BDC4718FFB0B349FF753017417B1F93" blockId="12.[132,1457,1372,1480]" pageId="12" pageNumber="699">
|
|
<smallCapsWord id="8D3AD1C4FFB0B349FF75301740B61F37" baselines="1385,1385" box="[132,157,1372,1389]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="12" pageNumber="699">FIG</smallCapsWord>
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|
. 5. — Shell and carapaces of
|
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<taxonomicName id="4C633C9BFFB0B349FE683017428F1F36" authority="Wood, 1976" authorityName="Wood" authorityYear="1976" box="[409,676,1372,1389]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="699" phylum="Chordata" rank="species" species="colombiana">
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<emphasis id="B9179B0AFFB0B349FE68301742131F36" box="[409,568,1372,1389]" italics="true" pageId="12" pageNumber="699">Chelus colombiana</emphasis>
|
|
<bibRefCitation id="EFF23AE9FFB0B349FDCF3017428F1F36" author="WOOD R. C." box="[574,676,1372,1389]" pageId="12" pageNumber="699" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood, 1976</bibRefCitation>
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|
</taxonomicName>
|
|
from the Tatacoa of Colombia:
|
|
<emphasis id="B9179B0AFFB0B349FC5F301743901F36" bold="true" box="[942,955,1372,1389]" pageId="12" pageNumber="699">A</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB0B349FC34301743F91F36" bold="true" box="[965,978,1372,1389]" pageId="12" pageNumber="699">B</emphasis>
|
|
, UCMP-78762 holotype, articulated shell:
|
|
<emphasis id="B9179B0AFFB0B349FAC4301745691F36" bold="true" box="[1333,1346,1372,1389]" pageId="12" pageNumber="699">A</emphasis>
|
|
, shell in dorsal view;
|
|
<emphasis id="B9179B0AFFB0B349FF20303840F51FDF" bold="true" box="[209,222,1395,1412]" pageId="12" pageNumber="699">B</emphasis>
|
|
, shell in ventral view;
|
|
<emphasis id="B9179B0AFFB0B349FE64303841931FDF" bold="true" box="[405,440,1395,1412]" pageId="12" pageNumber="699">C -G</emphasis>
|
|
, VPPLT-1748:
|
|
<emphasis id="B9179B0AFFB0B349FDC63038426E1FDF" bold="true" box="[567,581,1395,1412]" pageId="12" pageNumber="699">C</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB0B349FDBE303842761FDF" bold="true" box="[591,605,1395,1412]" pageId="12" pageNumber="699">D</emphasis>
|
|
, carapace in dorsal view;
|
|
<emphasis id="B9179B0AFFB0B349FCC6303843681FDF" bold="true" box="[823,835,1395,1412]" pageId="12" pageNumber="699">E</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB0B349FCBF303843721FDF" bold="true" box="[846,857,1395,1412]" pageId="12" pageNumber="699">F</emphasis>
|
|
, carapace in ventral view;
|
|
<emphasis id="B9179B0AFFB0B349FBC63038446E1FDF" bold="true" box="[1079,1093,1395,1412]" pageId="12" pageNumber="699">G</emphasis>
|
|
, carapace in right lateral view;
|
|
<emphasis id="B9179B0AFFB0B349FAB8303845431FDF" bold="true" box="[1353,1384,1395,1412]" pageId="12" pageNumber="699">H -J</emphasis>
|
|
, MT-22:
|
|
<emphasis id="B9179B0AFFB0B349FF7530C240B91FC1" bold="true" box="[132,146,1417,1434]" pageId="12" pageNumber="699">H</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB0B349FF6C30C240881FC1" bold="true" box="[157,163,1417,1434]" pageId="12" pageNumber="699">I</emphasis>
|
|
, carapace in dorsal view;
|
|
<emphasis id="B9179B0AFFB0B349FE8F30C241A31FC1" bold="true" box="[382,392,1417,1434]" pageId="12" pageNumber="699">J</emphasis>
|
|
, carapace in left lateral view. Abbreviations:
|
|
<emphasis id="B9179B0AFFB0B349FD0E30C243341FC1" bold="true" box="[767,799,1417,1434]" pageId="12" pageNumber="699">axb</emphasis>
|
|
, axillary buttress;
|
|
<emphasis id="B9179B0AFFB0B349FC4930C243F21FC1" bold="true" box="[952,985,1417,1434]" pageId="12" pageNumber="699">bim</emphasis>
|
|
, bioerosion marks;
|
|
<emphasis id="B9179B0AFFB0B349FB8E30C244B41FC1" bold="true" box="[1151,1183,1417,1434]" pageId="12" pageNumber="699">Cer</emphasis>
|
|
, cervical scute;
|
|
<emphasis id="B9179B0AFFB0B349FAD630C245161FC1" bold="true" box="[1319,1341,1417,1434]" pageId="12" pageNumber="699">co</emphasis>
|
|
, costal bone;
|
|
<emphasis id="B9179B0AFFB0B349FF7530EB40891FEA" bold="true" box="[132,162,1440,1457]" pageId="12" pageNumber="699">ilsc</emphasis>
|
|
, iliac scar;
|
|
<emphasis id="B9179B0AFFB0B349FEF030EB41371FEA" bold="true" box="[257,284,1440,1457]" pageId="12" pageNumber="699">ing</emphasis>
|
|
, inguinal scar;
|
|
<emphasis id="B9179B0AFFB0B349FE6B30EB419A1FEA" bold="true" box="[410,433,1440,1457]" pageId="12" pageNumber="699">lcr</emphasis>
|
|
, left costal ridge;
|
|
<emphasis id="B9179B0AFFB0B349FDB630EB42731FEA" bold="true" box="[583,600,1440,1457]" pageId="12" pageNumber="699">M</emphasis>
|
|
, marginal scute;
|
|
<emphasis id="B9179B0AFFB0B349FD1630EB42D61FEA" bold="true" box="[743,765,1440,1457]" pageId="12" pageNumber="699">ne</emphasis>
|
|
, neural bone;
|
|
<emphasis id="B9179B0AFFB0B349FC8430EB43B91FEA" bold="true" box="[885,914,1440,1457]" pageId="12" pageNumber="699">ner</emphasis>
|
|
, neural ridge;
|
|
<emphasis id="B9179B0AFFB0B349FBF830EB443E1FEA" bold="true" box="[1033,1045,1440,1457]" pageId="12" pageNumber="699">P</emphasis>
|
|
, pleural scute;
|
|
<emphasis id="B9179B0AFFB0B349FB6730EB44841FEA" bold="true" box="[1174,1199,1440,1457]" pageId="12" pageNumber="699">rcr</emphasis>
|
|
, right costal ridge;
|
|
<emphasis id="B9179B0AFFB0B349FAA130EB45771FEA" bold="true" box="[1360,1372,1440,1457]" pageId="12" pageNumber="699">V</emphasis>
|
|
, vertebral scute. Scale bar: 20 cm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BDC4718FFB0B349FCB2337844151DB0" blockId="12.[811,1457,1523,2027]" pageId="12" pageNumber="699">
|
|
The plastron of IGMp-002045 is well-preserved, without any evidence of crushing, only some minor cracks, and missing most of the anterior plastral lobe (
|
|
<figureCitation id="13585B9DFFB0B349FB0D333845591CD6" box="[1276,1394,1651,1677]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="12" pageNumber="699">Fig. 6D, E</figureCitation>
|
|
). The sutures between the bones, as well as the sulci left by the scutes are clearly identifiable. The preserved sulci indicate that the femoral was longer than the abdominal, and the anal was the shortest one of these three scutes. The xiphiplastra exhibit a large and deep U-shaped anal notch and long and narrow posterior tips. The plastron is characterized by a deep. medial concavity (
|
|
<figureCitation id="13585B9DFFB0B349FBA3321944B81D37" box="[1106,1171,1874,1900]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="12" pageNumber="699">Fig. 6</figureCitation>
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E-H). These features suggest that the specimen potentially represents a male. Even though most of the anterior plastral lobe is missing, its left margin is preserved enough to indicate that this lobe was wider than the posterior.
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</paragraph>
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<caption id="DF1C1790FFB1B348FF77301542641FBB" pageId="13" pageNumber="700" startId="13.[134,143,1374,1391]" targetBox="[153,1457,219,1327]" targetPageId="13" targetType="figure">
|
|
<paragraph id="8BDC4718FFB1B348FF77301542641FBB" blockId="13.[132,1457,1374,1504]" pageId="13" pageNumber="700">
|
|
<smallCapsWord id="8D3AD1C4FFB1B348FF77301540B61F35" baselines="1387,1387" box="[134,157,1374,1391]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="13" pageNumber="700">FIG</smallCapsWord>
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|
. 6. — Articulated shell of
|
|
<taxonomicName id="4C633C9BFFB1B348FE83301542501F34" authority="Wood, 1976" authorityName="Wood" authorityYear="1976" box="[370,635,1374,1391]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="13" pageNumber="700" phylum="Chordata" rank="species" species="colombiana">
|
|
<emphasis id="B9179B0AFFB1B348FE833015423B1F34" box="[370,528,1374,1391]" italics="true" pageId="13" pageNumber="700">Chelus colombiana</emphasis>
|
|
<bibRefCitation id="EFF23AE9FFB1B348FDE4301542501F34" author="WOOD R. C." box="[533,635,1374,1391]" pageId="13" pageNumber="700" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood, 1976</bibRefCitation>
|
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</taxonomicName>
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|
IGMp-002045 from the Tatacoa of Colombia:
|
|
<emphasis id="B9179B0AFFB1B348FC0B3015442C1F34" bold="true" box="[1018,1031,1374,1391]" pageId="13" pageNumber="700">A</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB1B348FBE1301544361F34" bold="true" box="[1040,1053,1374,1391]" pageId="13" pageNumber="700">B</emphasis>
|
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, shell in dorsal view;
|
|
<emphasis id="B9179B0AFFB1B348FB3F301544F71F34" bold="true" box="[1230,1244,1374,1391]" pageId="13" pageNumber="700">C</emphasis>
|
|
, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (
|
|
<emphasis id="B9179B0AFFB1B348FD5B303F43201FDE" bold="true" box="[682,779,1396,1413]" pageId="13" pageNumber="700">red arrows</emphasis>
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|
);
|
|
<emphasis id="B9179B0AFFB1B348FCEB303F43031FDE" bold="true" box="[794,808,1396,1413]" pageId="13" pageNumber="700">D</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB1B348FCC2303F43141FDE" bold="true" box="[819,831,1396,1413]" pageId="13" pageNumber="700">E</emphasis>
|
|
, shell in ventral view;
|
|
<emphasis id="B9179B0AFFB1B348FC06303F44291FDE" bold="true" box="[1015,1026,1396,1413]" pageId="13" pageNumber="700">F</emphasis>
|
|
,
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|
<emphasis id="B9179B0AFFB1B348FBFC303F44301FDE" bold="true" box="[1037,1051,1396,1413]" pageId="13" pageNumber="700">G</emphasis>
|
|
, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;
|
|
<emphasis id="B9179B0AFFB1B348FD5C30C042901FC7" bold="true" box="[685,699,1419,1436]" pageId="13" pageNumber="700">H</emphasis>
|
|
, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations:
|
|
<emphasis id="B9179B0AFFB1B348FF3030E940CE1FE8" bold="true" box="[193,229,1442,1459]" pageId="13" pageNumber="700">Abd</emphasis>
|
|
, abdominal scute;
|
|
<emphasis id="B9179B0AFFB1B348FE7330E941891FE8" bold="true" box="[386,418,1442,1459]" pageId="13" pageNumber="700">ace</emphasis>
|
|
, acetabulum;
|
|
<emphasis id="B9179B0AFFB1B348FDE930E942101FE8" bold="true" box="[536,571,1442,1459]" pageId="13" pageNumber="700">Ana</emphasis>
|
|
, anal scute;
|
|
<emphasis id="B9179B0AFFB1B348FD5530E942E01FE8" bold="true" box="[676,715,1442,1459]" pageId="13" pageNumber="700">Fem</emphasis>
|
|
, femoral scute;
|
|
<emphasis id="B9179B0AFFB1B348FCBE30E943441FE8" bold="true" box="[847,879,1442,1459]" pageId="13" pageNumber="700">hyo</emphasis>
|
|
, hyoplastron;
|
|
<emphasis id="B9179B0AFFB1B348FC1730E9442D1FE8" bold="true" box="[998,1030,1442,1459]" pageId="13" pageNumber="700">hyp</emphasis>
|
|
, hypoplastron;
|
|
<emphasis id="B9179B0AFFB1B348FB7630E944BE1FE8" bold="true" box="[1159,1173,1442,1459]" pageId="13" pageNumber="700">ili</emphasis>
|
|
, ilium;
|
|
<emphasis id="B9179B0AFFB1B348FB2130E944C11FE8" bold="true" box="[1232,1258,1442,1459]" pageId="13" pageNumber="700">isc</emphasis>
|
|
, ischium;
|
|
<emphasis id="B9179B0AFFB1B348FACF30E9457E1FE8" bold="true" box="[1342,1365,1442,1459]" pageId="13" pageNumber="700">lcr</emphasis>
|
|
, left costal ridge;
|
|
<emphasis id="B9179B0AFFB1B348FF4830F340E11F92" bold="true" box="[185,202,1464,1481]" pageId="13" pageNumber="700">M</emphasis>
|
|
, marginal scute;
|
|
<emphasis id="B9179B0AFFB1B348FEA830F3415D1F92" bold="true" box="[345,374,1464,1481]" pageId="13" pageNumber="700">ner</emphasis>
|
|
, neural ridge;
|
|
<emphasis id="B9179B0AFFB1B348FE1F30F3422F1F92" bold="true" box="[494,516,1464,1481]" pageId="13" pageNumber="700">pe</emphasis>
|
|
, peripheral bone;
|
|
<emphasis id="B9179B0AFFB1B348FD6D30F342831F92" bold="true" box="[668,680,1464,1481]" pageId="13" pageNumber="700">P</emphasis>
|
|
, pleural scute;
|
|
<emphasis id="B9179B0AFFB1B348FCD830F343601F92" bold="true" box="[809,843,1464,1481]" pageId="13" pageNumber="700">Pec</emphasis>
|
|
, pectoral scute;
|
|
<emphasis id="B9179B0AFFB1B348FC2630F343D21F92" bold="true" box="[983,1017,1464,1481]" pageId="13" pageNumber="700">pub</emphasis>
|
|
, pubis;
|
|
<emphasis id="B9179B0AFFB1B348FBCD30F3447E1F92" bold="true" box="[1084,1109,1464,1481]" pageId="13" pageNumber="700">rcr</emphasis>
|
|
, right costal ridge;
|
|
<emphasis id="B9179B0AFFB1B348FB0730F345391F92" bold="true" box="[1270,1298,1464,1481]" pageId="13" pageNumber="700">tov</emphasis>
|
|
, thoracic vertebra;
|
|
<emphasis id="B9179B0AFFB1B348FF75308440BB1FBB" bold="true" box="[132,144,1487,1504]" pageId="13" pageNumber="700">V</emphasis>
|
|
, vertebral scute;
|
|
<emphasis id="B9179B0AFFB1B348FED0308441101FBB" bold="true" box="[289,315,1487,1504]" pageId="13" pageNumber="700">xip</emphasis>
|
|
, xiphiplastron. Scale bar: 10 cm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BDC4718FFB1B348FF6A335842891D57" blockId="13.[131,777,1555,2028]" pageId="13" pageNumber="700">
|
|
The pelvic girdle of IGMp-002045 is well preserved, showing a strong sutural contact between the ilia and the carapace and the ischia and pubes with the plastron (
|
|
<figureCitation id="13585B9DFFB1B348FD98331842831C36" box="[617,680,1619,1645]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="13" pageNumber="700">Fig. 6</figureCitation>
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F-H). In left lateral view (
|
|
<figureCitation id="13585B9DFFB1B348FEC63338418E1CD6" box="[311,421,1651,1677]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="13" pageNumber="700">Fig. 6F, G</figureCitation>
|
|
), the acetabulum is well defined, only slightly dislocated from its original anatomical orientation due to some fractures present in the left ischium. The ilia are robust at their contact with costals 8, suprapygal and possibly reaching a portion of the pygal (
|
|
<figureCitation id="13585B9DFFB1B348FDCF33B942BF1D57" box="[574,660,1778,1804]" captionStart="FIG" captionStartId="13.[134,143,1374,1391]" captionTargetBox="[153,1457,219,1327]" captionTargetId="figure-500@13.[697,1266,223,787]" captionTargetPageId="13" captionText="FIG. 6. — Articulated shell of Chelus colombiana Wood, 1976 IGMp-002045 from the Tatacoa of Colombia: A, B, shell in dorsal view; C, view of the most anterovisceral view of the shell showing the large costovertebral tunnel (red arrows); D, E, shell in ventral view; F, G, left posterolateral view of the shell showing the left pelvic girdle and the posterior end of the costovertebral tunnel;H, posterior view of the shell showing both pelvic girdles and the xiphiplastral notch. Abbreviations: Abd, abdominal scute; ace, acetabulum; Ana, anal scute; Fem, femoral scute; hyo, hyoplastron; hyp, hypoplastron; ili, ilium; isc, ischium; lcr, left costal ridge; M, marginal scute; ner, neural ridge; pe, peripheral bone; P, pleural scute; Pec, pectoral scute; pub, pubis; rcr, right costal ridge; tov, thoracic vertebra; V, vertebral scute; xip, xiphiplastron. Scale bar: 10 cm." pageId="13" pageNumber="700">Fig. 6H</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB1B348FF6A325945831DD7" blockId="13.[131,777,1555,2028]" lastBlockId="13.[811,1458,1555,2028]" pageId="13" pageNumber="700">
|
|
VPPLT-380 consists of a nearly complete plastron, only missing the most lateral portions of the left hyoplastron and hypoplastron (
|
|
<figureCitation id="13585B9DFFB1B348FEAE3219418A1D36" box="[351,417,1874,1901]" captionStart="FIG" captionStartId="14.[131,142,1602,1619]" captionTargetBox="[151,1441,215,1560]" captionTargetId="figure-288@14.[687,1256,789,1366]" captionTargetPageId="14" captionText="FIG. 7. — Complete plastra of Chelus colombiana Wood, 1976 VPPLT-380 and MT-22 from the Tatacoa of Colombia: A, B, VPPLT-380 specimen in dorsal view; C, D, VPPLT-380 specimen in ventral view; E, F, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Ext II, extragular scute II; Fem, femoral scute; Gul, gular scute; isc, ischium scar; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; Pec, pectoral scute; psc, pubis scar; xip, xiphiplastron. Scale bar: 20 cm." pageId="13" pageNumber="700">Fig. 7</figureCitation>
|
|
). In visceral view (
|
|
<figureCitation id="13585B9DFFB1B348FD8E321942D21D37" box="[639,761,1874,1901]" captionStart="FIG" captionStartId="14.[131,142,1602,1619]" captionTargetBox="[151,1441,215,1560]" captionTargetId="figure-288@14.[687,1256,789,1366]" captionTargetPageId="14" captionText="FIG. 7. — Complete plastra of Chelus colombiana Wood, 1976 VPPLT-380 and MT-22 from the Tatacoa of Colombia: A, B, VPPLT-380 specimen in dorsal view; C, D, VPPLT-380 specimen in ventral view; E, F, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Ext II, extragular scute II; Fem, femoral scute; Gul, gular scute; isc, ischium scar; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; Pec, pectoral scute; psc, pubis scar; xip, xiphiplastron. Scale bar: 20 cm." pageId="13" pageNumber="700">Fig. 7A, B</figureCitation>
|
|
), the pubic scars are oval in shape and the ischia scars are nearly trapezoidal in shape indicating that the ischia almost touch each other medially. In ventral view (
|
|
<figureCitation id="13585B9DFFB1B348FDA832F942FB1D97" box="[601,720,1970,1996]" captionStart="FIG" captionStartId="14.[131,142,1602,1619]" captionTargetBox="[151,1441,215,1560]" captionTargetId="figure-288@14.[687,1256,789,1366]" captionTargetPageId="14" captionText="FIG. 7. — Complete plastra of Chelus colombiana Wood, 1976 VPPLT-380 and MT-22 from the Tatacoa of Colombia: A, B, VPPLT-380 specimen in dorsal view; C, D, VPPLT-380 specimen in ventral view; E, F, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Ext II, extragular scute II; Fem, femoral scute; Gul, gular scute; isc, ischium scar; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; Pec, pectoral scute; psc, pubis scar; xip, xiphiplastron. Scale bar: 20 cm." pageId="13" pageNumber="700">Fig. 7C, D</figureCitation>
|
|
), the sutures between the bones and the sulci left by scutes are well-preserved. The anterior plastral lobe is wider than the posterior, having a large entoplastron and a relatively long medial contact between both epiplastra. The hyoplastra are slightly larger than the hypoplastra. There were two pairs of extragular scutes (I and II), enclosing together with the humerals the gular scute, which had a hexagonal shape and barely reached the anteromedial margin of the plastral lobe. The plastral formula was Abd> Fem> Pec = Hum = Gul> Ana> ExtII. The xiphiplastra exhibit a large and deep U-opened anal notch, long and narrow posterior tips and a medial concavity at the medial sutural contact between each other, indicating that the specimen was potentially a male.
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB1B34BFCB232D941871D37" blockId="13.[811,1458,1555,2028]" lastBlockId="14.[132,775,1746,1901]" lastPageId="14" lastPageNumber="701" pageId="13" pageNumber="700">
|
|
VPPLT-1756. It is an articulated shell, preserving most of the plastron (missing the anterior regions of epiplastra and entoplastron), most of the peripherals and the pygal (missing portions of peripherals 1 and the nuchal), and most of costals 1 and lateral portions of costals 5 (
|
|
<figureCitation id="13585B9DFFB2B34BFD4633B942DC1D57" box="[695,759,1778,1804]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="14" pageNumber="701">Fig. 8</figureCitation>
|
|
). It also preserves the right femur, most of the right scapula, the right coracoid, cervical vertebrae 7 and 8, and remains of some thoracic vertebrae.
|
|
</paragraph>
|
|
<caption id="DF1C1790FFB2B34BFF72330944F21CCC" pageId="14" pageNumber="701" startId="14.[131,142,1602,1619]" targetBox="[151,1441,215,1560]" targetPageId="14" targetType="figure">
|
|
<paragraph id="8BDC4718FFB2B34BFF72330944F21CCC" blockId="14.[131,1457,1602,1687]" pageId="14" pageNumber="701">
|
|
<smallCapsWord id="8D3AD1C4FFB2B34BFF72330940B61C08" baselines="1615,1616" box="[131,157,1602,1619]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="14" pageNumber="701">FIG</smallCapsWord>
|
|
. 7. — Complete plastra of
|
|
<taxonomicName id="4C633C9BFFB2B34BFE8D330942AD1C08" authority="Wood, 1976" authorityName="Wood" authorityYear="1976" box="[380,646,1602,1619]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="14" pageNumber="701" phylum="Chordata" rank="species" species="colombiana">
|
|
<emphasis id="B9179B0AFFB2B34BFE8D330942301C08" box="[380,539,1602,1619]" italics="true" pageId="14" pageNumber="701">Chelus colombiana</emphasis>
|
|
<bibRefCitation id="EFF23AE9FFB2B34BFDD1330942AD1C08" author="WOOD R. C." box="[544,646,1602,1619]" pageId="14" pageNumber="701" pagination="1 - 26" refId="ref14041" refString="WOOD R. C. 1976. - Two new species of Chelus (Testudines: Pleurodira) from the Late Tertiary of northern South America. Breviora 435: 1 - 26. https: // www. biodiversitylibrary. org / page / 6590474" type="journal article" year="1976">Wood, 1976</bibRefCitation>
|
|
</taxonomicName>
|
|
VPPLT-380 and MT-22 from the Tatacoa of Colombia:
|
|
<emphasis id="B9179B0AFFB2B34BFBA0330944751C08" bold="true" box="[1105,1118,1602,1619]" pageId="14" pageNumber="701">A</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB2B34BFB993309445E1C08" bold="true" box="[1128,1141,1602,1619]" pageId="14" pageNumber="701">B</emphasis>
|
|
, VPPLT-380 specimen in dorsal view;
|
|
<emphasis id="B9179B0AFFB2B34BFF75331240B91C31" bold="true" box="[132,146,1625,1642]" pageId="14" pageNumber="701">C</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB2B34BFF6F331240871C31" bold="true" box="[158,172,1625,1642]" pageId="14" pageNumber="701">D</emphasis>
|
|
, VPPLT-380 specimen in ventral view;
|
|
<emphasis id="B9179B0AFFB2B34BFE0B3312422D1C31" bold="true" box="[506,518,1625,1642]" pageId="14" pageNumber="701">E</emphasis>
|
|
,
|
|
<emphasis id="B9179B0AFFB2B34BFDE3331242361C31" bold="true" box="[530,541,1625,1642]" pageId="14" pageNumber="701">F</emphasis>
|
|
, MT-22 specimen, plastron articulated with carapace in ventral view. Abbreviations:
|
|
<emphasis id="B9179B0AFFB2B34BFB1F331245391C31" bold="true" box="[1262,1298,1625,1642]" pageId="14" pageNumber="701">Abd</emphasis>
|
|
, abdominal scute;
|
|
<emphasis id="B9179B0AFFB2B34BFF75333B408C1CDA" bold="true" box="[132,167,1648,1665]" pageId="14" pageNumber="701">Ana</emphasis>
|
|
, anal scute;
|
|
<emphasis id="B9179B0AFFB2B34BFEE3333B41051CDA" bold="true" box="[274,302,1648,1665]" pageId="14" pageNumber="701">ent</emphasis>
|
|
, entoplastron;
|
|
<emphasis id="B9179B0AFFB2B34BFE5D333B41EC1CDA" bold="true" box="[428,455,1648,1665]" pageId="14" pageNumber="701">epi</emphasis>
|
|
, epiplastron;
|
|
<emphasis id="B9179B0AFFB2B34BFDC8333B42491CDA" bold="true" box="[569,610,1648,1665]" pageId="14" pageNumber="701">Ext I</emphasis>
|
|
, extragular scute I;
|
|
<emphasis id="B9179B0AFFB2B34BFCF6333B431F1CDA" bold="true" box="[775,820,1648,1665]" pageId="14" pageNumber="701">Ext II</emphasis>
|
|
, extragular scute II;
|
|
<emphasis id="B9179B0AFFB2B34BFC2E333B442D1CDA" bold="true" box="[991,1030,1648,1665]" pageId="14" pageNumber="701">Fem</emphasis>
|
|
, femoral scute;
|
|
<emphasis id="B9179B0AFFB2B34BFB7D333B44811CDA" bold="true" box="[1164,1194,1648,1665]" pageId="14" pageNumber="701">Gul</emphasis>
|
|
, gular scute;
|
|
<emphasis id="B9179B0AFFB2B34BFAED333B451D1CDA" bold="true" box="[1308,1334,1648,1665]" pageId="14" pageNumber="701">isc</emphasis>
|
|
, ischium scar;
|
|
<emphasis id="B9179B0AFFB2B34BFF7533CD40851CCC" bold="true" box="[132,174,1670,1687]" pageId="14" pageNumber="701">Hum</emphasis>
|
|
, humeral scute;
|
|
<emphasis id="B9179B0AFFB2B34BFEC833CD41721CCC" bold="true" box="[313,345,1670,1687]" pageId="14" pageNumber="701">hyo</emphasis>
|
|
, hyoplastron;
|
|
<emphasis id="B9179B0AFFB2B34BFE2133CD41DB1CCC" bold="true" box="[464,496,1670,1687]" pageId="14" pageNumber="701">hyp</emphasis>
|
|
, hypoplastron;
|
|
<emphasis id="B9179B0AFFB2B34BFD8333CD42BF1CCC" bold="true" box="[626,660,1670,1687]" pageId="14" pageNumber="701">Pec</emphasis>
|
|
, pectoral scute;
|
|
<emphasis id="B9179B0AFFB2B34BFCD133CD436B1CCC" bold="true" box="[800,832,1670,1687]" pageId="14" pageNumber="701">psc</emphasis>
|
|
, pubis scar;
|
|
<emphasis id="B9179B0AFFB2B34BFC5A33CD43EE1CCC" bold="true" box="[939,965,1670,1687]" pageId="14" pageNumber="701">xip</emphasis>
|
|
, xiphiplastron. Scale bar: 20 cm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BDC4718FFB2B34BFF7532D940CF1DF7" blockId="14.[132,228,1938,1964]" box="[132,228,1938,1964]" pageId="14" pageNumber="701">
|
|
<heading id="D094F074FFB2B34BFF7532D940CF1DF7" box="[132,228,1938,1964]" fontSize="11" level="2" pageId="14" pageNumber="701" reason="8">
|
|
<emphasis id="B9179B0AFFB2B34BFF7532D940CF1DF7" box="[132,228,1938,1964]" italics="true" pageId="14" pageNumber="701">Carapace</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB2B34BFF7532F9432D1DB0" blockId="14.[131,775,1970,2028]" pageId="14" pageNumber="701">The shape of the carapace is rectangular, slightly curved at its anterior and posterior margins. Both costals 1 are preserved;</paragraph>
|
|
<paragraph id="8BDC4718FFB2B34BFCDD3399459B1D17" blockId="14.[811,1456,1746,1869]" pageId="14" pageNumber="701">
|
|
including ventrally the long axillary buttress that likely reached costals 2. Other preserved regions of the carapace are the lateral portions of costals 5, most of the peripherals and the pygal. However, peripherals 1 and the nuchal are missing (
|
|
<figureCitation id="13585B9DFFB2B34BFAA03279458F1D17" box="[1361,1444,1842,1868]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="14" pageNumber="701">Fig. 8A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB2B34BFCDC323943A91DD7" blockId="14.[813,898,1906,1932]" box="[813,898,1906,1932]" pageId="14" pageNumber="701">
|
|
<heading id="D094F074FFB2B34BFCDC323943A91DD7" box="[813,898,1906,1932]" fontSize="11" level="2" pageId="14" pageNumber="701" reason="8">
|
|
<emphasis id="B9179B0AFFB2B34BFCDC323943A91DD7" box="[813,898,1906,1932]" italics="true" pageId="14" pageNumber="701">Plastron</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB2B34AFCDC32D941E91BAA" blockId="14.[812,1455,1938,2028]" lastBlockId="15.[130,777,216,1071]" lastPageId="15" lastPageNumber="702" pageId="14" pageNumber="701">
|
|
The plastron is almost completely preserved, missing only the most anterior portions of both epiplastra and the most anterior corner of the entoplastron (
|
|
<figureCitation id="13585B9DFFB2B34BFB41329A450B1DB7" box="[1200,1312,2001,2028]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="14" pageNumber="701">Fig. 8B, C</figureCitation>
|
|
). The sutures between bones, as well as the sulci left by the scutes are clearly identifiable. The femoral scute was the longest of the scutes, resulting in a plastral formula Fem> Abd> Pec> Ana = Hum. The xiphiplastra exhibit a large and deep U-shaped anal notch and long and narrow posterior tips. Portions of both pelvic girdles are still attached to the plastron via the ischia and pubis. The plastron lacks of deep concavity at the medial region of xiphiplastral, suggesting that it was possibly a female individual.
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB3B34AFF6A34BC41EB188A" blockId="15.[130,777,216,1071]" pageId="15" pageNumber="702">
|
|
The right femur is preserved, missing most of the major and minor trochanters (
|
|
<figureCitation id="13585B9DFFB3B34AFE69375C41F1186A" box="[408,474,535,561]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8</figureCitation>
|
|
D-F). The femoral shaft is arched and exhibits an oval-shaped outline. The femoral head is oval in shape, eroded at its more proximal region and slightly projected laterally. In all aspects, the femur of VPPLT-1756 resembles the femur of the extant specimens of
|
|
<taxonomicName id="4C633C9BFFB3B34AFF5337FD410E188B" authorityName="Schneider" authorityYear="1783" box="[162,293,694,720]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="702" phylum="Chordata" rank="species" species="fimbriata">
|
|
<emphasis id="B9179B0AFFB3B34AFF5337FD410E188B" box="[162,293,694,720]" italics="true" pageId="15" pageNumber="702">C. fimbriata</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C633C9BFFB3B34AFEAC37FD4192188B" authorityName="Wood" authorityYear="1976" box="[349,441,694,720]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="702" phylum="Chordata" rank="species" species="lewisi">
|
|
<emphasis id="B9179B0AFFB3B34AFEAC37FD4192188B" box="[349,441,694,720]" italics="true" pageId="15" pageNumber="702">C. lewisi</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB3B34AFF6A379D42D21E74" blockId="15.[130,777,216,1071]" pageId="15" pageNumber="702">
|
|
VPPLT-1756 preserved the right scapula and coracoid (
|
|
<figureCitation id="13585B9DFFB3B34AFF7C37BD410A194B" box="[141,289,758,784]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8G, H, J</figureCitation>
|
|
). The anterodorsal process of the scapula has an oval-shaped when seen in cross section, and it is broken, missing most of its distal end. There is a moderate notch in proximity to the glenoid fossa. The ventromedial prong (acromial process) is flatter than the anterodorsal and exhibits a dorsomedial ridge.The internal angle between the two processes is 85° (
|
|
<figureCitation id="13585B9DFFB3B34AFF2236FE41071994" box="[211,300,949,975]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8H</figureCitation>
|
|
). In
|
|
<taxonomicName id="4C633C9BFFB3B34AFE9536FD41CC1994" authorityName="Schneider" authorityYear="1783" box="[356,487,949,976]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="702" phylum="Chordata" rank="species" species="fimbriata">
|
|
<emphasis id="B9179B0AFFB3B34AFE9536FD41CC1994" box="[356,487,949,976]" italics="true" pageId="15" pageNumber="702">C. fimbriata</emphasis>
|
|
</taxonomicName>
|
|
this angle is 75° (
|
|
<figureCitation id="13585B9DFFB3B34AFD5F36FE42D31994" box="[686,760,949,975]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8I</figureCitation>
|
|
). The coracoid has a flat distal blade which, although broken, is preserved enough to indicate that this region formed a broad surface, similarly to the coracoid of
|
|
<taxonomicName id="4C633C9BFFB3B34AFDF7315E42A31E74" authorityName="Schneider" authorityYear="1783" box="[518,648,1045,1071]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="702" phylum="Chordata" rank="species" species="fimbriata">
|
|
<emphasis id="B9179B0AFFB3B34AFDF7315E42A31E74" box="[518,648,1045,1071]" italics="true" pageId="15" pageNumber="702">C. fimbriata</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="13585B9DFFB3B34AFD69315E42C71E74" box="[664,748,1045,1071]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8K</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB3B34AFF75311E40CA1E34" blockId="15.[132,225,1109,1135]" box="[132,225,1109,1135]" pageId="15" pageNumber="702">
|
|
<heading id="D094F074FFB3B34AFF75311E40CA1E34" box="[132,225,1109,1135]" fontSize="11" level="2" pageId="15" pageNumber="702" reason="8">
|
|
<emphasis id="B9179B0AFFB3B34AFF75311E40CA1E34" box="[132,225,1109,1135]" italics="true" pageId="15" pageNumber="702">Cervicals</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="8BDC4718FFB3B34AFF75313F41BA1C76" blockId="15.[131,776,1140,1582]" pageId="15" pageNumber="702">
|
|
Cervicals 7 and 8 are preserved slightly crushed (
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<figureCitation id="13585B9DFFB3B34AFD7D313F42E21ED4" box="[652,713,1140,1167]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8</figureCitation>
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L-U). Cervical 7 has a biconcave centrum, exhibiting a long and robust dorsoventrally projected neural spine, with postzygapophyses facing ventrally and meeting medially, and a long ventral keel (
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<figureCitation id="13585B9DFFB3B34AFEE131BF41661F55" box="[272,333,1268,1294]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8</figureCitation>
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L-P). Cervical 8 has a biconvex centrum, exhibiting also a long dorsoventrally projected neural spine that ends at the postzygapophyses as cervical 7, which face ventrolaterally and meet medially. It is shorter than cervical 7, and it has a very long ventral keel that ends in a dropshape anterior tip (
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<figureCitation id="13585B9DFFB3B34AFEBE30D841A01FF5" box="[335,395,1427,1454]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8</figureCitation>
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Q-U). VPPLT-1756 cervicals 7 and 8 resemble the centra of the same vertebrae in
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<taxonomicName id="4C633C9BFFB3B34AFD7230FF432C1F96" authorityName="Schneider" authorityYear="1783" box="[643,775,1459,1486]" class="Reptilia" family="Chelidae" genus="Chelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="15" pageNumber="702" phylum="Chordata" rank="species" species="fimbriata">
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<emphasis id="B9179B0AFFB3B34AFD7230FF432C1F96" box="[643,775,1459,1486]" italics="true" pageId="15" pageNumber="702">C. fimbriata</emphasis>
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</taxonomicName>
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(ICN-1781;
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<figureCitation id="13585B9DFFB3B34AFEE6309841591FB6" box="[279,370,1491,1517]" captionStart="FIG" captionStartId="16.[132,143,1825,1842]" captionTargetBox="[132,1445,217,1796]" captionTargetId="figure-294@16.[698,1267,702,1159]" captionTargetPageId="16" captionText="FIG. 8. — Chelus colombiana Wood, 1976 VPPLT-1756 specimen from the Tatacoa of Colombia: A, shell in dorsal view; B, C, plastron in ventral view; D-F, right femur in: D, dorsal view; E, ventral view; F, anterior view; G, H, right scapula in anterior view; I, right scapula Chelus fimbriata Schneider, 1783 UR-uncatalogued specimen in ventral view; J, right coracoid in ventral view; K, right coracoid C. fimbriata UR-uncatalogued specimen in ventral view; L-P, cervical vertebra 7 in: L, anterior view; M, dorsal view; N, posterior view; O, ventral view; P, right lateral view; Q-U, cervical vertebra 8 in: Q, anterior view; R, dorsal view; S, posterior view; T, ventral view; U, right lateral view; V, cervicals 7 and 8 of C. fimbriata ICN-1781 specimen in right lateral view, with schematic centra articulation below. Abbreviations: Abd, abdominal scute; Ana, anal scute; ent, entoplastron; epi, epiplastron; Ext I, extragular scute I; Fem, femoral scute; fh, femoral head; Gul, gular scute; Hum, humeral scute; hyo, hyoplastron; hyp, hypoplastron; ltp, lateral process; nec, neural canal; nes, neural spine; ntc, notch; Pec, pectoral scute; poz, postzygapophysis; prz, prezygapophysis; tia, tibial articulation; tma, trochanter major; tmin, trochanter min; vek, ventral keel; xip, xiphiplastron. Scale bars: A-C, 10 cm; D-V, 5 cm." pageId="15" pageNumber="702">Fig. 8V</figureCitation>
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). In the extant taxon, however, the neural spines of these vertebrae are shorter, broader and more dorsally projected.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |