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<mods:title id="D387C7B58F0154296C828C44AD1DD45B">A new species of the Scinax cruentomma group (Anura: Hylidae) from the Ucayali River basin of Loreto, Peru</mods:title>
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<mods:namePart id="8CF2E631E454FE8D09AEF019C5304E23">Padial, José M.</mods:namePart>
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<mods:affiliation id="02FAE3AD9367367BADD41443FE7AD3CE">División Herpetología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” - CONICET, Av. Ángel Gallardo 470, C 1405 DJR, CABA, Argentina &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, C 1428 EGA, CABA, Argentina</mods:affiliation>
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(
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sp.
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18— 
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Araujo-Vieira 
<emphasis id="B917EAACFFFCFFB8EA9D07FB3193FC5C" box="[458,509,975,999]" italics="true" pageId="3" pageNumber="404">et al.</emphasis>
2023
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. Phylogenetic analysis and recognition as confirmed candidate species.
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CRBIIAP 2912 (
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male collected on 
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</paragraph>
<paragraph id="8BDC36BEFFFCFFB8EB9000AA323BFA2C" blockId="3.[151,1436,1045,2008]" pageId="3" pageNumber="404">
<materialsCitation id="3B0B3CE3FFFCFFB8EB9000AA3406FB7B" collectionCode="GGU, MACN" isTailEnumeration="true" pageId="3" pageNumber="404" specimenCode="GGU 5313, 5353, 5358, 5468, 7618, MACN 59716, GGU 7617" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B917EAACFFFCFFB8EB9000AA3128FB20" bold="true" box="[199,326,1153,1179]" pageId="3" pageNumber="404">
<typeStatus id="54D8881CFFFCFFB8EB9000AA312FFB20" box="[199,321,1153,1179]" pageId="3" pageNumber="404" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D65FD37FFFCFFB8EA1800AA318FFB20" box="[335,481,1153,1179]" count="12" pageId="3" pageNumber="404" type="adult">Twelve adult</specimenCount>
males and 
<specimenCount id="9D65FD37FFFCFFB8E93200A93288FB20" box="[613,742,1153,1180]" count="1" pageId="3" pageNumber="404" type="female">one female</specimenCount>
. CRBIIAP 1570, 1595, 1597, 1656, 2913 (
<specimenCode id="DBC59EC5FFFCFFB8EF9900AA353EFB27" box="[1230,1360,1153,1180]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 5313</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EE0A00AA35F6FB27" box="[1373,1432,1153,1180]" collectionCode="GGU" pageId="3" pageNumber="404">5353</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EBC0008E30BCFB7B" box="[151,210,1189,1216]" collectionCode="GGU" pageId="3" pageNumber="404">5358</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EB8A008E3176FB7B" box="[221,280,1189,1216]" collectionCode="GGU" pageId="3" pageNumber="404">5468</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EA75008E3130FB04" box="[290,350,1189,1215]" collectionCode="GGU" pageId="3" pageNumber="404">7618</specimenCode>
), 
<specimenCode id="DBC59EC5FFFCFFB8EA26008E327FFB7B" box="[369,529,1189,1216]" collectionCode="MACN" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="404" type="Museum">MACN 59716</specimenCode>
(
<specimenCode id="DBC59EC5FFFCFFB8E948008E32CEFB04" box="[543,672,1189,1215]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 7617</specimenCode>
), 
<specimenCode id="DBC59EC5FFFCFFB8E9E5008E3352FB7B" box="[690,828,1189,1216]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="404">MCP 14562</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8E811008E33E2FB7B" box="[838,908,1189,1216]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="404">14564</specimenCode>
(
<specimenCode id="DBC59EC5FFFCFFB8E8CD008E3474FB7B" box="[922,1050,1189,1216]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 5355</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EF73008E340EFB7B" box="[1060,1120,1189,1216]" collectionCode="GGU" pageId="3" pageNumber="404">5586</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0B3CE3FFFCFFB8EF24008E34DDFA97" collectingDateMax="2022-05-31" collectingDateMin="2017-04-30" collectionCode="GGU, MCP" country="Peru" county="Samantha Solis-Ruiz" elevation="123" latitude="-4.901539" location="Giussepe Gagliardi-Urrutia" longLatPrecision="1" longitude="-73.63347" municipality="Katyuscia Araujo-Vieira" pageId="3" pageNumber="404" specimenCode="GGU 5578, MCP 14565, 15108, GGU 5099, 6305, 6301, 5117" specimenCount="1" stateProvince="Loreto" typeStatus="paratype">
males, and CRBIIAP 1712 (
<specimenCode id="DBC59EC5FFFCFFB8EBC800E2314FFB5F" box="[159,289,1225,1252]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 5578</specimenCode>
), female, collected on different dates in 
<collectingDate id="EF99E996FFFCFFB8E9B000E234E1FB58" box="[743,1167,1225,1252]" isEnumeration="true" pageId="3" pageNumber="404">April and May 2017 and March 2022</collectingDate>
by 
<location id="8EBC6065FFFCFFB8EFEC00E23085FABC" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8EFEC00E23085FABC" county="Samantha Solis-Ruiz" municipality="Katyuscia Araujo-Vieira" name="Giussepe Gagliardi-Urrutia" pageId="3" pageNumber="404">Giussepe Gagliardi-Urrutia</location>
, 
<location id="8EBC6065FFFCFFB8EBA000C631D6FABC" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8EBA000C631D6FABC" box="[247,440,1261,1287]" county="Samantha Solis-Ruiz" municipality="Katyuscia Araujo-Vieira" name="Ehiko Rios-Alva" pageId="3" pageNumber="404">Ehiko Rios-Alva</location>
, 
<location id="8EBC6065FFFCFFB8EA9300C6319AFABC" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8EA9300C6319AFABC" box="[452,500,1261,1287]" county="Samantha Solis-Ruiz" municipality="Katyuscia Araujo-Vieira" name="Jose" pageId="3" pageNumber="404">José</location>
M. 
<location id="8EBC6065FFFCFFB8E97200C63200FABC" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8E97200C63200FABC" box="[549,622,1261,1287]" county="Samantha Solis-Ruiz" municipality="Katyuscia Araujo-Vieira" name="Padial" pageId="3" pageNumber="404">Padial</location>
, 
<collectingMunicipality id="6BB8ACC4FFFCFFB8E92D00C633E0FABC" box="[634,910,1261,1287]" pageId="3" pageNumber="404">Katyuscia Araujo-Vieira</collectingMunicipality>
, and 
<collectingCounty id="62BD4E32FFFCFFB8E89C00C634D6FABC" box="[971,1208,1261,1287]" pageId="3" pageNumber="404">Samantha Solís-Ruiz</collectingCounty>
at the same locality as the holotype; CRBIIAP 1851, 1905, 
<specimenCode id="DBC59EC5FFFCFFB8E909013A3284FA97" box="[606,746,1297,1324]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="404">MCP 14565</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8E9AF013A3350FA97" box="[760,830,1297,1324]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="404">15108</specimenCode>
(
<specimenCode id="DBC59EC5FFFCFFB8E818013A33BFFA97" box="[847,977,1297,1324]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 5099</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8E888013A3474FA97" box="[991,1050,1297,1324]" collectionCode="GGU" pageId="3" pageNumber="404">6305</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EF70013A340CFA90" box="[1063,1122,1297,1323]" collectionCode="GGU" pageId="3" pageNumber="404">6301</specimenCode>
, 
<specimenCode id="DBC59EC5FFFCFFB8EF27013A34C5FA97" box="[1136,1195,1297,1324]" collectionCode="GGU" pageId="3" pageNumber="404">5117</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0B3CE3FFFCFFB8EF97013A323EFA2C" collectingDateMax="2018-04-30" collectingDateMin="2017-05-31" collectionCode="WGS" country="Peru" county="Requena" elevation="131" latitude="-5.88903" location="Rio Blanco" longLatPrecision="1" longitude="-73.76537" municipality="Soplin" pageId="3" pageNumber="404" specimenCode="WGS84" specimenCount="1" stateProvince="Loreto" typeStatus="paratype">
males, collected on different dates in 
<collectingDate id="EF99E996FFFCFFB8EA35011E32CCFAF4" box="[354,674,1333,1359]" isEnumeration="true" pageId="3" pageNumber="404">March 2017 and April 2018</collectingDate>
by Giussepe Gagliardi-Urrutia, Ehiko Rios-Alva, Samantha Solís-Ruiz, and Layné Guerra-Vargas at 
<location id="8EBC6065FFFCFFB8E947017232E3FAC8" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8E947017232E3FAC8" box="[528,653,1369,1395]" country="Peru" county="Requena" latitude="-5.88903" longLatPrecision="1" longitude="-73.76537" municipality="Soplin" name="Rio Blanco" pageId="3" pageNumber="404" stateProvince="Loreto">Río Blanco</location>
(RB), 
<location id="8EBC6065FFFCFFB8E981017133E3FAC8" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8E981017133E3FAC8" box="[726,909,1369,1396]" country="Peru" county="Requena" latitude="-5.88903" longLatPrecision="1" longitude="-73.76537" municipality="Soplin" name="Frontera Village" pageId="3" pageNumber="404" stateProvince="Loreto">Frontera Village</location>
, 
<collectingMunicipality id="6BB8ACC4FFFCFFB8E8C20172338FFAC8" box="[917,993,1369,1395]" pageId="3" pageNumber="404">Soplín</collectingMunicipality>
, 
<collectingCounty id="62BD4E32FFFCFFB8E8BD01713421FACF" box="[1002,1103,1370,1396]" pageId="3" pageNumber="404">Requena</collectingCounty>
, 
<collectingRegion id="49A7F85CFFFCFFB8EF0F017134C8FACF" box="[1112,1190,1370,1396]" country="Peru" name="Loreto" pageId="3" pageNumber="404">Loreto</collectingRegion>
, 
<collectingCountry id="F374762EFFFCFFB8EFF90171348FFACF" box="[1198,1249,1370,1396]" name="Peru" pageId="3" pageNumber="404">Peru</collectingCountry>
(datum 
<specimenCode id="DBC59EC5FFFCFFB8EE6D017235F6FAC8" box="[1338,1432,1369,1395]" collectionCode="WGS" pageId="3" pageNumber="404">WGS84</specimenCode>
; 
<geoCoordinate id="EE575079FFFCFFB8EBC00156317BFA2C" box="[151,277,1405,1432]" degrees="5.88903" direction="south" orientation="latitude" pageId="3" pageNumber="404" precision="1" value="-5.88903">5.88903° S</geoCoordinate>
, 
<geoCoordinate id="EE575079FFFCFFB8EA75015631D6FA23" box="[290,440,1405,1432]" degrees="73.76537" direction="west" orientation="longitude" pageId="3" pageNumber="404" precision="1" value="-73.76537">73.76537° W</geoCoordinate>
, 
<elevation id="004ED18DFFFCFFB8EA9401563228FA2C" box="[451,582,1405,1432]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="404" unit="m" value="131.0">
<quantity id="4C9B9B5BFFFCFFB8EA9401563264FA23" box="[451,522,1405,1432]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="404" unit="m" value="131.0">131 m</quantity>
a.s.l.
</elevation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36BEFFFCFFB8EB90018A34EEF993" blockId="3.[151,1436,1045,2008]" pageId="3" pageNumber="404">
<emphasis id="B917EAACFFFCFFB8EB90018A31D0FA00" bold="true" box="[199,446,1441,1467]" pageId="3" pageNumber="404">Referred specimens.</emphasis>
<materialsCitation id="3B0B3CE3FFFCFFB8EA9E018A3412F993" collectingDate="2017-04" collectionCode="MCP, GGU" collectorName="Jose M. Padial" country="Peru" county="Requena" elevation="123" latitude="-4.901539" location="Giussepe Gagliardi-Urrutia" longLatPrecision="1" longitude="-73.63347" municipality="Soplin" pageId="3" pageNumber="404" specimenCode="MCP 14561, GGU 5350, GGU 5152" specimenCount="1" stateProvince="Loreto">
<specimenCode id="DBC59EC5FFFCFFB8EA9E018A323AFA07" box="[457,596,1441,1468]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="3" pageNumber="404">MCP 14561</specimenCode>
(
<specimenCode id="DBC59EC5FFFCFFB8E930018A3283FA07" box="[615,749,1441,1468]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 5350</specimenCode>
), juvenile, collected on 
<date id="FFDD107EFFFCFFB8EF51018A34EBFA00" box="[1030,1157,1441,1467]" pageId="3" pageNumber="404" value="2017-04">
<collectingDate id="EF99E996FFFCFFB8EF51018A34EBFA00" box="[1030,1157,1441,1467]" pageId="3" pageNumber="404" value="2017-04">April 2017</collectingDate>
</date>
by Giussepe Gagliardi-Urrutia, 
<collectorName id="26965368FFFCFFB8EBA101EE31F3FA64" box="[246,413,1477,1504]" pageId="3" pageNumber="404">José M. Padial</collectorName>
, Ehiko Rios-Alva, and Samantha Solís-Ruiz at the same locality as the holotype; CRBIIAP 1883 (
<specimenCode id="DBC59EC5FFFCFFB8EB8B01C23132F9BF" box="[220,348,1513,1540]" collectionCode="GGU" pageId="3" pageNumber="404">GGU 5152</specimenCode>
), newly metamorphosed, collected in 
<date id="FFDD107EFFFCFFB8E9AA01C2331BF9B8" box="[765,885,1513,1539]" pageId="3" pageNumber="404" value="2017-04">
<collectingDate id="EF99E996FFFCFFB8E9AA01C2331BF9B8" box="[765,885,1513,1539]" pageId="3" pageNumber="404" value="2017-04">April 2017</collectingDate>
</date>
by 
<location id="8EBC6065FFFCFFB8E8CA01C234BEF9B8" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8E8CA01C234BEF9B8" box="[925,1232,1513,1539]" country="Peru" county="Requena" municipality="Soplin" name="Giussepe Gagliardi-Urrutia" pageId="3" pageNumber="404" stateProvince="Loreto">Giussepe Gagliardi-Urrutia</location>
, 
<location id="8EBC6065FFFCFFB8EF8D01C235F6F9B8" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8EF8D01C235F6F9B8" box="[1242,1432,1513,1539]" country="Peru" county="Requena" municipality="Soplin" name="Ehiko Rios-Alva" pageId="3" pageNumber="404" stateProvince="Loreto">Ehiko Rios-Alva</location>
, and 
<location id="8EBC6065FFFCFFB8EB90022631DCF99C" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8EB90022631DCF99C" box="[199,434,1549,1575]" country="Peru" county="Requena" municipality="Soplin" name="Samantha Solis-Ruiz" pageId="3" pageNumber="404" stateProvince="Loreto">Samantha Solís-Ruiz</location>
at 
<location id="8EBC6065FFFCFFB8EA8202263238F99C" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8EA8202263238F99C" box="[469,598,1549,1575]" country="Peru" county="Requena" municipality="Soplin" name="Rio Blanco" pageId="3" pageNumber="404" stateProvince="Loreto">Río Blanco</location>
, 
<location id="8EBC6065FFFCFFB8E93602253375F99C" LSID="urn:lsid:plazi:treatment:03CA87A8FFFCFFB7EBC0074C3492FE6A:8EBC6065FFFCFFB8E93602253375F99C" box="[609,795,1549,1576]" country="Peru" county="Requena" municipality="Soplin" name="Frontera Village" pageId="3" pageNumber="404" stateProvince="Loreto">Frontera Village</location>
, 
<collectingMunicipality id="6BB8ACC4FFFCFFB8E8720226331FF99C" box="[805,881,1549,1575]" pageId="3" pageNumber="404">Soplín</collectingMunicipality>
, 
<collectingCounty id="62BD4E32FFFCFFB8E82B0225338FF993" box="[892,993,1550,1576]" pageId="3" pageNumber="404">Requena</collectingCounty>
, 
<collectingRegion id="49A7F85CFFFCFFB8E8BB02253454F993" box="[1004,1082,1550,1576]" country="Peru" name="Loreto" pageId="3" pageNumber="404">Loreto</collectingRegion>
, 
<collectingCountry id="F374762EFFFCFFB8EF1202253412F993" box="[1093,1148,1550,1576]" name="Peru" pageId="3" pageNumber="404">Peru</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796535FFFCFFB8EB90021A3100F8FC" pageId="3" pageNumber="404" type="diagnosis">
<paragraph id="8BDC36BEFFFCFFB8EB90021A3100F8FC" blockId="3.[151,1436,1045,2008]" pageId="3" pageNumber="404">
<emphasis id="B917EAACFFFCFFB8EB90021A312CF9F0" bold="true" box="[199,322,1585,1611]" pageId="3" pageNumber="404">Diagnosis.</emphasis>
(1) male SVL 
<quantity id="4C9B9B5BFFFCFFB8EABE021A32E1F9F7" box="[489,655,1585,1612]" metricMagnitude="-2" metricUnit="m" metricValue="2.155" metricValueMax="2.29" metricValueMin="2.02" pageId="3" pageNumber="404" unit="mm" value="21.549999999999997" valueMax="22.9" valueMin="20.2">20.2–22.9 mm</quantity>
; (2) nostrils not protruding dorsolaterally; (3) canthus rostralis poorly defined, straight or slightly convex; (4) loreal region slightly concave; (5) vocal sac weakly bilobate, ventrally not reaching the pectoral region; (6) dorsum uniform light to dark brown with scattered darker blotches of different sizes; (7) iris with median horizontal red streak, in life; (8) physiological chlorosis present, bones bluish-green; (9) moderately developed pre- and postaxial webbing of Toe IV, reaching the proximal half of the penultimate phalanx; (10) m. depressor mandibulae originating from the dorsal fascia that covers the suprascapula; and (11) advertisement call composed of a single pulsed note with duration of 4–62 ms, with 1–3 pulses emitted at a rate of 45.3–69.0 pulses/s.
</paragraph>
</subSubSection>
<subSubSection id="C3796535FFFCFFB0EB90037A339BFA26" lastPageId="11" lastPageNumber="412" pageId="3" pageNumber="404" type="description">
<paragraph id="8BDC36BEFFFCFFBFEB90037A342EFE16" blockId="3.[151,1436,1045,2008]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="405" pageId="3" pageNumber="404">
<emphasis id="B917EAACFFFCFFB8EB90037A327DF8D0" bold="true" box="[199,531,1873,1899]" pageId="3" pageNumber="404">
Description of the 
<typeStatus id="54D8881CFFFCFFB8EAF2037A3260F8D0" box="[421,526,1873,1899]" pageId="3" pageNumber="404" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, 
<quantity id="4C9B9B5BFFFCFFB8E9F2037A3364F8D7" box="[677,778,1873,1900]" metricMagnitude="-2" metricUnit="m" metricValue="2.06" pageId="3" pageNumber="404" unit="mm" value="20.6">20.6 mm</quantity>
SVL (
<figureCitation id="13582A3BFFFCFFB8E802037A33DEF8D0" box="[853,944,1873,1899]" captionStart="FIGURE 2" captionStartId="4.[151,250,1127,1151]" captionTargetBox="[158,1430,453,1104]" captionTargetId="figure-157@4.[158,1430,453,1104]" captionTargetPageId="4" captionText="FIGURE 2. Scinax ushiniauae, holotype (CRBIIAP 2912). (A) Dorsal and (B) ventral views." figureDoi="http://doi.org/10.5281/zenodo.10628012" httpUri="https://zenodo.org/record/10628012/files/figure.png" pageId="3" pageNumber="404">Fig. 2A</figureCitation>
). Head longer than wide, HW 89% HL; HL 39% SVL; HW 35% SVL. Snout rounded in dorsal and lateral views; END equal to 76% ED; nostrils elliptical, not protruding dorsolaterally; the region between nostrils slightly concave; canthus rostralis poorly defined and slightly convex; loreal region slightly concave. ED 46% HW and 41% HL. Pupil horizontal. Palpebral membrane translucent, not reticulated. Tympanum distinct, round, small, TD 39% ED; tympanic annulus rounded, with upper portion hidden by the supratympanic fold. Supratympanic fold evident from the upper part of the tympanum to close to the insertion of the arm. Tongue ovoid, free laterally and posteriorly, shallowly notched posteriorly. Vomerine teeth in two barely separated transverse series, each bearing six (right) and five (left) teeth. Choanae oval. Vocal slits present, extending from the lateral base of tongue to the corner of the mouth. Vocal sac subgular, median, externally expanded, ventrally not reaching the pectoral region and not occupying space between head and body, with a slight medial constriction in the posterior portion (
<figureCitation id="13582A3BFFFBFFBFE9D805443286FE32" box="[655,744,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1127,1151]" captionTargetBox="[158,1430,453,1104]" captionTargetId="figure-157@4.[158,1430,453,1104]" captionTargetPageId="4" captionText="FIGURE 2. Scinax ushiniauae, holotype (CRBIIAP 2912). (A) Dorsal and (B) ventral views." figureDoi="http://doi.org/10.5281/zenodo.10628012" httpUri="https://zenodo.org/record/10628012/files/figure.png" pageId="4" pageNumber="405">Fig. 2B</figureCitation>
), which gives it a bilobate shape when deflated and inflated. Pectoral fold present, with pre-and postaxillar elements. Axillary membrane absent.
</paragraph>
<caption id="DF1C6636FFFBFFBFEBC0004C3423FB3B" ID-DOI="http://doi.org/10.5281/zenodo.10628012" ID-Zenodo-Dep="10628012" box="[151,1101,1127,1152]" httpUri="https://zenodo.org/record/10628012/files/figure.png" pageId="4" pageNumber="405" startId="4.[151,250,1127,1151]" targetBox="[158,1430,453,1104]" targetPageId="4" targetType="figure">
<paragraph id="8BDC36BEFFFBFFBFEBC0004C3423FB3B" blockId="4.[151,1101,1127,1152]" box="[151,1101,1127,1152]" pageId="4" pageNumber="405">
<emphasis id="B917EAACFFFBFFBFEBC0004C317DFB3B" bold="true" box="[151,275,1127,1152]" pageId="4" pageNumber="405">FIGURE 2.</emphasis>
<taxonomicName id="4C634D3DFFFBFFBFEA4E004331BCFB3B" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[281,466,1128,1152]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFFBFFBFEA4E004331BCFB3B" box="[281,466,1128,1152]" italics="true" pageId="4" pageNumber="405">Scinax ushiniauae</emphasis>
</taxonomicName>
, holotype (CRBIIAP 2912). 
<emphasis id="B917EAACFFFBFFBFE9AF004C3375FBC4" bold="true" box="[760,795,1127,1151]" pageId="4" pageNumber="405">(A)</emphasis>
Dorsal and 
<emphasis id="B917EAACFFFBFFBFE8C1004C33D6FBC4" bold="true" box="[918,952,1127,1151]" pageId="4" pageNumber="405">(B)</emphasis>
ventral views.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFFBFFBFEB9000F933D7F9B7" blockId="4.[151,1437,1234,1944]" pageId="4" pageNumber="405">
Forelimbs not hypertrophied; upper arm thinner than forearm. Ulnar tubercles absent. Fingers fringed; relative finger length II&lt;III&lt;V&lt;IV (
<figureCitation id="13582A3BFFFBFFBFEA9B00DD3248FAAB" box="[460,550,1270,1296]" captionStart="FIGURE 3" captionStartId="5.[151,250,1150,1174]" captionTargetBox="[202,1385,181,1127]" captionTargetId="figure-18@5.[202,1385,181,1127]" captionTargetPageId="5" captionText="FIGURE 3. Scinax ushiniauae, holotype (CRBIIAP 2912). (A) Left hand and (B) left foot in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10628014" httpUri="https://zenodo.org/record/10628014/files/figure.png" pageId="4" pageNumber="405">Fig. 3A</figureCitation>
). Discs elliptical, wider than long, 3FD 33% ED and 85% TD; disc of Finger II smaller than others. Subarticular tubercles single, conical on fingers II and III, rounded and quadrangular on fingers IV and V, respectively. Supernumerary tubercles small, single, and rounded. Inner metacarpal tubercle single, flattened, and elliptical; outer metacarpal tubercle bifid and flattened. Basal webbing between fingers, more reduced between fingers II and III. Nuptial pad present on Finger II, thickened, light-colored, covering the metacarpal II dorsomedially, extending from the base of the inner metacarpal tubercle, obscuring its proximal and outer margins to distal margin of the subarticular tubercle ventrally, with distinct epidermal projections. Nuptial pad of finger III covers the metacarpal III dorsomedially, without epidermal projections.
</paragraph>
<paragraph id="8BDC36BEFFFBFFBFEB90023D35E2F95F" blockId="4.[151,1437,1234,1944]" pageId="4" pageNumber="405">
Hind limbs proportionally long, TL 50% SVL, THL 45% SVL, FL 41% SVL. Tarsal fold and tubercles absent. Tubercles absent on heel. Toes fringed; relative toe length I&lt;II&lt;III≈V&lt;IV (
<figureCitation id="13582A3BFFFBFFBFE89A0211344DF9EF" box="[973,1059,1594,1620]" captionStart="FIGURE 3" captionStartId="5.[151,250,1150,1174]" captionTargetBox="[202,1385,181,1127]" captionTargetId="figure-18@5.[202,1385,181,1127]" captionTargetPageId="5" captionText="FIGURE 3. Scinax ushiniauae, holotype (CRBIIAP 2912). (A) Left hand and (B) left foot in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10628014" httpUri="https://zenodo.org/record/10628014/files/figure.png" pageId="4" pageNumber="405">Fig. 3B</figureCitation>
). Discs elliptical, wider than long, 4TD 27% ED and 69% TD. Subarticular tubercles single, conical on Toe I, and rounded on toes II–V. Supernumerary tubercles small, single, and rounded. Inner metatarsal tubercle single and elliptical; outer metatarsal tubercle single and elliptical (right toe) or round (left toe), three times smaller than the inner metatarsal tubercle. Webbing formula I 2 
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– 
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II 
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– 
<superScript id="7C169BF6FFFBFFBFEA1602E1310DF963" attach="left" box="[321,355,1738,1764]" fontSize="6" pageId="4" pageNumber="405">21/2</superScript>
III 
<superScript id="7C169BF6FFFBFFBFEADD02E131C2F963" attach="none" box="[394,428,1738,1764]" fontSize="6" pageId="4" pageNumber="405">11/2</superScript>
– 
<superScript id="7C169BF6FFFBFFBFEAEC02E131B2F963" attach="left" box="[443,476,1738,1764]" fontSize="6" pageId="4" pageNumber="405">21/3</superScript>
IV 
<superScript id="7C169BF6FFFBFFBFE95202E13249F963" attach="none" box="[517,551,1738,1764]" fontSize="6" pageId="4" pageNumber="405">22/3</superScript>
–1 
<superScript id="7C169BF6FFFBFFBFE91302E13223F963" attach="left" box="[580,589,1738,1752]" fontSize="6" pageId="4" pageNumber="405">+</superScript>
V (
<figureCitation id="13582A3BFFFBFFBFE92102E132A0F95F" box="[630,718,1738,1764]" captionStart="FIGURE 3" captionStartId="5.[151,250,1150,1174]" captionTargetBox="[202,1385,181,1127]" captionTargetId="figure-18@5.[202,1385,181,1127]" captionTargetPageId="5" captionText="FIGURE 3. Scinax ushiniauae, holotype (CRBIIAP 2912). (A) Left hand and (B) left foot in ventral view." figureDoi="http://doi.org/10.5281/zenodo.10628014" httpUri="https://zenodo.org/record/10628014/files/figure.png" pageId="4" pageNumber="405">Fig. 3B</figureCitation>
). Cloacal opening directed posteriorly at upper level of thighs.
</paragraph>
<paragraph id="8BDC36BEFFFBFFBFEB9002C53466F823" blockId="4.[151,1437,1234,1944]" pageId="4" pageNumber="405">Skin texture on dorsal surface of head, loreal region, upper eyelid, and tympanum region shagreen; small, flat, and scattered tubercles present on dorsal surface of trunk and flanks. Skin smooth on dorsal surface of limbs, gular and inguinal regions, and on hidden and ventral surfaces of forelimbs, tibiae, and tarsi. Low granules on abdomen, ventral surface of thighs, and subcloacal region. Glandular concentrations or spicule-shaped papillary epidermal projections absent on inner margin of forelimbs, pectoral and inguinal regions.</paragraph>
<caption id="DF1C6636FFFAFFBEEBC0005534BFFB2C" ID-DOI="http://doi.org/10.5281/zenodo.10628014" ID-Zenodo-Dep="10628014" box="[151,1233,1150,1175]" httpUri="https://zenodo.org/record/10628014/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,250,1150,1174]" targetBox="[202,1385,181,1127]" targetPageId="5" targetType="figure">
<paragraph id="8BDC36BEFFFAFFBEEBC0005534BFFB2C" blockId="5.[151,1233,1150,1175]" box="[151,1233,1150,1175]" pageId="5" pageNumber="406">
<emphasis id="B917EAACFFFAFFBEEBC00055317DFB2C" bold="true" box="[151,275,1150,1175]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
<taxonomicName id="4C634D3DFFFAFFBEEA4E005431BCFB2C" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[281,466,1151,1175]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFFAFFBEEA4E005431BCFB2C" box="[281,466,1151,1175]" italics="true" pageId="5" pageNumber="406">Scinax ushiniauae</emphasis>
</taxonomicName>
, holotype (CRBIIAP 2912). 
<emphasis id="B917EAACFFFAFFBEE9AF00553375FB2D" bold="true" box="[760,795,1150,1174]" pageId="5" pageNumber="406">(A)</emphasis>
Left hand and 
<emphasis id="B917EAACFFFAFFBEE8E3005533B8FB2D" bold="true" box="[948,982,1150,1174]" pageId="5" pageNumber="406">(B)</emphasis>
left foot in ventral view.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFFAFFBEEB9000C23236FA9C" blockId="5.[151,1437,1257,1859]" pageId="5" pageNumber="406">
<emphasis id="B917EAACFFFAFFBEEB9000C232F3FAB8" bold="true" box="[199,669,1257,1283]" pageId="5" pageNumber="406">
Measurements of the 
<typeStatus id="54D8881CFFFAFFBEEA9E00C2325EFAB8" box="[457,560,1257,1283]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
SVL 20.6; HL 8.1; HW 7.2; IND 1.9; IOD 2.3; ED 3.3; EN 2.5; TD 1.3; TL 10.4; FL 9.7; 3FD 1.1; 4TD 0.9.
</paragraph>
<paragraph id="8BDC36BEFFFAFFBEEB90011A3392F96C" blockId="5.[151,1437,1257,1859]" pageId="5" pageNumber="406">
<emphasis id="B917EAACFFFAFFBEEB90011A3248FAF0" bold="true" box="[199,550,1329,1355]" pageId="5" pageNumber="406">
Color in life of the 
<typeStatus id="54D8881CFFFAFFBEEAEE011A324CFAF0" box="[441,546,1329,1355]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsum golden light brown with scattered black or dark brown blotches of different sizes (
<figureCitation id="13582A3BFFFAFFBEEA10017E31AAFAD4" box="[327,452,1365,1391]" captionStart="FIGURE 4" captionStartId="6.[151,250,1407,1431]" captionTargetBox="[158,1430,181,1383]" captionTargetId="figure-24@6.[158,1430,181,1383]" captionTargetPageId="6" captionText="FIGURE 4. Scinax ushiniauae, holotype in life (CRBIIAP 2912). Body in (A) dorsolateral, (B) anterior, and (C) ventral views." figureDoi="http://doi.org/10.5281/zenodo.10628016" httpUri="https://zenodo.org/record/10628016/files/figure.png" pageId="5" pageNumber="406">Fig. 4A, B</figureCitation>
). Canthus rostralis with a diffuse black line, incomplete towards the nostril. Upper lip light brown or cream, with diffuse dark brown blotches below the eyes. Dorsal surfaces of forelimbs and hindlimbs conspicuously spotted with irregular transverse dark brown markings on forearms and dark brown blotches on shanks. Supracloacal region and knees with a light golden blotch dorsally. Flanks light yellow with a dark brown lateral irregular line from the posterior corner of the eye and through the upper part of the tympanic membrane and supratympanic fold to the inguinal region. Hidden surfaces of thighs, shanks, and inguinal region uniformly brown. Ventrally, fore- and hindlimbs light green. Soles and legs conspicuously spotted with brown. Palms, margins of gular region, area around arm insertion, and lateral edges of arms, thighs, and shanks finely spotted with brown. Pectoral region and abdomen white, immaculate; gular region yellow (
<figureCitation id="13582A3BFFFAFFBEE806025E33C5F934" box="[849,939,1653,1679]" captionStart="FIGURE 4" captionStartId="6.[151,250,1407,1431]" captionTargetBox="[158,1430,181,1383]" captionTargetId="figure-24@6.[158,1430,181,1383]" captionTargetPageId="6" captionText="FIGURE 4. Scinax ushiniauae, holotype in life (CRBIIAP 2912). Body in (A) dorsolateral, (B) anterior, and (C) ventral views." figureDoi="http://doi.org/10.5281/zenodo.10628016" httpUri="https://zenodo.org/record/10628016/files/figure.png" pageId="5" pageNumber="406">Fig. 4C</figureCitation>
). Iris silvery bronze with black reticulations and median horizontal red streak; red coloration also bordering the pupil; black blotches on the sides of the pupil. Physiological chlorosis present (bluish-green coloration in bones and lymph).
</paragraph>
<paragraph id="8BDC36BEFFFAFFBEEB9002CA31B5F8F8" blockId="5.[151,1437,1257,1859]" pageId="5" pageNumber="406">
<emphasis id="B917EAACFFFAFFBEEB9002CA32E7F940" bold="true" box="[199,649,1761,1787]" pageId="5" pageNumber="406">
Color in preservative of the 
<typeStatus id="54D8881CFFFAFFBEE94B02CA32EBF940" box="[540,645,1761,1787]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
.
</emphasis>
The coloration pattern is like that of the specimen in life, but darker. Golden tones faded and disappeared. Yellow, light green, and white colors became light beige or cream white. The iris coloration became black.
</paragraph>
<caption id="DF1C6636FFF9FFBDEBC0015430B7FA00" ID-DOI="http://doi.org/10.5281/zenodo.10628016" ID-Zenodo-Dep="10628016" httpUri="https://zenodo.org/record/10628016/files/figure.png" pageId="6" pageNumber="407" startId="6.[151,250,1407,1431]" targetBox="[158,1430,181,1383]" targetPageId="6" targetType="figure">
<paragraph id="8BDC36BEFFF9FFBDEBC0015430B7FA00" blockId="6.[151,1436,1407,1467]" pageId="6" pageNumber="407">
<emphasis id="B917EAACFFF9FFBDEBC001543179FA2C" bold="true" box="[151,279,1407,1431]" pageId="6" pageNumber="407">FIGURE 4.</emphasis>
<taxonomicName id="4C634D3DFFF9FFBDEA7601AB31B0FA2C" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[289,478,1407,1431]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF9FFBDEA7601AB31B0FA2C" box="[289,478,1407,1431]" italics="true" pageId="6" pageNumber="407">Scinax ushiniauae</emphasis>
</taxonomicName>
, holotype in life (CRBIIAP 2912). Body in 
<emphasis id="B917EAACFFF9FFBDE8EC015433B0FA2C" bold="true" box="[955,990,1407,1431]" pageId="6" pageNumber="407">(A)</emphasis>
dorsolateral, 
<emphasis id="B917EAACFFF9FFBDEF27015434FCFA2C" bold="true" box="[1136,1170,1407,1431]" pageId="6" pageNumber="407">(B)</emphasis>
anterior, and 
<emphasis id="B917EAACFFF9FFBDEE7E01543522FA2C" bold="true" box="[1321,1356,1407,1431]" pageId="6" pageNumber="407">(C)</emphasis>
ventral views.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFF9FFBDEB9002263339F80F" blockId="6.[151,1437,1549,1972]" pageId="6" pageNumber="407">
<emphasis id="B917EAACFFF9FFBDEB900226327CF99C" bold="true" box="[199,530,1549,1576]" pageId="6" pageNumber="407">Variation in the type series.</emphasis>
Some measurements are presented in 
<tableCitation id="C6E10305FFF9FFBDE89202253474F993" box="[965,1050,1550,1576]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionTargetPageId="7" captionText="TABLE 1. Measurements (in mm) of the type series of Scinax ushiniauae (including the holotype). See Material and methods for abbreviations." httpUri="http://table.plazi.org/id/DF1C6636FFF8FFBCEBC004BC31DBFF68" pageId="6" pageNumber="407" tableUuid="DF1C6636FFF8FFBCEBC004BC31DBFF68">Table 1</tableCitation>
. Vomerine teeth in two transverse series barely separated or separated by a space equivalent to half the length of the dentigerous process. Vomerine teeth 3–5 on the right side and 2–7 on the left side. Nuptial pad on finger II like that of the 
<typeStatus id="54D8881CFFF9FFBDEF87027D355CF9CB" box="[1232,1330,1622,1648]" pageId="6" pageNumber="407" type="holotype">holotype</typeStatus>
in 
<specimenCount id="9D65FD37FFF9FFBDEE0B027D3162F92F" count="7" pageId="6" pageNumber="407" type="generic">seven specimens</specimenCount>
(
<figureCitation id="13582A3BFFF9FFBDEA4C025131F8F92F" box="[283,406,1658,1685]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1582]" captionTargetBox="[196,1390,812,1535]" captionTargetId="figure-159@7.[196,1390,812,1535]" captionTargetPageId="7" captionText="FIGURE 5. Nuptial pad on fingers II and III of the left hand of Scinax ushiniauae, paratype (MACN 59716). Hand in (A) dorsal (digitally reversed) and (B) ventral views. The arrow indicates the glandular acini of the nuptial pad on finger III." figureDoi="http://doi.org/10.5281/zenodo.10628018" httpUri="https://zenodo.org/record/10628018/files/figure.png" pageId="6" pageNumber="407">Fig. 5A, B</figureCitation>
), thinner in 
<specimenCount id="9D65FD37FFF9FFBDE949025132ABF92F" box="[542,709,1658,1684]" count="2" pageId="6" pageNumber="407" type="generic">two specimens</specimenCount>
(CRBIIAP 1656, MCP 14564), and covering approximately half of metacarpal II dorsomedial surface in 
<specimenCount id="9D65FD37FFF9FFBDE90602B5336BF903" box="[593,773,1694,1720]" count="3" pageId="6" pageNumber="407" type="generic">three specimens</specimenCount>
(CRBIIAP 1595, 1556, MCP 14562). Toe webbing formula varies as follows: I 2 
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– (
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– 
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) II (
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–1 
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) – (
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– 
<superScript id="7C169BF6FFF9FFBDE98802E9336FF96B" attach="none" box="[735,769,1730,1756]" fontSize="6" pageId="6" pageNumber="407">21/4</superScript>
) III (
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–1 
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) – (2 
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– 2) IV (
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–2 
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) – (
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–1) V. Skin texture on dorsum smooth or shagreen; smooth or with small, flat, and scattered tubercles on trunk and flanks; smooth or areolate on vocal sac and pectoral regions. The only available mature female (CRBIIAP 1712) is larger than males (
<quantity id="4C9B9B5BFFF9FFBDEBBF03053120F8F3" box="[232,334,1838,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="6" pageNumber="407" unit="mm" value="27.3">27.3 mm</quantity>
versus 
<quantity id="4C9B9B5BFFF9FFBDEAF203053227F8F3" box="[421,585,1838,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.155" metricValueMax="2.29" metricValueMin="2.02" pageId="6" pageNumber="407" unit="mm" value="21.549999999999997" valueMax="22.9" valueMin="20.2">20.2–22.9 mm</quantity>
in males) and has webbing toe formula I 2 
<superScript id="7C169BF6FFF9FFBDEF7803053456F887" attach="none" box="[1071,1080,1838,1852]" fontSize="6" pageId="6" pageNumber="407">+</superScript>
– 
<superScript id="7C169BF6FFF9FFBDEF1003053407F887" attach="left" box="[1095,1129,1838,1864]" fontSize="6" pageId="6" pageNumber="407">21/4</superScript>
II 1–2 
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III 1–2 
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IV 2 
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–1 V. The horizontal pupil contracts extensively along the predominant (horizontal) and non predominant axis (vertical), generating in the most extreme contraction that we have documented (
<figureCitation id="13582A3BFFF9FFBDE8EB035D347FF82B" box="[956,1041,1910,1936]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="6" pageNumber="407">Fig. 6F</figureCitation>
) a pupil that retains the horizontal shape, with an evident inflexion point along its ventral margin.
</paragraph>
<caption id="DF1C6636FFF8FFBCEBC004BC31DBFF68" ID-Table-UUID="DF1C6636FFF8FFBCEBC004BC31DBFF68" httpUri="http://table.plazi.org/id/DF1C6636FFF8FFBCEBC004BC31DBFF68" pageId="7" pageNumber="408" startId="7.[151,239,151,175]" targetBox="[159,1295,228,785]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="8BDC36BEFFF8FFBCEBC004BC31DBFF68" blockId="7.[151,1437,151,211]" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCEBC004BC3162FF14" bold="true" box="[151,268,151,175]" pageId="7" pageNumber="408">TABLE 1.</emphasis>
Measurements (in mm) of the type series of 
<taxonomicName id="4C634D3DFFF8FFBCE85704B333ABFF14" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[768,965,151,175]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF8FFBCE85704B333ABFF14" box="[768,965,151,175]" italics="true" pageId="7" pageNumber="408">Scinax ushiniauae</emphasis>
</taxonomicName>
(including the holotype). See Material and methods for abbreviations.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFF8FFBCEBC804CF34FEFCAA" pageId="7" pageNumber="408">
<table id="F963C41EFFF80044EBC804CF3561FCAA" box="[159,1295,228,785]" gridcols="4" gridrows="14" pageId="7" pageNumber="408">
<tr id="355334FCFFF80044EBC804CF3561FF41" box="[159,1295,228,250]" gridrow="0" pageId="7" pageNumber="408" rowspan-2="1">
<th id="76825D80FFF80044EBC804CF315BFF41" box="[159,309,228,250]" gridcol="0" gridrow="0" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCEBC804CF315BFF42" bold="true" box="[159,309,228,249]" pageId="7" pageNumber="408">Measurements</emphasis>
</th>
<th id="76825D80FFF80044EA8904CF321EFF41" box="[478,624,228,250]" gridcol="1" gridrow="0" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCEA8904CF321EFF41" bold="true" box="[478,624,228,250]" pageId="7" pageNumber="408">
Males (
<emphasis id="B917EAACFFF8FFBCE97D04CE3259FF41" bold="true" box="[554,567,229,250]" italics="true" pageId="7" pageNumber="408">n</emphasis>
= 13)
</emphasis>
</th>
<th id="76825D80FFF80044EF2504CF3561FF41" box="[1138,1295,228,250]" gridcol="3" gridrow="0" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCEF2504CF3561FF41" bold="true" box="[1138,1295,228,250]" pageId="7" pageNumber="408">
Females (
<emphasis id="B917EAACFFF8FFBCEF8304CE348FFF41" bold="true" box="[1236,1249,229,250]" italics="true" pageId="7" pageNumber="408">n</emphasis>
= 1)
</emphasis>
</th>
</tr>
<tr id="355334FCFFF80044EBC805273561FE99" box="[159,1295,268,290]" gridrow="1" pageId="7" pageNumber="408" rowspan-0="1" rowspan-3="1">
<td id="76825D80FFF80044EA890527321EFE99" box="[478,624,268,290]" gridcol="1" gridrow="1" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCEA890527323FFE99" bold="true" box="[478,593,268,290]" pageId="7" pageNumber="408">Mean ± SD</emphasis>
</td>
<td id="76825D80FFF80044E878052733E1FE99" box="[815,911,268,290]" gridcol="2" gridrow="1" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCE8780527331EFE9A" bold="true" box="[815,880,268,289]" pageId="7" pageNumber="408">Range</emphasis>
</td>
</tr>
<tr id="355334FCFFF80044EBC8051E3561FEF0" box="[159,1295,309,331]" gridrow="2" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC8051E315BFEF0" box="[159,309,309,331]" gridcol="0" gridrow="2" pageId="7" pageNumber="408">SVL</th>
<td id="76825D80FFF80044EA89051E321EFEF0" box="[478,624,309,331]" gridcol="1" gridrow="2" pageId="7" pageNumber="408">21.5 ± 0.8</td>
<td id="76825D80FFF80044E878051E33E1FEF0" box="[815,911,309,331]" gridcol="2" gridrow="2" pageId="7" pageNumber="408">20.2–22.9</td>
<td id="76825D80FFF80044EF25051E3561FEF0" box="[1138,1295,309,331]" gridcol="3" gridrow="2" pageId="7" pageNumber="408">27.3</td>
</tr>
<tr id="355334FCFFF80044EBC805753561FECF" box="[159,1295,350,372]" gridrow="3" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC80575315BFECF" box="[159,309,350,372]" gridcol="0" gridrow="3" pageId="7" pageNumber="408">HL</th>
<td id="76825D80FFF80044EA890575321EFECF" box="[478,624,350,372]" gridcol="1" gridrow="3" pageId="7" pageNumber="408">8.0 ± 0.3</td>
<td id="76825D80FFF80044E878057533E1FECF" box="[815,911,350,372]" gridcol="2" gridrow="3" pageId="7" pageNumber="408">7.6–8.8</td>
<td id="76825D80FFF80044EF2505753561FECF" box="[1138,1295,350,372]" gridcol="3" gridrow="3" pageId="7" pageNumber="408">10.9</td>
</tr>
<tr id="355334FCFFF80044EBC805A33561FE25" box="[159,1295,392,414]" gridrow="4" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC805A3315BFE25" box="[159,309,392,414]" gridcol="0" gridrow="4" pageId="7" pageNumber="408">HW</th>
<td id="76825D80FFF80044EA8905A3321EFE25" box="[478,624,392,414]" gridcol="1" gridrow="4" pageId="7" pageNumber="408">7.5 ± 0.3</td>
<td id="76825D80FFF80044E87805A333E1FE25" box="[815,911,392,414]" gridcol="2" gridrow="4" pageId="7" pageNumber="408">
7.0 
<emphasis id="B917EAACFFF8FFBCE81A05A33337FE26" box="[845,857,392,413]" italics="true" pageId="7" pageNumber="408">–</emphasis>
7.9
</td>
<td id="76825D80FFF80044EF2505A33561FE25" box="[1138,1295,392,414]" gridcol="3" gridrow="4" pageId="7" pageNumber="408">9.6</td>
</tr>
<tr id="355334FCFFF80044EBC8059A3561FE7C" box="[159,1295,433,455]" gridrow="5" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC8059A315BFE7C" box="[159,309,433,455]" gridcol="0" gridrow="5" pageId="7" pageNumber="408">IND</th>
<td id="76825D80FFF80044EA89059A321EFE7C" box="[478,624,433,455]" gridcol="1" gridrow="5" pageId="7" pageNumber="408">1.9 ± 0.3</td>
<td id="76825D80FFF80044E878059A33E1FE7C" box="[815,911,433,455]" gridcol="2" gridrow="5" pageId="7" pageNumber="408">
1.6 
<emphasis id="B917EAACFFF8FFBCE81A05993337FE7C" box="[845,857,434,455]" italics="true" pageId="7" pageNumber="408">–</emphasis>
2.7
</td>
<td id="76825D80FFF80044EF25059A3561FE7C" box="[1138,1295,433,455]" gridcol="3" gridrow="5" pageId="7" pageNumber="408">2.3</td>
</tr>
<tr id="355334FCFFF80044EBC805F13561FE4B" box="[159,1295,474,496]" gridrow="6" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC805F1315BFE4B" box="[159,309,474,496]" gridcol="0" gridrow="6" pageId="7" pageNumber="408">IOD</th>
<td id="76825D80FFF80044EA8905F1321EFE4B" box="[478,624,474,496]" gridcol="1" gridrow="6" pageId="7" pageNumber="408">2.7 ± 0.2</td>
<td id="76825D80FFF80044E87805F133E1FE4B" box="[815,911,474,496]" gridcol="2" gridrow="6" pageId="7" pageNumber="408">
2.3 
<emphasis id="B917EAACFFF8FFBCE81A05F03337FE4B" box="[845,857,475,496]" italics="true" pageId="7" pageNumber="408">–</emphasis>
2.9
</td>
<td id="76825D80FFF80044EF2505F13561FE4B" box="[1138,1295,474,496]" gridcol="3" gridrow="6" pageId="7" pageNumber="408">3.6</td>
</tr>
<tr id="355334FCFFF80044EBC8062F3561FDA2" box="[159,1295,516,537]" gridrow="7" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC8062F315BFDA2" box="[159,309,516,537]" gridcol="0" gridrow="7" pageId="7" pageNumber="408">ED</th>
<td id="76825D80FFF80044EA89062F321EFDA2" box="[478,624,516,537]" gridcol="1" gridrow="7" pageId="7" pageNumber="408">3.3 ± 0.3</td>
<td id="76825D80FFF80044E878062F33E1FDA2" box="[815,911,516,537]" gridcol="2" gridrow="7" pageId="7" pageNumber="408">
2.8 
<emphasis id="B917EAACFFF8FFBCE81A062F3337FDA2" box="[845,857,516,537]" italics="true" pageId="7" pageNumber="408">–</emphasis>
3.9
</td>
<td id="76825D80FFF80044EF25062F3561FDA2" box="[1138,1295,516,537]" gridcol="3" gridrow="7" pageId="7" pageNumber="408">4.4</td>
</tr>
<tr id="355334FCFFF80044EBC806063561FDF8" box="[159,1295,557,579]" gridrow="8" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC80606315BFDF8" box="[159,309,557,579]" gridcol="0" gridrow="8" pageId="7" pageNumber="408">EN</th>
<td id="76825D80FFF80044EA890606321EFDF8" box="[478,624,557,579]" gridcol="1" gridrow="8" pageId="7" pageNumber="408">2.6 ± 0.1</td>
<td id="76825D80FFF80044E878060633E1FDF8" box="[815,911,557,579]" gridcol="2" gridrow="8" pageId="7" pageNumber="408">
2.5 
<emphasis id="B917EAACFFF8FFBCE81A06053337FDF8" box="[845,857,558,579]" italics="true" pageId="7" pageNumber="408">–</emphasis>
2.9
</td>
<td id="76825D80FFF80044EF2506063561FDF8" box="[1138,1295,557,579]" gridcol="3" gridrow="8" pageId="7" pageNumber="408">3.4</td>
</tr>
<tr id="355334FCFFF80044EBC8067D3561FDD7" box="[159,1295,598,620]" gridrow="9" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC8067D315BFDD7" box="[159,309,598,620]" gridcol="0" gridrow="9" pageId="7" pageNumber="408">TD</th>
<td id="76825D80FFF80044EA89067D321EFDD7" box="[478,624,598,620]" gridcol="1" gridrow="9" pageId="7" pageNumber="408">1.3 ± 0.2</td>
<td id="76825D80FFF80044E878067D33E1FDD7" box="[815,911,598,620]" gridcol="2" gridrow="9" pageId="7" pageNumber="408">
1.1 
<emphasis id="B917EAACFFF8FFBCE81A067C3337FDD7" box="[845,857,599,620]" italics="true" pageId="7" pageNumber="408">–</emphasis>
1.8
</td>
<td id="76825D80FFF80044EF25067D3561FDD7" box="[1138,1295,598,620]" gridcol="3" gridrow="9" pageId="7" pageNumber="408">1.7</td>
</tr>
<tr id="355334FCFFF80044EBC806AB3561FD2D" box="[159,1295,640,662]" gridrow="10" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC806AB315BFD2D" box="[159,309,640,662]" gridcol="0" gridrow="10" pageId="7" pageNumber="408">TL</th>
<td id="76825D80FFF80044EA8906AB321EFD2D" box="[478,624,640,662]" gridcol="1" gridrow="10" pageId="7" pageNumber="408">10.8 ± 0.4</td>
<td id="76825D80FFF80044E87806AB33E1FD2D" box="[815,911,640,662]" gridcol="2" gridrow="10" pageId="7" pageNumber="408">
10.4 
<emphasis id="B917EAACFFF8FFBCE80E06AB330BFD2E" box="[857,869,640,661]" italics="true" pageId="7" pageNumber="408">–</emphasis>
11.6
</td>
<td id="76825D80FFF80044EF2506AB3561FD2D" box="[1138,1295,640,662]" gridcol="3" gridrow="10" pageId="7" pageNumber="408">14.0</td>
</tr>
<tr id="355334FCFFF80044EBC806823561FD04" box="[159,1295,681,703]" gridrow="11" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC80682315BFD04" box="[159,309,681,703]" gridcol="0" gridrow="11" pageId="7" pageNumber="408">FL</th>
<td id="76825D80FFF80044EA890682321EFD04" box="[478,624,681,703]" gridcol="1" gridrow="11" pageId="7" pageNumber="408">8.9 ± 0.5</td>
<td id="76825D80FFF80044E878068233E1FD04" box="[815,911,681,703]" gridcol="2" gridrow="11" pageId="7" pageNumber="408">
8.3 
<emphasis id="B917EAACFFF8FFBCE81A06813337FD04" box="[845,857,682,703]" italics="true" pageId="7" pageNumber="408">–</emphasis>
9.6
</td>
<td id="76825D80FFF80044EF2506823561FD04" box="[1138,1295,681,703]" gridcol="3" gridrow="11" pageId="7" pageNumber="408">12.3</td>
</tr>
<tr id="355334FCFFF80044EBC806F93561FD53" box="[159,1295,722,744]" gridrow="12" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC806F9315BFD53" box="[159,309,722,744]" gridcol="0" gridrow="12" pageId="7" pageNumber="408">3FD</th>
<td id="76825D80FFF80044EA8906F9321EFD53" box="[478,624,722,744]" gridcol="1" gridrow="12" pageId="7" pageNumber="408">1.1 ± 0.1</td>
<td id="76825D80FFF80044E87806F933E1FD53" box="[815,911,722,744]" gridcol="2" gridrow="12" pageId="7" pageNumber="408">
0.9 
<emphasis id="B917EAACFFF8FFBCE81A06F83337FD53" box="[845,857,723,744]" italics="true" pageId="7" pageNumber="408">–</emphasis>
1.2
</td>
<td id="76825D80FFF80044EF2506F93561FD53" box="[1138,1295,722,744]" gridcol="3" gridrow="12" pageId="7" pageNumber="408">1.5</td>
</tr>
<tr id="355334FCFFF80044EBC806D03561FCAA" box="[159,1295,763,785]" gridrow="13" pageId="7" pageNumber="408">
<th id="76825D80FFF80044EBC806D0315BFCAA" box="[159,309,763,785]" gridcol="0" gridrow="13" pageId="7" pageNumber="408">4TD</th>
<td id="76825D80FFF80044EA8906D0321EFCAA" box="[478,624,763,785]" gridcol="1" gridrow="13" pageId="7" pageNumber="408">1.1 ± 0.1</td>
<td id="76825D80FFF80044E87806D033E1FCAA" box="[815,911,763,785]" gridcol="2" gridrow="13" pageId="7" pageNumber="408">
0.9 
<emphasis id="B917EAACFFF8FFBCE81A06D73337FCAA" box="[845,857,764,785]" italics="true" pageId="7" pageNumber="408">–</emphasis>
1.3
</td>
<td id="76825D80FFF80044EF2506D03561FCAA" box="[1138,1295,763,785]" gridcol="3" gridrow="13" pageId="7" pageNumber="408">1.6</td>
</tr>
</table>
</paragraph>
<caption id="DF1C6636FFF8FFBCEBC0023D357AF9E9" ID-DOI="http://doi.org/10.5281/zenodo.10628018" ID-Zenodo-Dep="10628018" httpUri="https://zenodo.org/record/10628018/files/figure.png" pageId="7" pageNumber="408" startId="7.[151,250,1558,1582]" targetBox="[196,1390,812,1535]" targetPageId="7" targetType="figure">
<paragraph id="8BDC36BEFFF8FFBCEBC0023D357AF9E9" blockId="7.[151,1436,1558,1619]" pageId="7" pageNumber="408">
<emphasis id="B917EAACFFF8FFBCEBC0023D317CF994" bold="true" box="[151,274,1558,1583]" pageId="7" pageNumber="408">FIGURE 5.</emphasis>
Nuptial pad on fingers II and III of the left hand of 
<taxonomicName id="4C634D3DFFF8FFBCE844023C33A5F994" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[787,971,1559,1583]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF8FFBCE844023C33A5F994" box="[787,971,1559,1583]" italics="true" pageId="7" pageNumber="408">Scinax ushiniauae</emphasis>
</taxonomicName>
, paratype (MACN 59716). Hand in 
<emphasis id="B917EAACFFF8FFBCEE61023D3537F995" bold="true" box="[1334,1369,1558,1582]" pageId="7" pageNumber="408">(A)</emphasis>
dorsal (digitally reversed) and 
<emphasis id="B917EAACFFF8FFBCEADD021131C2F9E9" bold="true" box="[394,428,1594,1618]" pageId="7" pageNumber="408">(B)</emphasis>
ventral views. The arrow indicates the glandular acini of the nuptial pad on finger III.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFF8FFB3EB9002B631DFFEDE" blockId="7.[151,1437,1693,2043]" lastBlockId="8.[151,1436,150,357]" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">
In life, dorsal color golden light brown, brownish gray to dark brown, with scattered or conspicuously distributed dark brown or black blotches of different sizes (see 
<figureCitation id="13582A3BFFF8FFBCE9A402EA3357F960" box="[755,825,1729,1755]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6</figureCitation>
); small light blotches are present on dorsum in most individuals (e.g., CRBIIAP 1851, 1570, 1597, 2905, MCP 14562; 
<figureCitation id="13582A3BFFF8FFBCE83D02CE33D0F944" box="[874,958,1765,1791]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6E</figureCitation>
). Canthal black line present (e.g., CRBIIAP 1597, 1656; 
<figureCitation id="13582A3BFFF8FFBCEA7F032231CCF898" box="[296,418,1801,1827]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6C, E</figureCitation>
), absent (e.g., CRBIIAP 1570, MCP 14562; 
<figureCitation id="13582A3BFFF8FFBCE8F503223393F898" box="[930,1021,1801,1827]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6A</figureCitation>
), or indistinct (e.g., CRBIIAP 1595; 
<figureCitation id="13582A3BFFF8FFBCEBC003063080F8FC" box="[151,238,1837,1863]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6B</figureCitation>
); when present, it covers the canthus rostralis completely (e.g., CRBIIAP 1587, MACN 59716; 
<figureCitation id="13582A3BFFF8FFBCEE7503063518F8FC" box="[1314,1398,1837,1863]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6E</figureCitation>
) or just a small portion of it (e.g., CRBIIAP 1656; 
<figureCitation id="13582A3BFFF8FFBCE9E7037A3366F8D0" box="[688,776,1873,1899]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6C</figureCitation>
). Number of transverse dark brown markings on forearms (e.g., 2–3 markings) and blotches on shanks (few or many blotches) variable among individuals; transversal dark brown markings on thighs present in some individuals (e.g., CRBIIAP 1597, MCP 14565; 
<figureCitation id="13582A3BFFF8FFBCEFF003B2356EF808" box="[1191,1280,1945,1971]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6E</figureCitation>
). Flanks light or dark brown with complete (e.g., CRBIIAP 1570; 
<figureCitation id="13582A3BFFF8FFBCE9B503963357F86C" box="[738,825,1981,2007]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6A</figureCitation>
) or interrupted (e.g., CRBIIAP 2913; 
<figureCitation id="13582A3BFFF8FFBCEFB103963554F86C" box="[1254,1338,1981,2007]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="7" pageNumber="408">Fig. 6F</figureCitation>
), lateral, dark brown to black irregular line, extending from the posterior corner of the eye to the upper part of the tympanic membrane and supratympanic fold (e.g., CRBIIAP 1851, MCP 15108), to the middle of the flanks (e.g., CRBIIAP 1595, 1712, 1905; 
<figureCitation id="13582A3BFFF7FFB3EA3D0490318CFF6E" box="[362,482,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409">Fig. 6B, D</figureCitation>
), or the groin (e.g., CRBIIAP 1570, 1656; 
<figureCitation id="13582A3BFFF7FFB3E89704903454FF6E" box="[960,1082,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409">Fig. 6A, C</figureCitation>
). Ventrally, fore- and hindlimbs and gular region more conspicuously spotted with brown in MCP 14565; abdomen finely pigmented with dark brown in some individuals (e.g., CRBIIAP 1656, 1712). Iris silvery bronze or bronze, with finely or conspicuously distributed black reticulations. Gular region cream to white, finely spotted with brown on the mental area in the female (CRBIIAP 1712).
</paragraph>
<caption id="DF1C6636FFF7FFB3EBC002B03405F96C" ID-DOI="http://doi.org/10.5281/zenodo.10628020" ID-Zenodo-Dep="10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409" startId="8.[151,250,1691,1715]" targetBox="[170,1416,381,1667]" targetPageId="8" targetType="figure">
<paragraph id="8BDC36BEFFF7FFB3EBC002B03405F96C" blockId="8.[151,1437,1691,1751]" pageId="8" pageNumber="409">
<emphasis id="B917EAACFFF7FFB3EBC002B0317DF908" bold="true" box="[151,275,1691,1715]" pageId="8" pageNumber="409">FIGURE 6.</emphasis>
Paratypes of 
<taxonomicName id="4C634D3DFFF7FFB3EACA02B03238F908" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[413,598,1691,1715]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="409" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF7FFB3EACA02B03238F908" box="[413,598,1691,1715]" italics="true" pageId="8" pageNumber="409">Scinax ushiniauae</emphasis>
</taxonomicName>
in life. 
<emphasis id="B917EAACFFF7FFB3E9F302B032A6F908" bold="true" box="[676,712,1691,1715]" pageId="8" pageNumber="409">(A)</emphasis>
CRBIIAP 1570 (male), 
<emphasis id="B917EAACFFF7FFB3E8E902B0338EF908" bold="true" box="[958,992,1691,1715]" pageId="8" pageNumber="409">(B)</emphasis>
CRBIIAP 1595 (male), 
<emphasis id="B917EAACFFF7FFB3EF8002B03494F908" bold="true" box="[1239,1274,1691,1715]" pageId="8" pageNumber="409">(C)</emphasis>
CRBIIAP 1656 (male), 
<emphasis id="B917EAACFFF7FFB3EBB102943167F96C" bold="true" box="[230,265,1727,1751]" pageId="8" pageNumber="409">(D)</emphasis>
CRBIIAP 1712 (female), 
<emphasis id="B917EAACFFF7FFB3E94302943258F96C" bold="true" box="[532,566,1727,1751]" pageId="8" pageNumber="409">(E)</emphasis>
CRBIIAP 1597 (male), and 
<emphasis id="B917EAACFFF7FFB3E80F02943317F96C" bold="true" box="[856,889,1727,1751]" pageId="8" pageNumber="409">(F)</emphasis>
CRBIIAP 2913 (male).
</paragraph>
</caption>
<paragraph id="8BDC36BEFFF7FFB3EB9003013334F84C" blockId="8.[151,1437,1833,2040]" pageId="8" pageNumber="409">
In preservative, coloration patterns are similar, but some specimens became darker (CRBIIAP 1712; 
<figureCitation id="13582A3BFFF7FFB3EE32030230D2F8DC" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409">Figs. 6D</figureCitation>
, 
<figureCitation id="13582A3BFFF7FFB3EB9F036630B9F8DC" box="[200,215,1869,1895]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1918]" captionTargetId="figure-18@9.[203,1384,181,1918]" captionTargetPageId="9" captionText="FIGURE 7. Paratypes of Scinax ushiniauae in preservative. Color pattern variation of body in (A) dorsal and (B) ventral views. From left to right and top to bottom: CRBIIAP 1851, 1597, 1905, MCP 15108, CRBIIAP 2913, MACN 59716, CRBIIAP 1595, MCP 14562, all males, and CRBIIAP 1712, female. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10628022" httpUri="https://zenodo.org/record/10628022/files/figure.png" pageId="8" pageNumber="409">7</figureCitation>
) or lighter (CRBIIAP 2913; 
<figureCitation id="13582A3BFFF7FFB3E94D03663219F8DC" box="[538,631,1869,1895]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409">Figs. 6F</figureCitation>
, 
<figureCitation id="13582A3BFFF7FFB3E9D5036632FFF8DC" box="[642,657,1869,1895]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1918]" captionTargetId="figure-18@9.[203,1384,181,1918]" captionTargetPageId="9" captionText="FIGURE 7. Paratypes of Scinax ushiniauae in preservative. Color pattern variation of body in (A) dorsal and (B) ventral views. From left to right and top to bottom: CRBIIAP 1851, 1597, 1905, MCP 15108, CRBIIAP 2913, MACN 59716, CRBIIAP 1595, MCP 14562, all males, and CRBIIAP 1712, female. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10628022" httpUri="https://zenodo.org/record/10628022/files/figure.png" pageId="8" pageNumber="409">7</figureCitation>
) after fixation in 10% formalin. Dorsally, dark and white blotches on dorsum disappeared (CRBIIAP 1597; 
<figureCitation id="13582A3BFFF7FFB3E91F035A32C7F830" box="[584,681,1905,1931]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409">Figs. 6E</figureCitation>
, 
<figureCitation id="13582A3BFFF7FFB3E9E1035A32B4F830" box="[694,730,1905,1931]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1918]" captionTargetId="figure-18@9.[203,1384,181,1918]" captionTargetPageId="9" captionText="FIGURE 7. Paratypes of Scinax ushiniauae in preservative. Color pattern variation of body in (A) dorsal and (B) ventral views. From left to right and top to bottom: CRBIIAP 1851, 1597, 1905, MCP 15108, CRBIIAP 2913, MACN 59716, CRBIIAP 1595, MCP 14562, all males, and CRBIIAP 1712, female. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10628022" httpUri="https://zenodo.org/record/10628022/files/figure.png" pageId="8" pageNumber="409">7A</figureCitation>
) or became indistinct (CRBIIAP 1712; 
<figureCitation id="13582A3BFFF7FFB3EFC9035A356CF830" box="[1182,1282,1905,1931]" captionStart="FIGURE 6" captionStartId="8.[151,250,1691,1715]" captionTargetBox="[170,1416,381,1667]" captionTargetId="figure-115@8.[170,1416,381,1667]" captionTargetPageId="8" captionText="FIGURE 6. Paratypes of Scinax ushiniauae in life. (A) CRBIIAP 1570 (male), (B) CRBIIAP 1595 (male), (C) CRBIIAP 1656 (male), (D) CRBIIAP 1712 (female), (E) CRBIIAP 1597 (male), and (F) CRBIIAP 2913 (male)." figureDoi="http://doi.org/10.5281/zenodo.10628020" httpUri="https://zenodo.org/record/10628020/files/figure.png" pageId="8" pageNumber="409">Figs. 6D</figureCitation>
, 
<figureCitation id="13582A3BFFF7FFB3EE58035A355DF830" box="[1295,1331,1905,1931]" captionStart="FIGURE 7" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1918]" captionTargetId="figure-18@9.[203,1384,181,1918]" captionTargetPageId="9" captionText="FIGURE 7. Paratypes of Scinax ushiniauae in preservative. Color pattern variation of body in (A) dorsal and (B) ventral views. From left to right and top to bottom: CRBIIAP 1851, 1597, 1905, MCP 15108, CRBIIAP 2913, MACN 59716, CRBIIAP 1595, MCP 14562, all males, and CRBIIAP 1712, female. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10628022" httpUri="https://zenodo.org/record/10628022/files/figure.png" pageId="8" pageNumber="409">7A</figureCitation>
) in some specimens. Lateral lines on flanks faded and disappeared in the area immediately posterior to tympanum in some specimens (CRBIIAP 1597, 1851, 1905, MCP 15108). Two individuals (CRBIIAP 1851, MCP 15108) turned darker in some regions (head, nostril, and sacral region) after fixation.
</paragraph>
<caption id="DF1C6636FFF6FFB2EBC003BC3315F84C" ID-DOI="http://doi.org/10.5281/zenodo.10628022" ID-Zenodo-Dep="10628022" httpUri="https://zenodo.org/record/10628022/files/figure.png" pageId="9" pageNumber="410" startId="9.[151,250,1943,1967]" targetBox="[203,1384,181,1918]" targetPageId="9" targetType="figure">
<paragraph id="8BDC36BEFFF6FFB2EBC003BC3315F84C" blockId="9.[151,1437,1943,2039]" pageId="9" pageNumber="410">
<emphasis id="B917EAACFFF6FFB2EBC003BC317CF814" bold="true" box="[151,274,1943,1967]" pageId="9" pageNumber="410">FIGURE 7.</emphasis>
Paratypes of 
<taxonomicName id="4C634D3DFFF6FFB2EACC03B3323DF814" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[411,595,1943,1967]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF6FFB2EACC03B3323DF814" box="[411,595,1943,1967]" italics="true" pageId="9" pageNumber="410">Scinax ushiniauae</emphasis>
</taxonomicName>
in preservative. Color pattern variation of body in 
<emphasis id="B917EAACFFF6FFB2EF0603BC341AF814" bold="true" box="[1105,1140,1943,1967]" pageId="9" pageNumber="410">(A)</emphasis>
dorsal and 
<emphasis id="B917EAACFFF6FFB2EFB003BC3567F814" bold="true" box="[1255,1289,1943,1967]" pageId="9" pageNumber="410">(B)</emphasis>
ventral views. From left to right and top to bottom: CRBIIAP 1851, 1597, 1905, MCP 15108, CRBIIAP 2913, MACN 59716, CRBIIAP 1595, MCP 14562, all males, and CRBIIAP 1712, female. Scale bars = 10 mm.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFF5FFB1EB9004BD32E6FE16" blockId="10.[151,1437,150,429]" pageId="10" pageNumber="411">
<emphasis id="B917EAACFFF5FFB1EB9004BD3290FF0B" bold="true" box="[199,766,150,177]" pageId="10" pageNumber="411">Advertisement call (mean ± SD in parentheses).</emphasis>
The advertisement call of 
<taxonomicName id="4C634D3DFFF5FFB1EF7D04BC34AFFF0A" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[1066,1217,151,177]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF5FFB1EF7D04BC34AFFF0A" box="[1066,1217,151,177]" italics="true" pageId="10" pageNumber="411">S. ushiniauae</emphasis>
</taxonomicName>
consists of a single pulsed note, with pulses of high amplitude preceded by pulses with lower energy (
<figureCitation id="13582A3BFFF5FFB1EF7104903407FF6E" box="[1062,1129,187,213]" captionStart="FIGURE 8" captionStartId="10.[151,250,1593,1617]" captionTargetBox="[205,1383,466,1556]" captionTargetId="figure-168@10.[192,1396,453,1569]" captionTargetPageId="10" captionText="FIGURE 8. Advertisement call of Scinax ushiniauae (CRBIIAP 1905, paratype, FZ-SOUND-CODE: 14638) from Frontera Village, Soplín, Requena, Loreto, Peru. Recorded at 19:30 h and 24.5°C. (A)Waveform of five call series (4.5 s). (B) Spectrogram and oscillogram of the third call series from A. (C) Oscillogram of the second note from B." figureDoi="http://doi.org/10.5281/zenodo.10628024" httpUri="https://zenodo.org/record/10628024/files/figure.png" pageId="10" pageNumber="411">Fig. 8</figureCitation>
). Calls are emitted in series (n= 79 out of 88 analyzed calls) or as isolated notes (n = 9). Series are composed of 2–5 calls per series (2.6±0.2) with a call interval of 70–160 ms (60±0) and a duration of 90–380 ms (200±6). The individual call duration is 4–62 ms (29±14), and each is composed of 1–3 pulses (1.7±0.4) of 1–33 ms (9±2). In cases in which calls include two or more pulses (44 out of 88), the pulse rate is 45.3–69.0 pulses/s (58.2±6.7) with a pulse interval of 10–210 s (20±0). The dominant frequency is 
<geoCoordinate id="EE575079FFF5FFB1EA9B05443264FE32" box="[460,522,367,393]" degrees="2,713" direction="north" orientation="latitude" pageId="10" pageNumber="411" precision="55" value="2.713">2,713</geoCoordinate>
<geoCoordinate id="EE575079FFF5FFB1E95C05443236FE32" box="[523,600,367,393]" degrees="4,219" direction="west" orientation="longitude" pageId="10" pageNumber="411" precision="55" value="-4.219">–4,219</geoCoordinate>
Hz (3,382±238), the 95% frequency 
<geoCoordinate id="EE575079FFF5FFB1E8AA05443452FE32" box="[1021,1084,367,393]" degrees="3,704" direction="north" orientation="latitude" pageId="10" pageNumber="411" precision="55" value="3.704">3,704</geoCoordinate>
<geoCoordinate id="EE575079FFF5FFB1EF6C054434E7FE32" box="[1083,1161,367,393]" degrees="6,460" direction="west" orientation="longitude" pageId="10" pageNumber="411" precision="55" value="-6.46">–6,460</geoCoordinate>
Hz (4729±516), and the 25% frequency 
<geoCoordinate id="EE575079FFF5FFB1EA1F05B831E6FE16" box="[328,392,403,429]" degrees="2,627" direction="north" orientation="latitude" pageId="10" pageNumber="411" precision="55" value="2.627">2,627</geoCoordinate>
<geoCoordinate id="EE575079FFF5FFB1EADF05B831BBFE16" box="[392,469,403,429]" degrees="3,141" direction="west" orientation="longitude" pageId="10" pageNumber="411" precision="55" value="-3.141">–3,141</geoCoordinate>
Hz (2,887±29).
</paragraph>
<caption id="DF1C6636FFF5FFB1EBC00212345CF923" ID-DOI="http://doi.org/10.5281/zenodo.10628024" ID-Zenodo-Dep="10628024" httpUri="https://zenodo.org/record/10628024/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,250,1593,1617]" targetBox="[205,1383,466,1556]" targetPageId="10" targetType="figure">
<paragraph id="8BDC36BEFFF5FFB1EBC00212345CF923" blockId="10.[151,1436,1593,1689]" pageId="10" pageNumber="411">
<emphasis id="B917EAACFFF5FFB1EBC00212317BF9EA" bold="true" box="[151,277,1593,1617]" pageId="10" pageNumber="411">FIGURE 8.</emphasis>
Advertisement call of 
<taxonomicName id="4C634D3DFFF5FFB1E952021232AEF9EA" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[517,704,1593,1617]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF5FFB1E952021232AEF9EA" box="[517,704,1593,1617]" italics="true" pageId="10" pageNumber="411">Scinax ushiniauae</emphasis>
</taxonomicName>
(CRBIIAP 1905, paratype, FZ-SOUND-CODE: 14638) from Frontera Village, Soplín, Requena, Loreto, Peru. Recorded at 19:30 h and 24.5°C. 
<emphasis id="B917EAACFFF5FFB1E83E027633E3F9CE" bold="true" box="[873,909,1629,1653]" pageId="10" pageNumber="411">(A)</emphasis>
Waveform of five call series (4.5 s). 
<emphasis id="B917EAACFFF5FFB1EFA202763579F9CE" bold="true" box="[1269,1303,1629,1653]" pageId="10" pageNumber="411">(B)</emphasis>
Spectrogram and oscillogram of the third call series from A. 
<emphasis id="B917EAACFFF5FFB1E92002AA32F4F922" bold="true" box="[631,666,1665,1689]" pageId="10" pageNumber="411">(C)</emphasis>
Oscillogram of the second note from B.
</paragraph>
</caption>
<paragraph id="8BDC36BEFFF5FFB1EB9002C83190F8D2" blockId="10.[151,1437,1763,2042]" pageId="10" pageNumber="411">
<emphasis id="B917EAACFFF5FFB1EB9002C83103F946" bold="true" box="[199,365,1763,1789]" pageId="10" pageNumber="411">Comparisons.</emphasis>
Based on the phylogenetic hypothesis of 
<bibRefCitation id="EFF24B4FFFF5FFB1E86202C8340EF946" author="Araujo-Vieira, K. &amp; Lourenco, A. C. C. &amp; Lacerda, J. V. A. &amp; Lyra, M. L. &amp; Blotto, B. L. &amp; Ron, S. R. &amp; Baldo, D. &amp; Pereyra, M. O. &amp; Mayorga, A. M. &amp; Baeta, R. &amp; Ferreira, R. B. &amp; Barrio-Amoros, C. L. &amp; Borteiro, C. &amp; Brandao, R. A. &amp; Brasileiro, C. A. &amp; Donnelly, M. A. &amp; Dubeux, M. J. M. &amp; Kohler, J. &amp; Kolenc, F. &amp; Leite, F. S. F. &amp; Maciel, N. M. &amp; Nunes, I. &amp; Orrico, V. G. D. &amp; Peloso, P. L. V. &amp; Pezzuti, T. L. &amp; Reichle, S. &amp; Rojas-Runjaic, F. J. M. &amp; Silva, H. R. da &amp; Sturaro, M. J. &amp; Langone, J. A. &amp; Garcia, P. C. A. &amp; Rodrigues, M. T. &amp; Frost, D. R. &amp; Wheeler, W. C. &amp; Grant, T. &amp; Pombal, Jr., J. P. &amp; Haddad, C. F. B. &amp; Faivovich, J." box="[821,1120,1763,1789]" pageId="10" pageNumber="411" pagination="1 - 143" refId="ref9892" refString="Araujo-Vieira, K., Lourenco, A. C. C., Lacerda, J. V. A., Lyra, M. L., Blotto, B. L., Ron, S. R., Baldo, D., Pereyra, M. O., Suarez- Mayorga, A. M., Baeta, R., Ferreira, R. B., Barrio-Amoros, C. L., Borteiro, C., Brandao, R. A., Brasileiro, C. A., Donnelly, M. A., Dubeux, M. J. M., Kohler, J., Kolenc, F., Leite, F. S. F., Maciel, N. M., Nunes, I., Orrico, V. G. D., Peloso, P. L. V., Pezzuti, T. L., Reichle, S., Rojas-Runjaic, F. J. M., Silva, H. R. da, Sturaro, M. J., Langone, J. A., Garcia, P. C. A., Rodrigues, M. T., Frost, D. R., Wheeler, W. C., Grant, T., Pombal, Jr., J. P., Haddad, C. F. B. &amp; Faivovich, J. (2023) Treefrog diversity in the Neotropics: Phylogenetic relationships of Scinaxini (Anura: Hylidae: Hylinae). South American Journal of Herpetology, 27 (Special Issue), 1 - 143. https: // doi. org / 10.2994 / SAJH-D- 22 - 00038.1" type="journal article" year="2023">
Araujo-Vieira 
<emphasis id="B917EAACFFF5FFB1E88102CF3463F946" box="[982,1037,1763,1789]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
(2023)
</bibRefCitation>
, we restrict our comparisons to the species of the 
<taxonomicName id="4C634D3DFFF5FFB1EA2C032C324CF89A" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[379,546,1799,1825]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="cruentomma">
<emphasis id="B917EAACFFF5FFB1EA2C032C324CF89A" box="[379,546,1799,1825]" italics="true" pageId="10" pageNumber="411">S. cruentomma</emphasis>
</taxonomicName>
group. The smaller SVL (in mm) in males (20.2–22.9, n = 13) distinguishes the new species from 
<taxonomicName id="4C634D3DFFF5FFB1EA37030031D7F8FE" baseAuthorityName="Fouquette &amp; Pyburn" baseAuthorityYear="1972" box="[352,441,1835,1861]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="blairi">
<emphasis id="B917EAACFFF5FFB1EA37030031D7F8FE" box="[352,441,1835,1861]" italics="true" pageId="10" pageNumber="411">S. blairi</emphasis>
</taxonomicName>
(male SVL 27.8–30.1, n = 13; 
<bibRefCitation id="EFF24B4FFFF5FFB1E8460307345AF8FE" author="Fouquette, M. J. Jr. &amp; Pyburn, W. F." box="[785,1076,1835,1862]" pageId="10" pageNumber="411" pagination="176 - 181" refId="ref12027" refString="Fouquette, M. J. Jr. &amp; Pyburn, W. F. (1972) A new Colombian treefrog of the Hyla rubra complex. Herpetologica, 28, 176 - 181." type="journal article" year="1972">Fouquette &amp; Pyburn 1972</bibRefCitation>
) and 
<taxonomicName id="4C634D3DFFF5FFB1EF2603003573F8FE" baseAuthorityName="Pyburn" baseAuthorityYear="1993" box="[1137,1309,1835,1861]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="karenanneae">
<emphasis id="B917EAACFFF5FFB1EF2603003573F8FE" box="[1137,1309,1835,1861]" italics="true" pageId="10" pageNumber="411">S. karenanneae</emphasis>
</taxonomicName>
(male SVL 26.6–28.9, n = 2; 
<bibRefCitation id="EFF24B4FFFF5FFB1EA090364319CF8D2" author="Pyburn, W. F." box="[350,498,1871,1897]" pageId="10" pageNumber="411" pagination="46 - 50" refId="ref13892" refString="Pyburn, W. F. (1993). A new species of dimorphic tree frog, genus Hyla (Amphibia: Anura: Hylidae), from the Vaupes River of Colombia. Proceedings of the Biological Society of Washington, 106, 46 - 50." type="journal article" year="1993">Pyburn 1993</bibRefCitation>
).
</paragraph>
<paragraph id="8BDC36BEFFF5FFB1EB90035831AFF842" blockId="10.[151,1437,1763,2042]" pageId="10" pageNumber="411">
The weakly bilobate vocal sac, not reaching the pectoral region, distinguishes the new species from 
<taxonomicName id="4C634D3DFFF5FFB1EE17035835FBF836" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[1344,1429,1907,1933]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="altae">
<emphasis id="B917EAACFFF5FFB1EE17035835FBF836" box="[1344,1429,1907,1933]" italics="true" pageId="10" pageNumber="411">S. altae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF5FFB1EBC003BC313BF80A" baseAuthorityName="Rivero" baseAuthorityYear="1961" box="[151,341,1943,1969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="baumgardneri">
<emphasis id="B917EAACFFF5FFB1EBC003BC313BF80A" box="[151,341,1943,1969]" italics="true" pageId="10" pageNumber="411">S. baumgardneri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF5FFB1EA3103BC31ADF80A" baseAuthorityName="Fouquette &amp; Pyburn" baseAuthorityYear="1972" box="[358,451,1943,1969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="blairi">
<emphasis id="B917EAACFFF5FFB1EA3103BC31ADF80A" box="[358,451,1943,1969]" italics="true" pageId="10" pageNumber="411">S. blairi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF5FFB1EA8303BC3211F80A" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[468,639,1943,1969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="cruentomma">
<emphasis id="B917EAACFFF5FFB1EA8303BC3211F80A" box="[468,639,1943,1969]" italics="true" pageId="10" pageNumber="411">S. cruentomma</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF5FFB1E9C703BC336BF80A" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[656,773,1943,1969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="exiguus">
<emphasis id="B917EAACFFF5FFB1E9C703BC336BF80A" box="[656,773,1943,1969]" italics="true" pageId="10" pageNumber="411">S. exiguus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF5FFB1E84203BC33C9F80A" box="[789,935,1943,1969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="manriquei">
<emphasis id="B917EAACFFF5FFB1E84203BC33C9F80A" box="[789,935,1943,1969]" italics="true" pageId="10" pageNumber="411">S. manriquei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF5FFB1E8EF03BC345AF80A" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[952,1076,1943,1969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B917EAACFFF5FFB1E8EF03BC345AF80A" box="[952,1076,1943,1969]" italics="true" pageId="10" pageNumber="411">S. staufferi</emphasis>
</taxonomicName>
(single vocal sac, reaching the pectoral region; 
<bibRefCitation id="EFF24B4FFFF5FFB1EA07039031B0F86E" author="Rivero, J. A." box="[336,478,1979,2005]" pageId="10" pageNumber="411" pagination="1 - 207" refId="ref13987" refString="Rivero, J. A. (1961) Salientia of Venezuela. Bulletin of the Museum of Comparative Zoology, 126, 1 - 207." type="journal article" year="1961">Rivero 1961</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF5FFB1EABC039032F2F86E" author="Duellman, W. E." box="[491,668,1979,2005]" pageId="10" pageNumber="411" pagination="265 - 271" refId="ref11147" refString="Duellman, W. E. (1972) A new species of Hyla from Amazonian Ecuador. Copeia, 1972, 265 - 271. https: // doi. org / 10.2307 / 1442487" type="journal article" year="1972">Duellman 1972</bibRefCitation>
, 
<bibRefCitation id="EFF24B4FFFF5FFB1E9F00390328CF86E" author="Duellman, W. E." box="[679,738,1979,2005]" pageId="10" pageNumber="411" pagination="864 - 870" refId="ref11183" refString="Duellman, W. E. (1986) Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986, 864 - 870. https: // doi. org / 10.2307 / 1445281" type="journal article" year="1986">1986</bibRefCitation>
, 
<bibRefCitation id="EFF24B4FFFF5FFB1E9B903903347F86E" author="Duellman, W. E." box="[750,809,1979,2005]" pageId="10" pageNumber="411" refId="ref11225" refString="Duellman, W. E. (2001) Hylid Frogs of Middle America. 2 Vols. Revised Edition. Society for the Study of Amphibians and Reptiles, Ithaca, New York, xi + 753 pp., 324 figs., 72 pls." type="book" year="2001">2001</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF5FFB1E86203973433F86E" author="Fouquette, M. J. Jr. &amp; Pyburn, W. F." box="[821,1117,1979,2006]" pageId="10" pageNumber="411" pagination="176 - 181" refId="ref12027" refString="Fouquette, M. J. Jr. &amp; Pyburn, W. F. (1972) A new Colombian treefrog of the Hyla rubra complex. Herpetologica, 28, 176 - 181." type="journal article" year="1972">Fouquette &amp; Pyburn 1972</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF5FFB1EF3D039035F6F86E" author="Barrio-Amoros, C. L. &amp; Orellana, A. &amp; Chacon, A." box="[1130,1432,1979,2005]" pageId="10" pageNumber="411" pagination="105 - 112" refId="ref10317" refString="Barrio-Amoros, C. L., Orellana, A. &amp; Chacon, A. (2004) A new species of Scinax (Anura: Hylidae) from the Andes of Venezuela. Journal of Herpetology, 38, 105 - 112. https: // doi. org / 10.1670 / 82 - 01 N" type="journal article" year="2004">
Barrio-Amorós 
<emphasis id="B917EAACFFF5FFB1EE4B0397353BF86E" box="[1308,1365,1979,2005]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
2004
</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF5FFB1EBC003F431D8F842" author="Araujo-Vieira, K. &amp; Lourenco, A. C. C. &amp; Lacerda, J. V. A. &amp; Lyra, M. L. &amp; Blotto, B. L. &amp; Ron, S. R. &amp; Baldo, D. &amp; Pereyra, M. O. &amp; Mayorga, A. M. &amp; Baeta, R. &amp; Ferreira, R. B. &amp; Barrio-Amoros, C. L. &amp; Borteiro, C. &amp; Brandao, R. A. &amp; Brasileiro, C. A. &amp; Donnelly, M. A. &amp; Dubeux, M. J. M. &amp; Kohler, J. &amp; Kolenc, F. &amp; Leite, F. S. F. &amp; Maciel, N. M. &amp; Nunes, I. &amp; Orrico, V. G. D. &amp; Peloso, P. L. V. &amp; Pezzuti, T. L. &amp; Reichle, S. &amp; Rojas-Runjaic, F. J. M. &amp; Silva, H. R. da &amp; Sturaro, M. J. &amp; Langone, J. A. &amp; Garcia, P. C. A. &amp; Rodrigues, M. T. &amp; Frost, D. R. &amp; Wheeler, W. C. &amp; Grant, T. &amp; Pombal, Jr., J. P. &amp; Haddad, C. F. B. &amp; Faivovich, J." box="[151,438,2015,2042]" pageId="10" pageNumber="411" pagination="1 - 143" refId="ref9892" refString="Araujo-Vieira, K., Lourenco, A. C. C., Lacerda, J. V. A., Lyra, M. L., Blotto, B. L., Ron, S. R., Baldo, D., Pereyra, M. O., Suarez- Mayorga, A. M., Baeta, R., Ferreira, R. B., Barrio-Amoros, C. L., Borteiro, C., Brandao, R. A., Brasileiro, C. A., Donnelly, M. A., Dubeux, M. J. M., Kohler, J., Kolenc, F., Leite, F. S. F., Maciel, N. M., Nunes, I., Orrico, V. G. D., Peloso, P. L. V., Pezzuti, T. L., Reichle, S., Rojas-Runjaic, F. J. M., Silva, H. R. da, Sturaro, M. J., Langone, J. A., Garcia, P. C. A., Rodrigues, M. T., Frost, D. R., Wheeler, W. C., Grant, T., Pombal, Jr., J. P., Haddad, C. F. B. &amp; Faivovich, J. (2023) Treefrog diversity in the Neotropics: Phylogenetic relationships of Scinaxini (Anura: Hylidae: Hylinae). South American Journal of Herpetology, 27 (Special Issue), 1 - 143. https: // doi. org / 10.2994 / SAJH-D- 22 - 00038.1" type="journal article" year="2023">
Araujo-Vieira 
<emphasis id="B917EAACFFF5FFB1EA6D03CB3105F842" box="[314,363,2015,2041]" italics="true" pageId="10" pageNumber="411">et al</emphasis>
. 2023
</bibRefCitation>
).
</paragraph>
<paragraph id="8BDC36BEFFF4FFB0EB9004BC3418FE4E" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="412">
The dorsal color pattern of the body, uniformly light to dark brown with scattered darker blotches of different sizes, distinguishes 
<taxonomicName id="4C634D3DFFF4FFB0EA2A04903276FF6E" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[381,536,187,213]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF4FFB0EA2A04903276FF6E" box="[381,536,187,213]" italics="true" pageId="11" pageNumber="412">S. ushiniauae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C634D3DFFF4FFB0E93304903299FF6E" box="[612,759,187,213]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="albertinae">
<emphasis id="B917EAACFFF4FFB0E93304903299FF6E" box="[612,759,187,213]" italics="true" pageId="11" pageNumber="412">S. albertinae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E85E0490330FFF6E" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[777,865,187,213]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="altae">
<emphasis id="B917EAACFFF4FFB0E85E0490330FFF6E" box="[777,865,187,213]" italics="true" pageId="11" pageNumber="412">S. altae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E82404903387FF6F" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[883,1001,187,213]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="exiguus">
<emphasis id="B917EAACFFF4FFB0E82404903387FF6F" box="[883,1001,187,213]" italics="true" pageId="11" pageNumber="412">S. exiguus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E8AD04903419FF6E" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[1018,1143,187,213]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B917EAACFFF4FFB0E8AD04903419FF6E" box="[1018,1143,187,213]" italics="true" pageId="11" pageNumber="412">S. staufferi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C634D3DFFF4FFB0EFEA0490355AFF6E" baseAuthorityName="Pyburn &amp; Fouquette" baseAuthorityYear="1971" box="[1213,1332,187,213]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="wandae">
<emphasis id="B917EAACFFF4FFB0EFEA0490355AFF6E" box="[1213,1332,187,213]" italics="true" pageId="11" pageNumber="412">S. wandae</emphasis>
</taxonomicName>
(dorsum uniformly brown with longitudinal dark stripes; 
<bibRefCitation id="EFF24B4FFFF4FFB0E99104F43317FF42" author="Duellman, W. E." box="[710,889,223,249]" pageId="11" pageNumber="412" pagination="864 - 870" refId="ref11183" refString="Duellman, W. E. (1986) Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986, 864 - 870. https: // doi. org / 10.2307 / 1445281" type="journal article" year="1986">Duellman 1986</bibRefCitation>
, 
<bibRefCitation id="EFF24B4FFFF4FFB0E8D004F433ACFF42" author="Duellman, W. E." box="[903,962,223,249]" pageId="11" pageNumber="412" refId="ref11225" refString="Duellman, W. E. (2001) Hylid Frogs of Middle America. 2 Vols. Revised Edition. Society for the Study of Amphibians and Reptiles, Ithaca, New York, xi + 753 pp., 324 figs., 72 pls." type="book" year="2001">2001</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0E88704F4356EFF42" author="Pyburn, W. F. &amp; Fouquette, M. J. Jr." box="[976,1280,223,249]" pageId="11" pageNumber="412" pagination="97 - 101" refId="ref13941" refString="Pyburn, W. F. &amp; Fouquette, M. J. Jr. (1971) A new striped treefrog from central Colombia. Journal of Herpetology, 5, 97 - 101. https: // doi. org / 10.2307 / 1562731" type="journal article" year="1971">Pyburn &amp; Fouquette 1971</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EE5904F430BAFEA6" author="Ferrao, M. &amp; Moravec, J. &amp; Ferreira, A. S. &amp; Moraes, L. J. C. L. &amp; Hanken, J." pageId="11" pageNumber="412" pagination="1 - 36" refId="ref11907" refString="Ferrao, M., Moravec, J., Ferreira, A. S., Moraes, L. J. C. L. &amp; Hanken, J. (2022) A new snouted treefrog of the genus Scinax (Anura, Hylidae) from the white-sand forests of central Amazonia. Breviora, 573, 1 - 36. https: // doi. org / 10.3099 / 0006 - 9698 - 573.1.1" type="journal article" year="2022">
Ferrão 
<emphasis id="B917EAACFFF4FFB0EE3604F435F3FF42" box="[1377,1437,223,249]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2022
</bibRefCitation>
). The presence of a median horizontal red streak in the iris in living specimens distinguishes 
<taxonomicName id="4C634D3DFFF4FFB0EE54052835F2FEA6" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[1283,1436,259,285]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF4FFB0EE54052835F2FEA6" box="[1283,1436,259,285]" italics="true" pageId="11" pageNumber="412">S. ushiniauae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C634D3DFFF4FFB0EB80050C3106FEFA" box="[215,360,295,321]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="albertinae">
<emphasis id="B917EAACFFF4FFB0EB80050C3106FEFA" box="[215,360,295,321]" italics="true" pageId="11" pageNumber="412">S. albertinae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0EA2E050C3183FEFB" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[377,493,295,321]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="exiguus">
<emphasis id="B917EAACFFF4FFB0EA2E050C3183FEFB" box="[377,493,295,321]" italics="true" pageId="11" pageNumber="412">S. exiguus</emphasis>
</taxonomicName>
(median horizontal reddish brown streak in the iris; 
<bibRefCitation id="EFF24B4FFFF4FFB0EF18050C356FFEFA" author="Duellman, W. E." box="[1103,1281,295,321]" pageId="11" pageNumber="412" pagination="864 - 870" refId="ref11183" refString="Duellman, W. E. (1986) Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986, 864 - 870. https: // doi. org / 10.2307 / 1445281" type="journal article" year="1986">Duellman 1986</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EE58050C30BDFEDE" author="Ferrao, M. &amp; Moravec, J. &amp; Ferreira, A. S. &amp; Moraes, L. J. C. L. &amp; Hanken, J." pageId="11" pageNumber="412" pagination="1 - 36" refId="ref11907" refString="Ferrao, M., Moravec, J., Ferreira, A. S., Moraes, L. J. C. L. &amp; Hanken, J. (2022) A new snouted treefrog of the genus Scinax (Anura, Hylidae) from the white-sand forests of central Amazonia. Breviora, 573, 1 - 36. https: // doi. org / 10.3099 / 0006 - 9698 - 573.1.1" type="journal article" year="2022">
Ferrão 
<emphasis id="B917EAACFFF4FFB0EE35050C35F3FEFA" box="[1378,1437,295,321]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2022
</bibRefCitation>
), 
<taxonomicName id="4C634D3DFFF4FFB0EBBD0560312CFEDE" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[234,322,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="altae">
<emphasis id="B917EAACFFF4FFB0EBBD0560312CFEDE" box="[234,322,331,357]" italics="true" pageId="11" pageNumber="412">S. altae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0EA030560327DFEDE" baseAuthorityName="Rivero" baseAuthorityYear="1961" box="[340,531,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="baumgardneri">
<emphasis id="B917EAACFFF4FFB0EA030560327DFEDE" box="[340,531,331,357]" italics="true" pageId="11" pageNumber="412">S. baumgardneri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E972056032EDFEDE" baseAuthorityName="Fouquette &amp; Pyburn" baseAuthorityYear="1972" box="[549,643,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="blairi">
<emphasis id="B917EAACFFF4FFB0E972056032EDFEDE" box="[549,643,331,357]" italics="true" pageId="11" pageNumber="412">S. blairi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E9C205603346FEDF" box="[661,808,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="manriquei">
<emphasis id="B917EAACFFF4FFB0E9C205603346FEDF" box="[661,808,331,357]" italics="true" pageId="11" pageNumber="412">S. manriquei</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E86D05603387FEDE" baseAuthorityName="Pyburn" baseAuthorityYear="1993" box="[826,1001,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="karenanneae">
<emphasis id="B917EAACFFF4FFB0E86D05603387FEDE" box="[826,1001,331,357]" italics="true" pageId="11" pageNumber="412">S. karenanneae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E8AC0560341AFEDE" authorityName="Pyburn" authorityYear="1992" box="[1019,1140,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="lindsayi">
<emphasis id="B917EAACFFF4FFB0E8AC0560341AFEDE" box="[1019,1140,331,357]" italics="true" pageId="11" pageNumber="412">S. lindsayi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C634D3DFFF4FFB0EFEF0560355BFEDE" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[1208,1333,331,357]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B917EAACFFF4FFB0EFEF0560355BFEDE" box="[1208,1333,331,357]" italics="true" pageId="11" pageNumber="412">S. staufferi</emphasis>
</taxonomicName>
(median horizontal red streak in the iris absent; 
<bibRefCitation id="EFF24B4FFFF4FFB0E914054432BEFE32" author="Rivero, J. A." box="[579,720,367,393]" pageId="11" pageNumber="412" pagination="1 - 207" refId="ref13987" refString="Rivero, J. A. (1961) Salientia of Venezuela. Bulletin of the Museum of Comparative Zoology, 126, 1 - 207." type="journal article" year="1961">Rivero 1961</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0E98D054433E7FE32" author="Duellman, W. E." box="[730,905,367,393]" pageId="11" pageNumber="412" refId="ref11225" refString="Duellman, W. E. (2001) Hylid Frogs of Middle America. 2 Vols. Revised Edition. Society for the Study of Amphibians and Reptiles, Ithaca, New York, xi + 753 pp., 324 figs., 72 pls." type="book" year="2001">Duellman 2001</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0E8C3054434D9FE32" author="Fouquette, M. J. Jr. &amp; Pyburn, W. F." box="[916,1207,367,393]" pageId="11" pageNumber="412" pagination="176 - 181" refId="ref12027" refString="Fouquette, M. J. Jr. &amp; Pyburn, W. F. (1972) A new Colombian treefrog of the Hyla rubra complex. Herpetologica, 28, 176 - 181." type="journal article" year="1972">Fouquette &amp; Pyburn 1972</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EF960544353DFE32" author="Pyburn, W. F." box="[1217,1363,367,393]" pageId="11" pageNumber="412" pagination="405 - 411" refId="ref13855" refString="Pyburn, W. F. (1992) A new tree frog of the genus Scinax from the Vaupes River of northwestern Brazil. The Texas Journal of Science, 44, 405 - 411." type="journal article" year="1992">Pyburn 1992</bibRefCitation>
, 
<bibRefCitation id="EFF24B4FFFF4FFB0EE0B054435F9FE32" author="Pyburn, W. F." box="[1372,1431,367,393]" pageId="11" pageNumber="412" pagination="46 - 50" refId="ref13892" refString="Pyburn, W. F. (1993). A new species of dimorphic tree frog, genus Hyla (Amphibia: Anura: Hylidae), from the Vaupes River of Colombia. Proceedings of the Biological Society of Washington, 106, 46 - 50." type="journal article" year="1993">1993</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EBC005B831A4FE16" author="Barrio-Amoros, C. L. &amp; Orellana, A. &amp; Chacon, A." box="[151,458,403,429]" pageId="11" pageNumber="412" pagination="105 - 112" refId="ref10317" refString="Barrio-Amoros, C. L., Orellana, A. &amp; Chacon, A. (2004) A new species of Scinax (Anura: Hylidae) from the Andes of Venezuela. Journal of Herpetology, 38, 105 - 112. https: // doi. org / 10.1670 / 82 - 01 N" type="journal article" year="2004">
Barrio-Amorós 
<emphasis id="B917EAACFFF4FFB0EA1C05B831EBFE16" box="[331,389,403,429]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2004
</bibRefCitation>
). Furthermore, physiological chlorosis (impregnation of lymph, bones and some other tissues with biliverdin associated with its carrying serpin; 
<bibRefCitation id="EFF24B4FFFF4FFB0E872059C33C1FE6A" author="Barrio, A." box="[805,943,438,465]" pageId="11" pageNumber="412" refId="ref10252" refString="Barrio, A. (1965) Cloricia fisiologica en batracios anuros. Physis, 25,137 - 142." type="book" year="1965">Barrio 1965</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0E8EC059C34F5FE6A" author="Taboada, C. &amp; Brunetti, A. E. &amp; Lyra, M. L. &amp; Fitak, R. &amp; Faigon, A. &amp; Ron, S. &amp; Lagorio, M. G. &amp; Haddad, C. F. B. &amp; Lopes, N. P. &amp; Johnsen, S. &amp; Faivovich, J. &amp; Chemes, L. B. &amp; Bari, S." box="[955,1179,439,465]" pageId="11" pageNumber="412" pagination="18574 - 18581" refId="ref14353" refString="Taboada, C., Brunetti, A. E., Lyra, M. L., Fitak, R., Faigon, A., Ron, S., Lagorio, M. G., Haddad, C. F. B., Lopes, N. P., Johnsen, S., Faivovich, J., Chemes, L. B. &amp; Bari, S. (2020) Multiple origins of green coloration in frogs mediated by a novel biliverdinbinding serpin. Proceedings of the National Academy of Sciences, United States of America, 117, 18574 - 18581. https: // doi. org / 10.1073 / pnas. 2006771117" type="journal article" year="2020">
Taboada 
<emphasis id="B917EAACFFF4FFB0EF76059C343DFE6A" box="[1057,1107,439,465]" italics="true" pageId="11" pageNumber="412">et al</emphasis>
. 2020
</bibRefCitation>
) distinguishes the new species from 
<taxonomicName id="4C634D3DFFF4FFB0EA7B05F031EEFE4E" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[300,384,475,501]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="altae">
<emphasis id="B917EAACFFF4FFB0EA7B05F031EEFE4E" box="[300,384,475,501]" italics="true" pageId="11" pageNumber="412">S. altae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C634D3DFFF4FFB0EAE005F0325EFE4E" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[439,560,475,501]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B917EAACFFF4FFB0EAE005F0325EFE4E" box="[439,560,475,501]" italics="true" pageId="11" pageNumber="412">S. staufferi</emphasis>
</taxonomicName>
(chlorosis absent in these species; 
<bibRefCitation id="EFF24B4FFFF4FFB0E8EF05F03404FE4E" author="Duellman, W. E." box="[952,1130,475,501]" pageId="11" pageNumber="412" pagination="1 - 753" refId="ref11093" refString="Duellman, W. E. (1970) The Hylid Frogs of Middle America. Monograph of the Museum of Natural History, The University of Kansas, 1, 1 - 753. [2 Vols.] https: // doi. org / 10.5962 / bhl. title. 2835" type="journal article" year="1970">Duellman 1970</bibRefCitation>
).
</paragraph>
<paragraph id="8BDC36BEFFF4FFB0EB9005D434EAFD76" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="412">
The extension of pre- and postaxial webbings of Toe IV reaching the proximal half of the penultimate phalanx distinguishes the new species from 
<taxonomicName id="4C634D3DFFF4FFB0E96006083291FD86" box="[567,767,547,573]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="albertinae">
<emphasis id="B917EAACFFF4FFB0E96006083291FD86" box="[567,767,547,573]" italics="true" pageId="11" pageNumber="412">Scinax albertinae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C634D3DFFF4FFB0E846060833E9FD87" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[785,903,547,573]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="exiguus">
<emphasis id="B917EAACFFF4FFB0E846060833E9FD87" box="[785,903,547,573]" italics="true" pageId="11" pageNumber="412">S. exiguus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C634D3DFFF4FFB0E89C0608342CFD86" baseAuthorityName="Pyburn &amp; Fouquette" baseAuthorityYear="1971" box="[971,1090,547,573]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="wandae">
<emphasis id="B917EAACFFF4FFB0E89C0608342CFD86" box="[971,1090,547,573]" italics="true" pageId="11" pageNumber="412">S. wandae</emphasis>
</taxonomicName>
(pre- and postaxial webbings of Toe IV not surpassing the proximal half of antepenultimate phalanx anteriorly; 
<bibRefCitation id="EFF24B4FFFF4FFB0EF34066C35F9FDDA" author="Pyburn, W. F. &amp; Fouquette, M. J. Jr." box="[1123,1431,583,609]" pageId="11" pageNumber="412" pagination="97 - 101" refId="ref13941" refString="Pyburn, W. F. &amp; Fouquette, M. J. Jr. (1971) A new striped treefrog from central Colombia. Journal of Herpetology, 5, 97 - 101. https: // doi. org / 10.2307 / 1562731" type="journal article" year="1971">Pyburn &amp; Fouquette 1971</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EBC006403126FD3E" author="Duellman, W. E." box="[151,328,619,645]" pageId="11" pageNumber="412" pagination="864 - 870" refId="ref11183" refString="Duellman, W. E. (1986) Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986, 864 - 870. https: // doi. org / 10.2307 / 1445281" type="journal article" year="1986">Duellman 1986</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EA0406403271FD3E" author="Ferrao, M. &amp; Moravec, J. &amp; Ferreira, A. S. &amp; Moraes, L. J. C. L. &amp; Hanken, J." box="[339,543,619,645]" pageId="11" pageNumber="412" pagination="1 - 36" refId="ref11907" refString="Ferrao, M., Moravec, J., Ferreira, A. S., Moraes, L. J. C. L. &amp; Hanken, J. (2022) A new snouted treefrog of the genus Scinax (Anura, Hylidae) from the white-sand forests of central Amazonia. Breviora, 573, 1 - 36. https: // doi. org / 10.3099 / 0006 - 9698 - 573.1.1" type="journal article" year="2022">
Ferrão 
<emphasis id="B917EAACFFF4FFB0EAF4064031B2FD3E" box="[419,476,619,645]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2022
</bibRefCitation>
). The m. depressor mandibulae with an origin from the dorsal fascia that covers the suprascapula differentiates 
<taxonomicName id="4C634D3DFFF4FFB0EAB806A432D5FD12" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[495,699,655,681]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF4FFB0EAB806A432D5FD12" box="[495,699,655,681]" italics="true" pageId="11" pageNumber="412">Scinax ushiniauae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C634D3DFFF4FFB0E9AA06A4331BFD12" baseAuthorityName="Cope" baseAuthorityYear="1865" box="[765,885,655,681]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B917EAACFFF4FFB0E9AA06A4331BFD12" box="[765,885,655,681]" italics="true" pageId="11" pageNumber="412">S. staufferi</emphasis>
</taxonomicName>
(m. depressor mandibulae without an origin from the dorsal fascia that covers the suprascapula; 
<bibRefCitation id="EFF24B4FFFF4FFB0E9C90698333FFD76" author="Faivovich, J." box="[670,849,691,717]" pageId="11" pageNumber="412" pagination="367 - 393" refId="ref11508" refString="Faivovich, J. (2002) A cladistic analysis of Scinax (Anura: Hylidae). Cladistics, 18, 367 - 393. https: // doi. org / 10.1111 / j. 1096 - 0031.2002. tb 00157. x" type="journal article" year="2002">Faivovich 2002</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0E80D06983416FD76" author="Araujo-Vieira, K. &amp; Lourenco, A. C. C. &amp; Lacerda, J. V. A. &amp; Lyra, M. L. &amp; Blotto, B. L. &amp; Ron, S. R. &amp; Baldo, D. &amp; Pereyra, M. O. &amp; Mayorga, A. M. &amp; Baeta, R. &amp; Ferreira, R. B. &amp; Barrio-Amoros, C. L. &amp; Borteiro, C. &amp; Brandao, R. A. &amp; Brasileiro, C. A. &amp; Donnelly, M. A. &amp; Dubeux, M. J. M. &amp; Kohler, J. &amp; Kolenc, F. &amp; Leite, F. S. F. &amp; Maciel, N. M. &amp; Nunes, I. &amp; Orrico, V. G. D. &amp; Peloso, P. L. V. &amp; Pezzuti, T. L. &amp; Reichle, S. &amp; Rojas-Runjaic, F. J. M. &amp; Silva, H. R. da &amp; Sturaro, M. J. &amp; Langone, J. A. &amp; Garcia, P. C. A. &amp; Rodrigues, M. T. &amp; Frost, D. R. &amp; Wheeler, W. C. &amp; Grant, T. &amp; Pombal, Jr., J. P. &amp; Haddad, C. F. B. &amp; Faivovich, J." box="[858,1144,691,717]" pageId="11" pageNumber="412" pagination="1 - 143" refId="ref9892" refString="Araujo-Vieira, K., Lourenco, A. C. C., Lacerda, J. V. A., Lyra, M. L., Blotto, B. L., Ron, S. R., Baldo, D., Pereyra, M. O., Suarez- Mayorga, A. M., Baeta, R., Ferreira, R. B., Barrio-Amoros, C. L., Borteiro, C., Brandao, R. A., Brasileiro, C. A., Donnelly, M. A., Dubeux, M. J. M., Kohler, J., Kolenc, F., Leite, F. S. F., Maciel, N. M., Nunes, I., Orrico, V. G. D., Peloso, P. L. V., Pezzuti, T. L., Reichle, S., Rojas-Runjaic, F. J. M., Silva, H. R. da, Sturaro, M. J., Langone, J. A., Garcia, P. C. A., Rodrigues, M. T., Frost, D. R., Wheeler, W. C., Grant, T., Pombal, Jr., J. P., Haddad, C. F. B. &amp; Faivovich, J. (2023) Treefrog diversity in the Neotropics: Phylogenetic relationships of Scinaxini (Anura: Hylidae: Hylinae). South American Journal of Herpetology, 27 (Special Issue), 1 - 143. https: // doi. org / 10.2994 / SAJH-D- 22 - 00038.1" type="journal article" year="2023">
Araujo-Vieira 
<emphasis id="B917EAACFFF4FFB0E8AB0698345BFD76" box="[1020,1077,691,717]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
<paragraph id="8BDC36BEFFF4FFB0EB9006FC33CEFB52" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="412">
The advertisement call consisting of a single, very short pulsed note, characterized by the combination of note duration 4 to 62 ms, 1–3 pulses/note, and pulse rate of 45.3–69.0 pulses/s differentiates 
<taxonomicName id="4C634D3DFFF4FFB0EFD906D03533FCAE" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[1166,1373,763,789]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF4FFB0EFD906D03533FCAE" box="[1166,1373,763,789]" italics="true" pageId="11" pageNumber="412">Scinax ushiniauae</emphasis>
</taxonomicName>
from all species of the 
<taxonomicName id="4C634D3DFFF4FFB0EA0B0734326BFC83" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[348,517,799,825]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="cruentomma">
<emphasis id="B917EAACFFF4FFB0EA0B0734326BFC83" box="[348,517,799,825]" italics="true" pageId="11" pageNumber="412">S. cruentomma</emphasis>
</taxonomicName>
group whose calls have been described (see the Discussion section about the call of 
<taxonomicName id="4C634D3DFFF4FFB0EBEF07683124FCE7" box="[184,330,835,861]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="manriquei">
<emphasis id="B917EAACFFF4FFB0EBEF07683124FCE7" box="[184,330,835,861]" italics="true" pageId="11" pageNumber="412">S. manriquei</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EA0C076832FCFCE6" author="Barrio-Amoros, C. L. &amp; Orellana, A. &amp; Chacon, A." box="[347,658,835,861]" pageId="11" pageNumber="412" pagination="105 - 112" refId="ref10317" refString="Barrio-Amoros, C. L., Orellana, A. &amp; Chacon, A. (2004) A new species of Scinax (Anura: Hylidae) from the Andes of Venezuela. Journal of Herpetology, 38, 105 - 112. https: // doi. org / 10.1670 / 82 - 01 N" type="journal article" year="2004">
Barrio-Amorós 
<emphasis id="B917EAACFFF4FFB0E94707683224FCE6" box="[528,586,835,861]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2004
</bibRefCitation>
): 
<taxonomicName id="4C634D3DFFF4FFB0E9F107683356FCE6" box="[678,824,835,861]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="albertinae">
<emphasis id="B917EAACFFF4FFB0E9F107683356FCE6" box="[678,824,835,861]" italics="true" pageId="11" pageNumber="412">S. albertinae</emphasis>
</taxonomicName>
(note duration 500–650 ms, 79–105 pulses/note, and pulse rate of 111–200 pulses/s, n = 3; 
<bibRefCitation id="EFF24B4FFFF4FFB0E969074C3367FC3A" author="Ferrao, M. &amp; Moravec, J. &amp; Ferreira, A. S. &amp; Moraes, L. J. C. L. &amp; Hanken, J." box="[574,777,871,897]" pageId="11" pageNumber="412" pagination="1 - 36" refId="ref11907" refString="Ferrao, M., Moravec, J., Ferreira, A. S., Moraes, L. J. C. L. &amp; Hanken, J. (2022) A new snouted treefrog of the genus Scinax (Anura, Hylidae) from the white-sand forests of central Amazonia. Breviora, 573, 1 - 36. https: // doi. org / 10.3099 / 0006 - 9698 - 573.1.1" type="journal article" year="2022">
Ferrão 
<emphasis id="B917EAACFFF4FFB0E9D9074C32A8FC3A" box="[654,710,871,897]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2022
</bibRefCitation>
: figs. 6A, B, 13A), 
<taxonomicName id="4C634D3DFFF4FFB0E8B3074C3456FC3A" baseAuthorityName="Dunn" baseAuthorityYear="1933" box="[996,1080,871,897]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="altae">
<emphasis id="B917EAACFFF4FFB0E8B3074C3456FC3A" box="[996,1080,871,897]" italics="true" pageId="11" pageNumber="412">S. altae</emphasis>
</taxonomicName>
(note duration 140–180 ms and 110–130 pulses/s, n = 7; 
<bibRefCitation id="EFF24B4FFFF4FFB0EAE507A03241FC1E" author="Leon, J. R." box="[434,559,907,933]" pageId="11" pageNumber="412" pagination="505 - 545" refId="ref12566" refString="Leon, J. R. (1969) The systematics of the frogs of the Hyla rubra group in Middle America. University of Kansas Publications, Museum of Natural History, 18, 505 - 545. https: // doi. org / 10.5962 / bhl. part. 19991" type="journal article" year="1969">León 1969</bibRefCitation>
: pl. 4C; 
<bibRefCitation id="EFF24B4FFFF4FFB0E9D807A0332FFC1E" author="Duellman, W. E." box="[655,833,907,933]" pageId="11" pageNumber="412" pagination="1 - 753" refId="ref11093" refString="Duellman, W. E. (1970) The Hylid Frogs of Middle America. Monograph of the Museum of Natural History, The University of Kansas, 1, 1 - 753. [2 Vols.] https: // doi. org / 10.5962 / bhl. title. 2835" type="journal article" year="1970">Duellman 1970</bibRefCitation>
: table 17), 
<taxonomicName id="4C634D3DFFF4FFB0E8E907A03414FC1E" baseAuthorityName="Rivero" baseAuthorityYear="1961" box="[958,1146,907,933]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="baumgardneri">
<emphasis id="B917EAACFFF4FFB0E8E907A03414FC1E" box="[958,1146,907,933]" italics="true" pageId="11" pageNumber="412">S. baumgardneri</emphasis>
</taxonomicName>
(note duration 530–700 s and 170–180 pulses/s, n = 5; 
<bibRefCitation id="EFF24B4FFFF4FFB0EA8E078432E7FC72" author="Duellman, W. E." box="[473,649,943,969]" pageId="11" pageNumber="412" pagination="265 - 271" refId="ref11147" refString="Duellman, W. E. (1972) A new species of Hyla from Amazonian Ecuador. Copeia, 1972, 265 - 271. https: // doi. org / 10.2307 / 1442487" type="journal article" year="1972">Duellman 1972</bibRefCitation>
: fig. 4A), 
<taxonomicName id="4C634D3DFFF4FFB0E9AF0784333FFC72" baseAuthorityName="Fouquette &amp; Pyburn" baseAuthorityYear="1972" box="[760,849,943,969]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="blairi">
<emphasis id="B917EAACFFF4FFB0E9AF0784333FFC72" box="[760,849,943,969]" italics="true" pageId="11" pageNumber="412">S. blairi</emphasis>
</taxonomicName>
(note duration 140–160 ms and 18–22 pulses/note, n = 9; 
<bibRefCitation id="EFF24B4FFFF4FFB0EB9E07F83183FC56" author="Fouquette, M. J. Jr. &amp; Pyburn, W. F." box="[201,493,979,1005]" pageId="11" pageNumber="412" pagination="176 - 181" refId="ref12027" refString="Fouquette, M. J. Jr. &amp; Pyburn, W. F. (1972) A new Colombian treefrog of the Hyla rubra complex. Herpetologica, 28, 176 - 181." type="journal article" year="1972">Fouquette &amp; Pyburn 1972</bibRefCitation>
: fig. 3), 
<taxonomicName id="4C634D3DFFF4FFB0E91007F83280FC57" baseAuthorityName="Duellman" baseAuthorityYear="1972" box="[583,750,979,1005]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="cruentomma">
<emphasis id="B917EAACFFF4FFB0E91007F83280FC57" box="[583,750,979,1005]" italics="true" pageId="11" pageNumber="412">S. cruentomma</emphasis>
</taxonomicName>
(note duration 220–370 ms, 39–54 pulses/note, and pulse rate of 153.8–192.8 pulses/s, n = 6; 
<bibRefCitation id="EFF24B4FFFF4FFB0EAA107DC32C6FBAA" author="Duellman, W. E." box="[502,680,1015,1041]" pageId="11" pageNumber="412" pagination="265 - 271" refId="ref11147" refString="Duellman, W. E. (1972) A new species of Hyla from Amazonian Ecuador. Copeia, 1972, 265 - 271. https: // doi. org / 10.2307 / 1442487" type="journal article" year="1972">Duellman 1972</bibRefCitation>
: fig. 4B; 
<bibRefCitation id="EFF24B4FFFF4FFB0E85907DC3471FBAA" author="Duellman, W. E. &amp; Pyles, R. E." box="[782,1055,1015,1041]" pageId="11" pageNumber="412" pagination="639 - 649" refId="ref11273" refString="Duellman, W. E. &amp; Pyles, R. E. (1983) Acoustic resource partitioning in anuran communities. Copeia, 1983, 639 - 649. https: // doi. org / 10.2307 / 1444328" type="journal article" year="1983">Duellman &amp; Pyles 1983</bibRefCitation>
: table 2; 
<bibRefCitation id="EFF24B4FFFF4FFB0EFD307DC3502FBAA" author="Carvalho, T. R. &amp; Teixeira, B. F. V. &amp; Duellman, W. E. &amp; Giaretta, A. A." box="[1156,1388,1014,1041]" pageId="11" pageNumber="412" pagination="139 - 146" refId="ref10552" refString="Carvalho, T. R., Teixeira, B. F. V., Duellman, W. E. &amp; Giaretta, A. A. (2015) Scinax cruentommus (Anura: Hylidae) in the upper Rio Negro drainage, Amazonas state, Brazil, with the redescription of its advertisement call. Phyllomedusa, 14, 139 - 146. https: // doi. org / 10.11606 / issn. 2316 - 9079. v 14 i 2 p 139 - 146" type="journal article" year="2015">
Carvalho 
<emphasis id="B917EAACFFF4FFB0EFA607DC3547FBAA" box="[1265,1321,1015,1041]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2015
</bibRefCitation>
: fig. 2), 
<taxonomicName id="4C634D3DFFF4FFB0EBEA00303141FB8F" baseAuthorityName="Duellman" baseAuthorityYear="1986" box="[189,303,1051,1077]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="exiguus">
<emphasis id="B917EAACFFF4FFB0EBEA00303141FB8F" box="[189,303,1051,1077]" italics="true" pageId="11" pageNumber="412">S. exiguus</emphasis>
</taxonomicName>
(note duration 200–1,700 ms, 23–90 pulses/note, and pulse rate of 64–125 pulses/s, n = 9; 
<bibRefCitation id="EFF24B4FFFF4FFB0EE79003030BDFBE2" author="Duellman, W. E." pageId="11" pageNumber="412" pagination="864 - 870" refId="ref11183" refString="Duellman, W. E. (1986) Two new species of Ololygon (Anura: Hylidae) from the Venezuelan Guyana. Copeia, 1986, 864 - 870. https: // doi. org / 10.2307 / 1445281" type="journal article" year="1986">Duellman 1986</bibRefCitation>
: fig. 2C; 
<bibRefCitation id="EFF24B4FFFF4FFB0EA6800143242FBE2" author="Carvalho, T. R. &amp; Azarak P. &amp; Bang, D. L. &amp; Duellman, W. E. &amp; Giaretta, A. A." box="[319,556,1087,1113]" pageId="11" pageNumber="412" pagination="196 - 202" refId="ref10642" refString="Carvalho, T. R., Azarak P., Bang, D. L., Duellman, W. E. &amp; Giaretta, A. A. (2017) A reassessment of the vocalization and distribution of Scinax exiguus (Duellman, 1986) (Anura: Hylidae) in the Amazonian savanna of Roraima, northern Brazil, with description of its aggressive call. Neotropical Biodiversity, 3, 196 - 202. https: // doi. org / 10.1080 / 23766808.2017.1375813" type="journal article" year="2017">
Carvalho 
<emphasis id="B917EAACFFF4FFB0EAF900143186FBE2" box="[430,488,1087,1113]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2017
</bibRefCitation>
: fig. 2), 
<taxonomicName id="4C634D3DFFF4FFB0E9D90014336BFBE2" authorityName="Pyburn" authorityYear="1992" box="[654,773,1087,1113]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="lindsayi">
<emphasis id="B917EAACFFF4FFB0E9D90014336BFBE2" box="[654,773,1087,1113]" italics="true" pageId="11" pageNumber="412">S. lindsayi</emphasis>
</taxonomicName>
(note duration 80–100 ms, n = 3; 
<bibRefCitation id="EFF24B4FFFF4FFB0EFD80014354BFBE2" author="Pyburn, W. F." box="[1167,1317,1087,1113]" pageId="11" pageNumber="412" pagination="405 - 411" refId="ref13855" refString="Pyburn, W. F. (1992) A new tree frog of the genus Scinax from the Vaupes River of northwestern Brazil. The Texas Journal of Science, 44, 405 - 411." type="journal article" year="1992">Pyburn 1992</bibRefCitation>
: fig. 2); 
<taxonomicName id="4C634D3DFFF4FFB0EED00014309AFBC6" baseAuthorityName="Cope" baseAuthorityYear="1865" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="staufferi">
<emphasis id="B917EAACFFF4FFB0EED00014309AFBC6" italics="true" pageId="11" pageNumber="412">S. staufferi</emphasis>
</taxonomicName>
(note duration 130–230 ms and 100–130 pulses/s, n = 18; 
<bibRefCitation id="EFF24B4FFFF4FFB0E82F0048339DFBC6" author="Leon, J. R." box="[888,1011,1123,1149]" pageId="11" pageNumber="412" pagination="505 - 545" refId="ref12566" refString="Leon, J. R. (1969) The systematics of the frogs of the Hyla rubra group in Middle America. University of Kansas Publications, Museum of Natural History, 18, 505 - 545. https: // doi. org / 10.5962 / bhl. part. 19991" type="journal article" year="1969">León 1969</bibRefCitation>
: pl. 4B; 
<bibRefCitation id="EFF24B4FFFF4FFB0EF1A00483493FBC6" author="Duellman, W. E." box="[1101,1277,1123,1149]" pageId="11" pageNumber="412" pagination="1 - 753" refId="ref11093" refString="Duellman, W. E. (1970) The Hylid Frogs of Middle America. Monograph of the Museum of Natural History, The University of Kansas, 1, 1 - 753. [2 Vols.] https: // doi. org / 10.5962 / bhl. title. 2835" type="journal article" year="1970">Duellman 1970</bibRefCitation>
; pl. 26, fig. 2), 
<taxonomicName id="4C634D3DFFF4FFB0EBC000AC3123FB1B" box="[151,333,1159,1185]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="strussmannae">
<emphasis id="B917EAACFFF4FFB0EBC000AC3123FB1B" box="[151,333,1159,1185]" italics="true" pageId="11" pageNumber="412">S. strussmannae</emphasis>
</taxonomicName>
(note duration 100–110 ms, 23–27 pulses/note, and pulse rate of 200–250 pulses/s, n = 1; 
<bibRefCitation id="EFF24B4FFFF4FFB0EE6F00AC316DFB7E" author="Ferrao, M. &amp; Moravec, J. &amp; Kaefer, I. L. &amp; Fraga, R. &amp; Lima, A. P." pageId="11" pageNumber="412" pagination="473 - 486" refId="ref11761" refString="Ferrao, M., Moravec, J., Kaefer, I. L., Fraga, R. &amp; Lima, A. P. (2018 a) New species of Scinax (Anura: Hylidae) with red-striped eyes from Brazilian Amazonia. Journal of Herpetology, 52, 473 - 486. https: // doi. org / 10.1670 / 17 - 165" type="journal article" year="2018">
Ferrão 
<emphasis id="B917EAACFFF4FFB0EEDF00AC30DAFB7E" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2018a
</bibRefCitation>
: fig. 5), and 
<taxonomicName id="4C634D3DFFF4FFB0EADA0080326EFB7E" baseAuthorityName="Pyburn &amp; Fouquette" baseAuthorityYear="1971" box="[397,512,1195,1221]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="wandae">
<emphasis id="B917EAACFFF4FFB0EADA0080326EFB7E" box="[397,512,1195,1221]" italics="true" pageId="11" pageNumber="412">S. wandae</emphasis>
</taxonomicName>
(note duration 600–710 ms, 70–108 pulses/note, and pulse rate about 130 pulses/s, n = 5; 
<bibRefCitation id="EFF24B4FFFF4FFB0EBB700E43260FB52" author="Pyburn, W. F. &amp; Fouquette, M. J. Jr." box="[224,526,1231,1257]" pageId="11" pageNumber="412" pagination="97 - 101" refId="ref13941" refString="Pyburn, W. F. &amp; Fouquette, M. J. Jr. (1971) A new striped treefrog from central Colombia. Journal of Herpetology, 5, 97 - 101. https: // doi. org / 10.2307 / 1562731" type="journal article" year="1971">Pyburn &amp; Fouquette, 1971</bibRefCitation>
: fig. 2; 
<bibRefCitation id="EFF24B4FFFF4FFB0E93400E43354FB52" author="Pombal, J. P. Jr. &amp; Bilate, M. &amp; Gambale, P. G. &amp; Signorelli, L. &amp; Bastos, R. P." box="[611,826,1231,1257]" pageId="11" pageNumber="412" pagination="288 - 299" refId="ref13713" refString="Pombal, J. P. Jr., Bilate, M., Gambale, P. G., Signorelli, L. &amp; Bastos, R. P. (2011) A new miniature treefrog of the Scinax ruber clade from the Cerrado of central Brazil (Anura: Hylidae). Herpetologica, 67, 288 - 299. https: // doi. org / 10.1655 / HERPETOLOGICA-D- 10 - 00067.1" type="journal article" year="2011">
Pombal 
<emphasis id="B917EAACFFF4FFB0E99700E43296FB52" box="[704,760,1231,1257]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2011
</bibRefCitation>
: fig. 7F).
</paragraph>
<paragraph id="8BDC36BEFFF4FFB0EB9000D8339BFA26" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="412">
<taxonomicName id="4C634D3DFFF4FFB0EB9000D831FDFAB6" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[199,403,1267,1293]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF4FFB0EB9000D831FDFAB6" box="[199,403,1267,1293]" italics="true" pageId="11" pageNumber="412">Scinax ushiniauae</emphasis>
</taxonomicName>
morphologically resembles 
<taxonomicName id="4C634D3DFFF4FFB0E99900D833ECFAB7" box="[718,898,1267,1293]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="strussmannae">
<emphasis id="B917EAACFFF4FFB0E99900D833ECFAB7" box="[718,898,1267,1293]" italics="true" pageId="11" pageNumber="412">S. strussmannae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF24B4FFFF4FFB0E8C700D83409FAB6" author="Ferrao, M. &amp; Moravec, J. &amp; Kaefer, I. L. &amp; Fraga, R. &amp; Lima, A. P." box="[912,1127,1267,1293]" pageId="11" pageNumber="412" pagination="473 - 486" refId="ref11761" refString="Ferrao, M., Moravec, J., Kaefer, I. L., Fraga, R. &amp; Lima, A. P. (2018 a) New species of Scinax (Anura: Hylidae) with red-striped eyes from Brazilian Amazonia. Journal of Herpetology, 52, 473 - 486. https: // doi. org / 10.1670 / 17 - 165" type="journal article" year="2018">
Ferrão 
<emphasis id="B917EAACFFF4FFB0E8B600D83476FAB6" box="[993,1048,1267,1293]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2018a
</bibRefCitation>
: figs. 1–4), differing mainly in traits of their advertisement calls. However, the new species can be further differentiated from 
<taxonomicName id="4C634D3DFFF4FFB0EFB2013C35F2FA8B" box="[1253,1436,1303,1329]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="strussmannae">
<emphasis id="B917EAACFFF4FFB0EFB2013C35F2FA8B" box="[1253,1436,1303,1329]" italics="true" pageId="11" pageNumber="412">S. strussmannae</emphasis>
</taxonomicName>
by having nostrils not protruding dorsolaterally, canthus rostralis poorly defined, straight or slightly convex, and loreal region slightly concave (nostrils protruding dorsolaterally, canthus rostralis well defined, curved medially, and loreal region markedly concave in 
<taxonomicName id="4C634D3DFFF4FFB0E91B01A8336CFA27" box="[588,770,1411,1437]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="strussmannae">
<emphasis id="B917EAACFFF4FFB0E91B01A8336CFA27" box="[588,770,1411,1437]" italics="true" pageId="11" pageNumber="412">S. strussmannae</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0E84601A83387FA26" author="Ferrao, M. &amp; Moravec, J. &amp; Kaefer, I. L. &amp; Fraga, R. &amp; Lima, A. P." box="[785,1001,1411,1437]" pageId="11" pageNumber="412" pagination="473 - 486" refId="ref11761" refString="Ferrao, M., Moravec, J., Kaefer, I. L., Fraga, R. &amp; Lima, A. P. (2018 a) New species of Scinax (Anura: Hylidae) with red-striped eyes from Brazilian Amazonia. Journal of Herpetology, 52, 473 - 486. https: // doi. org / 10.1670 / 17 - 165" type="journal article" year="2018">
Ferrão 
<emphasis id="B917EAACFFF4FFB0E83601A833F7FA26" box="[865,921,1411,1437]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2018a
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3796535FFF4FFB7EB90018C3492FE6A" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412" type="etymology">
<paragraph id="8BDC36BEFFF4FFB0EB90018C3330F9EA" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="412">
<emphasis id="B917EAACFFF4FFB0EB90018C3120FA7A" bold="true" box="[199,334,1447,1473]" pageId="11" pageNumber="412">Etymology.</emphasis>
The specific name 
<taxonomicName id="4C634D3DFFF4FFB0E974018C32F0FA7A" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[547,670,1447,1473]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF4FFB0E974018C32F0FA7A" box="[547,670,1447,1473]" italics="true" pageId="11" pageNumber="412">ushiniauae</emphasis>
</taxonomicName>
honors Meri del Pilar Ushiñahua Álvarez, a recently retired professor from the Faculty of Biological Sciences of the Universidad Nacional de la Amazonía Peruana—UNAP. During decades of teaching, she has stimulated students with her enthusiasm and joy, her passion for the conservation and management of Amazonia, and her studies of Amazonian turtles. Her academic legacy lives through the large number of students she trained throughout her academic career.
</paragraph>
<paragraph id="8BDC36BEFFF4FFB7EB9002713303FF6E" blockId="11.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,465]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
<emphasis id="B917EAACFFF4FFB0EB90027132B7F9CF" bold="true" box="[199,729,1626,1653]" pageId="11" pageNumber="412">Geographic distribution and natural history.</emphasis>
The new species was collected in two Peruvian localities, CIJH and RB, in the province of Requena, department of Loreto, covering an area of just over 
<quantity id="4C9B9B5BFFF4FFB0EF3B025434AFF922" box="[1132,1217,1663,1689]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="412" unit="km" value="100.0">100 km</quantity>
in a straight line, in which there are no large rivers that fragment the populations (
<figureCitation id="13582A3BFFF4FFB0E81A028833E1F906" box="[845,911,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1113,1137]" captionTargetBox="[156,1434,186,1077]" captionTargetId="figure-24@2.[151,1435,179,1087]" captionTargetPageId="2" captionText="FIGURE 1. Geographic distribution of Scinax ushiniauae. Left: Western Amazonia and neighboring areas (inset map indicates its location within South America) showing main rivers (blue lines), country borders (black contours), and known localities of Scinax ushiniauae. Right: satellite image of the Ucayali-Tapiche-Yavarí interfluve and neighboring regions with labeled main rivers, cities, and conservation areas shaded with light green. Abbreviations: CIJH, Centro de Investigaciones Jenaro-Herrera; RB, Río Blanco." figureDoi="http://doi.org/10.5281/zenodo.10628010" httpUri="https://zenodo.org/record/10628010/files/figure.png" pageId="11" pageNumber="412">Fig. 1</figureCitation>
). The CIJH comprises 
<emphasis id="B917EAACFFF4FFB0EFC30288357EF906" box="[1172,1296,1699,1725]" italics="true" pageId="11" pageNumber="412">terra-firme</emphasis>
(unflooded) forest environments growing on clay and white-sand soils and forests growing on hydromorphic, seasonally flooded soils. The rainy season spans approximately four to six months, with the heaviest rainfall from December to March. Mean annual precipitation is 
<quantity id="4C9B9B5BFFF4FFB0EA8F03253225F892" box="[472,587,1806,1833]" metricMagnitude="0" metricUnit="m" metricValue="2.521" pageId="11" pageNumber="412" unit="mm" value="2521.0">2,521 mm</quantity>
, and the average yearly air temperature is 26 °C (22.5–31.9°C; 
<bibRefCitation id="EFF24B4FFFF4FFB0EE530324312CF8F6" author="Lopez-Parodi, J. &amp; Freitas, D." pageId="11" pageNumber="412" pagination="157 - 168" refId="ref12647" refString="Lopez-Parodi, J. &amp; Freitas, D. (1990) Geographical aspects of forested wetlands in the lower Ucayali, Peruvian Amazonia. Forest ecology and management, 33 / 34, 157 - 168. https: // doi. org / 10.1016 / 0378 - 1127 (90) 90190 - M" type="journal article" year="1990">López-Parodi &amp; Freitas 1990</bibRefCitation>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EA19031831A9F8F6" author="IIAP" box="[334,455,1843,1869]" pageId="11" pageNumber="412" refId="ref12264" refString="IIAP (2023) Centro de Investigacion Jenaro Herrera - Jose Lopez Parodi. Available from: http: // www. iiap. org. pe / web / ci-jenaroherrera. aspx (accessed 9 August 2023)" type="url" year="2023">IIAP 2023</bibRefCitation>
). RB is located approximately 
<quantity id="4C9B9B5BFFF4FFB0E8770318331AF8F6" box="[800,884,1843,1869]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="412" unit="km" value="110.0">110 km</quantity>
south of CIJH, on the east bank of the TapicheBlanco interfluve. Part of this interfluve is constituted by large areas of white-sand forests and other vegetation 
<typeStatus id="54D8881CFFF4FFB0EBC0035030BCF82E" box="[151,210,1915,1941]" pageId="11" pageNumber="412">types</typeStatus>
characteristic of white-sand soils (known locally as 
<emphasis id="B917EAACFFF4FFB0E866035033F5F82E" box="[817,923,1915,1941]" italics="true" pageId="11" pageNumber="412">varillales</emphasis>
and 
<emphasis id="B917EAACFFF4FFB0E8820350343FF82E" box="[981,1105,1915,1941]" italics="true" pageId="11" pageNumber="412">chamizales</emphasis>
; 
<bibRefCitation id="EFF24B4FFFF4FFB0EF3403503554F82E" author="Pitman, N. &amp; Vriesendorp, C. &amp; Rivera, L. C. &amp; Wachter, T. &amp; Alvira-Reyes, D. &amp; Campo, A. &amp; Gagliardi-Urrutia, G. &amp; Rivera, D. G. &amp; Trevejo, L. &amp; Rivera, D. G. &amp; Heilpern, S." box="[1123,1338,1914,1941]" pageId="11" pageNumber="412" refId="ref13567" refString="Pitman, N., Vriesendorp, C., Rivera, L. C., Wachter, T., Alvira-Reyes, D., del Campo, A., Gagliardi-Urrutia, G., Rivera, D. G., Trevejo, L., Rivera, D. G. &amp; Heilpern, S. (Eds.) (2015) Peru: Tapiche-Blanco. Rapid Biological and Social Inventories Report 27. The Field Museum, Chicago, Illinois, 400 pp." type="book" year="2015">
Pitman 
<emphasis id="B917EAACFFF4FFB0EFEB03503498F82E" box="[1212,1270,1915,1941]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2015
</bibRefCitation>
; GarcíaVillacorta 
<emphasis id="B917EAACFFF4FFB0EA5A03B43128F802" box="[269,326,1951,1977]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2016). Hydromorphic soils and 
<emphasis id="B917EAACFFF4FFB0E9E103B4335CF802" box="[694,818,1951,1977]" italics="true" pageId="11" pageNumber="412">terra-firme</emphasis>
clay-rich soils are also common at this site, home to a megadiverse plant community (
<bibRefCitation id="EFF24B4FFFF4FFB0EAA003E832A4F866" author="Pitman, N. &amp; Vriesendorp, C. &amp; Rivera, L. C. &amp; Wachter, T. &amp; Alvira-Reyes, D. &amp; Campo, A. &amp; Gagliardi-Urrutia, G. &amp; Rivera, D. G. &amp; Trevejo, L. &amp; Rivera, D. G. &amp; Heilpern, S." box="[503,714,1986,2013]" pageId="11" pageNumber="412" refId="ref13567" refString="Pitman, N., Vriesendorp, C., Rivera, L. C., Wachter, T., Alvira-Reyes, D., del Campo, A., Gagliardi-Urrutia, G., Rivera, D. G., Trevejo, L., Rivera, D. G. &amp; Heilpern, S. (Eds.) (2015) Peru: Tapiche-Blanco. Rapid Biological and Social Inventories Report 27. The Field Museum, Chicago, Illinois, 400 pp." type="book" year="2015">
Pitman 
<emphasis id="B917EAACFFF4FFB0E91803E832EEF866" box="[591,640,1987,2013]" italics="true" pageId="11" pageNumber="412">et al</emphasis>
. 2015
</bibRefCitation>
). RB is also in the buffer zone of the Matsés National Reserve. A population that may belong to the new species occurs in the city of Leticia, Departament of Amazonas, 
<collectingCountry id="F374762EFFF3FFB7EE7904BC35F2FF0A" box="[1326,1436,151,177]" name="Colombia" pageId="12" pageNumber="413">Colombia</collectingCountry>
(J. M. Padial, pers. obs.), although its identity remains uncertain.
</paragraph>
<paragraph id="8BDC36BEFFF3FFB7EB9004F43492FE6A" blockId="12.[151,1437,151,465]" pageId="12" pageNumber="413">
Most individuals of 
<taxonomicName id="4C634D3DFFF3FFB7EAFA04F4322BFF42" authorityName="Gagliardi-Urrutia &amp; Araujo-Vieira &amp; Padial &amp; Simões &amp; Faivovich &amp; Castroviejo-Fisher" authorityYear="2024" box="[429,581,223,249]" class="Amphibia" family="Hylidae" genus="Scinax" kingdom="Animalia" order="Anura" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ushiniauae">
<emphasis id="B917EAACFFF3FFB7EAFA04F4322BFF42" box="[429,581,223,249]" italics="true" pageId="12" pageNumber="413">S. ushiniauae</emphasis>
</taxonomicName>
(55 %) were found in white-sand forest environments, while the remaining 45 % were located in forests growing on clay-rich soils (
<bibRefCitation id="EFF24B4FFFF3FFB7E87B05283457FEA6" author="Gagliardi-Urrutia, G." box="[812,1081,259,285]" pageId="12" pageNumber="413" refId="ref12063" refString="Gagliardi-Urrutia, G. (2020) El papel de los bosques de arena blanca en la diversidad de anfibios de la Amazonia Oeste. Ph. D. dissertation, Pontificia Universidade Catolica do Rio Grande do Sul, Porto Alegre. [unknown pagination]" type="book" year="2020">Gagliardi-Urrutia 2020</bibRefCitation>
). Males were found vocalizing in horizontal or vertical positions, with the head upwards, while perched on 
<emphasis id="B917EAACFFF3FFB7E8A8050C3407FEFA" box="[1023,1129,295,321]" italics="true" pageId="12" pageNumber="413">varillales</emphasis>
and 
<emphasis id="B917EAACFFF3FFB7EFF4050C3571FEFA" box="[1187,1311,295,321]" italics="true" pageId="12" pageNumber="413">chamizales</emphasis>
vegetation (
<quantity id="4C9B9B5BFFF3FFB7EBC80561317FFEDE" box="[159,273,330,357]" metricMagnitude="0" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="12" pageNumber="413" unit="m" value="1.1" valueMax="1.5" valueMin="0.7">0.7–1.5 m</quantity>
above ground) that grow on the margin or adjacent areas of temporary ponds located at the edges of old timber extraction trails inside the forest, or ponds in natural depressions in the forest, always associated with white-sand soils (
<figureCitation id="13582A3BFFF3FFB7EA4105B831FCFE16" box="[278,402,403,429]" captionStart="FIGURE 9" captionStartId="12.[151,250,1697,1721]" captionTargetBox="[255,1333,489,1674]" captionTargetId="figure-169@12.[255,1333,489,1674]" captionTargetPageId="12" captionText="FIGURE 9. Habitats of Scinax ushiniauae. (A) Trail in Centro de Investigaciones Jenaro Herrera (CIJH), Requena, Loreto, Peru. (B) Stational pond at CIJH." figureDoi="http://doi.org/10.5281/zenodo.10628026" httpUri="https://zenodo.org/record/10628026/files/figure.png" pageId="12" pageNumber="413">Fig. 9A, B</figureCitation>
). These ponds were formed when rainwater filled depressions, resulting in land subsidence processes. The new species shares the same reproductive habitats with 
<taxonomicName id="4C634D3DFFF3FFB7E8E5059C3492FE6A" authority="Heyer." authorityName="Heyer." box="[946,1276,439,465]" class="Amphibia" family="Leptodactylidae" genus="Leptodactylus" kingdom="Animalia" order="Anura" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="diedrus">
<emphasis id="B917EAACFFF3FFB7E8E5059C34C4FE6A" box="[946,1194,439,465]" italics="true" pageId="12" pageNumber="413">Leptodactylus diedrus</emphasis>
Heyer.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>