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<mods:title id="B65C95F1FE11FB5B23EBE292E332082B">Systematic position of two Athamanthia Zhdanko, 1983 (Lepidoptera, Lycaenidae) taxa from the Iranian Plateau</mods:title>
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<treatment id="03B287C19266B01985F0850FFB12CACA" ID-DOI="http://doi.org/10.5281/zenodo.6042237" ID-GBIF-Taxon="127697048" ID-Zenodo-Dep="6042237" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA" httpUri="http://treatment.plazi.org/id/03B287C19266B01985F0850FFB12CACA" lastPageId="5" lastPageNumber="580" pageId="4" pageNumber="579">
<subSubSection id="C301655C9266B01885F0850FFD88CA71" pageId="4" pageNumber="579" type="nomenclature">
<paragraph id="8BA436D79266B01885F0850FFBF8CA16" blockId="4.[151,1127,1816,1876]" box="[151,1127,1816,1843]" pageId="4" pageNumber="579">
<heading id="D0EC81BB9266B01885F0850FFBF8CA16" bold="true" box="[151,1127,1816,1843]" fontSize="11" level="1" pageId="4" pageNumber="579" reason="1">
<emphasis id="B96FEAC59266B01885F0850FFBF8CA16" bold="true" box="[151,1127,1816,1843]" pageId="4" pageNumber="579">
<taxonomicName id="4C1B4D549266B01885F0850FFC4CCA16" authority="Eckweiler &amp; ten Hagen, 2001" authorityName="Eckweiler &amp; ten Hagen" authorityYear="2001" box="[151,979,1816,1843]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="balucha" status="comb. nov." subSpecies="athamantides">
<emphasis id="B96FEAC59266B01885F0850FFDD5CA17" bold="true" box="[151,586,1816,1842]" italics="true" pageId="4" pageNumber="579">Athamanthia balucha athamantides</emphasis>
(Eckweiler &amp; ten Hagen, 2001)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57BE9266B0188687850EFBF8CA16" box="[992,1127,1817,1843]" pageId="4" pageNumber="579">comb. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BA436D79266B01885F0852CFD88CA71" blockId="4.[151,1127,1816,1876]" box="[151,535,1851,1876]" pageId="4" pageNumber="579">(Plate 1, figs. 910; plate 2, fig. 4)</paragraph>
</subSubSection>
<subSubSection id="C301655C9266B01885F08568FB88CAB1" box="[151,1047,1918,1940]" pageId="4" pageNumber="579" type="description">
<paragraph id="8BA436D79266B01885F08568FB88CAB1" blockId="4.[151,1413,1918,2032]" box="[151,1047,1918,1940]" pageId="4" pageNumber="579">
<taxonomicName id="4C1B4D549266B01885F08568FD37CAB1" box="[151,680,1918,1940]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="phoenicurus" subGenus="Phoenicurusia" subSpecies="athamantides">
<emphasis id="B96FEAC59266B01885F08568FF73CAB1" box="[151,236,1919,1940]" italics="true" pageId="4" pageNumber="579">Lycaena</emphasis>
(
<emphasis id="B96FEAC59266B018859C8569FE10CAB1" box="[251,399,1918,1940]" italics="true" pageId="4" pageNumber="579">Phoenicurusia</emphasis>
)
<emphasis id="B96FEAC59266B01884FA8569FD37CAB1" box="[413,680,1918,1940]" italics="true" pageId="4" pageNumber="579">phoenicurus athamantides</emphasis>
</taxonomicName>
—Eckweiler &amp; ten Hagen 2001: 49.
</paragraph>
</subSubSection>
<subSubSection id="C301655C9266B01885F0858BFA1ACAD5" pageId="4" pageNumber="579" type="reference_group">
<paragraph id="8BA436D79266B01885F0858BFA1ACAD5" blockId="4.[151,1413,1918,2032]" pageId="4" pageNumber="579">
<taxonomicName id="4C1B4D549266B01885F0858BFCFBCA97" authority="Eckweiler &amp; ten Hagen, 2001" authorityName="Eckweiler &amp; ten Hagen" authorityYear="2001" box="[151,868,1948,1970]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="phoenicura" subSpecies="athamantides">
<emphasis id="B96FEAC59266B01885F0858BFDBDCA97" box="[151,546,1948,1970]" italics="true" pageId="4" pageNumber="579">Athamanthia phoenicura athamantides</emphasis>
(Eckweiler &amp; ten Hagen, 2001)
</taxonomicName>
<bibRefCitation id="EF8A4B269266B0188618858BFB2BCA97" author="Bozano" box="[895,1204,1948,1970]" pageId="4" pageNumber="579" refString="Bozano, G. C. &amp; Weidenhoffer, Z. (2001) Guide to the Butterflies of the Palearctic Region. Lycaenidae part I. Lycaeninae. Omnes Artes, Milano, 62 pp." type="book" year="2001">Bozano &amp; Weidenhoffer 2001</bibRefCitation>
: 40.
<taxonomicName id="4C1B4D549266B01885F085ABFCAFCAF4" authority="Eckweiler &amp; ten Hagen, 2001" authorityName="Eckweiler &amp; ten Hagen" authorityYear="2001" box="[151,816,1979,2001]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="phoenicurus" subSpecies="athamantides">
<emphasis id="B96FEAC59266B01885F085ABFE61CAF4" box="[151,510,1979,2001]" italics="true" pageId="4" pageNumber="579">Lycaena phoenicurus athamantides</emphasis>
Eckweiler &amp; ten Hagen, 2001
</taxonomicName>
<bibRefCitation id="EF8A4B269266B018862D85ACFC53CAF4" author="Nazari" box="[842,972,1979,2001]" pageId="4" pageNumber="579" refString="Nazari, V. (2003) Butterflies of Iran. Dayereh-Sabz, National Museum of Natural History of the Islamic Republic of Iran, Tehran, 568 pp. [in Persian]" type="book" year="2003">Nazari 2003</bibRefCitation>
: 122; col. pl. 22 23, figs. 4, 5.
<taxonomicName id="4C1B4D549266B01885F085CCFCAFCAD5" authority="Eckweiler &amp; ten Hagen, 2001" authorityName="Eckweiler &amp; ten Hagen" authorityYear="2001" box="[151,816,2010,2032]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="579" phylum="Arthropoda" rank="subSpecies" species="phoenicurus" subSpecies="athamantides">
<emphasis id="B96FEAC59266B01885F085CCFE61CAD5" box="[151,510,2010,2032]" italics="true" pageId="4" pageNumber="579">Lycaena phoenicurus athamantides</emphasis>
Eckweiler &amp; ten Hagen, 2001
</taxonomicName>
<bibRefCitation id="EF8A4B269266B018862D85CDFBDECAD5" author="Tshikolovets" box="[842,1089,2010,2032]" pageId="4" pageNumber="579" refString="Tshikolovets, V., Naderi, A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, 440 pp." type="book" year="2014">
Tshikolovets
<emphasis id="B96FEAC59266B01886B485CCFC60CAD5" box="[979,1023,2010,2032]" italics="true" pageId="4" pageNumber="579">et al</emphasis>
. 2014
</bibRefCitation>
: 134; col. pl. XXII, figs. 15, 16.
</paragraph>
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<emphasis id="B96FEAC59267B01985F08280FEAACD95" bold="true" box="[151,309,151,176]" pageId="5" pageNumber="580">
<typeStatus id="54A088759267B01985F08280FF4DCD95" box="[151,210,151,176]" pageId="5" pageNumber="580">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="49DFF8359267B019845B8280FE10CD95" box="[316,399,151,176]" country="Sri Lanka" name="Central" pageId="5" pageNumber="580">Central</collectingRegion>
<collectingCountry id="F30C76479267B01984F2828FFE5ACD8A" box="[405,453,152,175]" name="Iran" pageId="5" pageNumber="580">Iran</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B733C8A9267B01984B78280FCBECD8A" ID-GBIF-Occurrence="1434656517" box="[464,801,151,176]" country="Iran" location="Shir Kuh Mts." pageId="5" pageNumber="580" specimenCount="1" stateProvince="Yazd Province" typeStatus="holotype">
<collectingRegion id="49DFF8359267B01984B78280FDE8CD95" box="[464,631,151,176]" country="Iran" name="Yazd" pageId="5" pageNumber="580">Yazd Province</collectingRegion>
,
<location id="8EC4600C9267B01987E68280FCBECD8A" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B01987E68280FCBECD8A" box="[641,801,151,176]" country="Iran" name="Shir Kuh Mts." pageId="5" pageNumber="580" stateProvince="Yazd Province">Shir Kuh Mts.</location>
</materialsCitation>
</paragraph>
<paragraph id="8BA436D79267B01985A082ABFDE9CDDD" blockId="5.[151,1436,151,1221]" pageId="5" pageNumber="580">
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<emphasis id="B96FEAC59267B01985A082ABFE2ECDF0" bold="true" box="[199,433,188,213]" pageId="5" pageNumber="580">Material examined.</emphasis>
<specimenCount id="9D1DFD5E9267B01984D082ABFE7CCDF1" box="[439,483,188,212]" pageId="5" pageNumber="580" type="male">1 ♂</specimenCount>
, South
<collectingRegion id="49DFF8359267B019875F82AAFD13CDF1" box="[568,652,189,212]" country="American Samoa" name="Eastern" pageId="5" pageNumber="580">Eastern</collectingRegion>
<collectingCountry id="F30C76479267B01987F382AAFD5BCDF1" box="[660,708,189,212]" name="Iran" pageId="5" pageNumber="580">Iran</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B733C8A9267B01987A882AAFDE9CDDD" ID-GBIF-Occurrence="1434656528" collectingDate="2014-06-19" collectorName="I. G. Pljushtch" country="Iran" elevation="2450" location="Makhan" pageId="5" pageNumber="580" specimenCount="1" stateProvince="Kerman Province">
<collectingRegion id="49DFF8359267B01987A882AAFC08CDF0" box="[719,919,188,213]" country="Iran" name="Kerman" pageId="5" pageNumber="580">Kerman Province</collectingRegion>
,
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SE
<location id="8EC4600C9267B019817F82ABFBE8CDF0" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B019817F82ABFBE8CDF0" box="[1048,1143,188,213]" country="Iran" name="Makhan" pageId="5" pageNumber="580" stateProvince="Kerman Province">Makhan</location>
,
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</elevation>
,
<collectingDate id="EFE1E9FF9267B019818182ABFAF7CDF1" box="[1254,1384,188,212]" pageId="5" pageNumber="580" value="2014-06-19">
<date id="FFA510179267B019818182ABFAF7CDF1" box="[1254,1384,188,212]" pageId="5" pageNumber="580" value="2014-06-19">19.VI.2014</date>
</collectingDate>
,
<collectorName id="26EE53019267B019801382ABFF60CDDD" pageId="5" pageNumber="580">I.G. Pljushtch</collectorName>
leg., gen. prep. AK0068 (AKM).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C301655C9267B01985A08313FB12CACA" pageId="5" pageNumber="580" type="diagnosis">
<paragraph id="8BA436D79267B01985A08313FAD3CF3D" blockId="5.[151,1436,151,1221]" pageId="5" pageNumber="580">
<emphasis id="B96FEAC59267B01985A08313FE67CC38" bold="true" box="[199,504,260,285]" pageId="5" pageNumber="580">Diagnosis and discussion.</emphasis>
The taxon was originally described as a subspecies of
<taxonomicName id="4C1B4D549267B019810E8313FA96CC39" box="[1129,1289,260,284]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="phoenicura">
<emphasis id="B96FEAC59267B019810E8313FA96CC39" box="[1129,1289,260,284]" italics="true" pageId="5" pageNumber="580">A. phoenicura</emphasis>
</taxonomicName>
based solely on the wings colouration (Eckweiler &amp; ten Hagen 2001). According to the original description, the main diagnostic character of the subspecies is extended orange field on the forewing upperside of a female compared with
<taxonomicName id="4C1B4D549267B01980E3835BFE88CCAD" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="phoenicura">
<emphasis id="B96FEAC59267B01980E3835BFE88CCAD" italics="true" pageId="5" pageNumber="580">A. phoenicura</emphasis>
</taxonomicName>
. In 2014, a new population of
<emphasis id="B96FEAC59267B01987118367FC92CCAD" box="[630,781,368,392]" italics="true" pageId="5" pageNumber="580">athamantides</emphasis>
was discovered in
<collectingRegion id="49DFF8359267B01986808367FB32CCAD" box="[999,1197,367,392]" country="Iran" name="Kerman" pageId="5" pageNumber="580">Kerman Province</collectingRegion>
(
<quantity id="4CE39B329267B01981DA8367FA99CCAD" box="[1213,1286,367,392]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="580" unit="km" value="20.0">20 km</quantity>
SE Makhan) by the two junior authors and O. Pak. Our examination of the appearance and the male genitalia compared with
<taxonomicName id="4C1B4D549267B01980E38383FF6ECCF5" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B01980E38383FF6ECCF5" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
has revealed minor differences in the shape of valva and uncus (Plate 2, Fig. 4). Based on these differences and wide geographic discontinuity between Afghan and Iranian populations, we tentatively treat known Iranian populations as a subspecies of
<emphasis id="B96FEAC59267B01984888017FD1ACF32" box="[495,645,512,536]" italics="true" pageId="5" pageNumber="580">
<taxonomicName id="4C1B4D549267B01984888017FDF7CF3D" box="[495,616,512,536]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">A. balucha</taxonomicName>
</emphasis>
<taxonomicName id="4C1B4D549267B01987E38017FB5FCF3D" authority="Eckweiler &amp; ten Hagen, 2001" authorityName="Eckweiler &amp; ten Hagen" authorityYear="2001" box="[644,1216,511,536]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="subSpecies" species="phoenicurus" status="comb. nov." subSpecies="athamantides">
<emphasis id="B96FEAC59267B01987E38017FCC8CF3D" box="[644,855,512,536]" italics="true" pageId="5" pageNumber="580">A. b. athamantides</emphasis>
(Eckweiler &amp; ten Hagen, 2001)
</taxonomicName>
,
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<emphasis id="B96FEAC59267B01981AB83E8FAD3CF3D" bold="true" box="[1228,1356,511,536]" pageId="5" pageNumber="580">comb. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BA436D79267B01985A08033FE18CF82" blockId="5.[151,1436,151,1221]" pageId="5" pageNumber="580">
A potential
<taxonomicName id="4C1B4D549267B01984328033FDAECF19" box="[341,561,548,572]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="subSpecies" species="phoenicurus" subSpecies="athamantides">
<emphasis id="B96FEAC59267B01984328033FDAECF19" box="[341,561,548,572]" italics="true" pageId="5" pageNumber="580">A. b. athamantides</emphasis>
</taxonomicName>
population from
<collectingRegion id="49DFF8359267B01986658032FC53CF18" box="[770,972,548,573]" country="Iran" name="Kerman" pageId="5" pageNumber="580">Kerman Province</collectingRegion>
(vic. of Sar-e Siab) was mentioned by
<bibRefCitation id="EF8A4B269267B01985F08050FE24CF45" author="Tshikolovets" box="[151,443,583,608]" pageId="5" pageNumber="580" refString="Tshikolovets, V., Naderi, A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, 440 pp." type="book" year="2014">
Tshikolovets
<emphasis id="B96FEAC59267B0198457805EFEF3CF45" box="[304,364,584,608]" italics="true" pageId="5" pageNumber="580">et al.</emphasis>
(2014
</bibRefCitation>
: 133) as “
<taxonomicName id="4C1B4D549267B0198752805EFBFCCF45" authority="Eckweiler, 2004" authorityName="Eckweiler" authorityYear="2004" box="[565,1123,583,608]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="subSpecies" species="phoenicurus" subSpecies="monalisa">
<emphasis id="B96FEAC59267B0198752805EFC05CF45" box="[565,922,584,608]" italics="true" pageId="5" pageNumber="580">Lycaena phoenicurus monalisa</emphasis>
<bibRefCitation id="EF8A4B269267B01986C38050FBFCCF45" author="Eckweiler" box="[932,1123,583,608]" pageId="5" pageNumber="580" refString="Eckweiler, W. (2004) Lycaena (Phoenicurusia) phoenicurus monalisa ssp. n. (Lepidoptera: Lycaenidae). Nachrichten des Entomologischen Vereins Apollo, 25 (3), 104." type="journal article" year="2004">Eckweiler, 2004</bibRefCitation>
</taxonomicName>
”. The taxon was described from
<collectingRegion id="49DFF8359267B01985B3807BFE62CFA0" box="[212,509,620,645]" country="Iran" name="Khorasan-e Razavi" pageId="5" pageNumber="580">Razavi Khorasan Province</collectingRegion>
(mountains near Tabas) (
<bibRefCitation id="EF8A4B269267B0198671807BFC52CFA1" author="Eckweiler" box="[790,973,620,645]" pageId="5" pageNumber="580" refString="Eckweiler, W. (2004) Lycaena (Phoenicurusia) phoenicurus monalisa ssp. n. (Lepidoptera: Lycaenidae). Nachrichten des Entomologischen Vereins Apollo, 25 (3), 104." type="journal article" year="2004">Eckweiler 2004</bibRefCitation>
). Its status needs clarification; it may also belong to
<taxonomicName id="4C1B4D549267B01984608087FE1FCF8D" box="[263,384,656,680]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B01984608087FE1FCF8D" box="[263,384,656,680]" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA436D79267B01985A080A3FCACC910" blockId="5.[151,1436,151,1221]" pageId="5" pageNumber="580">
<materialsCitation id="3B733C8A9267B01985A080A3FD38CEED" ID-GBIF-Occurrence="1434656529" country="Iran" county="Sulaiman Mts." location="Kuh-e Taftan Mt." municipality="Balochistan Province" pageId="5" pageNumber="580" specimenCount="2" stateProvince="Kabul" typeStatus="holotype">
<emphasis id="B96FEAC59267B01985A080A3FDCCCFE8" bold="true" box="[199,595,692,717]" pageId="5" pageNumber="580">General distribution and biology.</emphasis>
As it is known so far, the range of
<taxonomicName id="4C1B4D549267B01986BC80A3FBCACFE9" box="[987,1109,692,716]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B01986BC80A3FBCACFE9" box="[987,1109,692,716]" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
is strongly disjunct (plate 3). The nominate subspecies is known from the
<typeStatus id="54A088759267B01987C280CFFD4ACFD5" box="[677,725,728,752]" pageId="5" pageNumber="580">type</typeStatus>
locality and two localities in the
<collectingCounty id="62C54E5B9267B019810480C0FA94CFCA" box="[1123,1291,727,752]" pageId="5" pageNumber="580">Sulaiman Mts.</collectingCounty>
in Pakistani Balochistan mentioned in the original description (
<bibRefCitation id="EF8A4B269267B019879E80EBFBE9CE30" author="Howarth" box="[761,1142,764,789]" pageId="5" pageNumber="580" refString="Howarth, T. G. &amp; Povolny, D. (1976) Beitrage zur Kenntnis der Fauna afghanistans. Casopis Moravskeho musea v Brne, 61, 139 - 170." type="journal article" year="1976">Howarth &amp; Povolny 1976: 150</bibRefCitation>
). A specimen treated as “
<taxonomicName id="4C1B4D549267B01985C48136FE0BCE1D" box="[163,404,800,824]" class="Insecta" family="Lycaenidae" genus="Lycaena" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="phoenicurus">
<emphasis id="B96FEAC59267B01985C48136FE0BCE1D" box="[163,404,800,824]" italics="true" pageId="5" pageNumber="580">Lycaena phoenicurus</emphasis>
</taxonomicName>
ssp.” from Sistan and
<collectingMunicipality id="6BC0ACAD9267B01987F08108FC18CE1D" box="[663,903,799,824]" pageId="5" pageNumber="580">Balochistan Province</collectingMunicipality>
in
<collectingCountry id="F30C76479267B01986CA8137FC45CE12" box="[941,986,800,823]" name="Iran" pageId="5" pageNumber="580">Iran</collectingCountry>
(
<location id="8EC4600C9267B019868D8108FB2ECE1D" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B019868D8108FB2ECE1D" box="[1002,1201,799,824]" country="Iran" county="Sulaiman Mts." municipality="Balochistan Province" name="Kuh-e Taftan Mt." pageId="5" pageNumber="580" stateProvince="Kabul">Kuh-e Taftan Mt.</location>
) was mentioned and pictured in
<bibRefCitation id="EF8A4B269267B019847B8153FDA2CE79" author="Tshikolovets" box="[284,573,836,861]" pageId="5" pageNumber="580" refString="Tshikolovets, V., Naderi, A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Tshikolovets Publications, Pardubice, 440 pp." type="book" year="2014">
Tshikolovets
<emphasis id="B96FEAC59267B01984D38152FE70CE79" box="[436,495,836,860]" italics="true" pageId="5" pageNumber="580">et al.</emphasis>
(2014
</bibRefCitation>
: 134; plate XXII, fig. 17). It shares all external diagnostic characters of
<taxonomicName id="4C1B4D549267B01980E38153FF6ECEA5" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B01980E38153FF6ECEA5" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
and probably belongs to the species in question, while its subspecific affinity is open to discussion.
<emphasis id="B96FEAC59267B0198001817FFF70CE81" italics="true" pageId="5" pageNumber="580">A. b. povolny</emphasis>
inhabits western foothills of the
<location id="8EC4600C9267B019873A819BFCBECE81" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B019873A819BFCBECE81" box="[605,801,908,933]" country="Iran" county="Sulaiman Mts." municipality="Balochistan Province" name="Hindu Kush Mts." pageId="5" pageNumber="580" stateProvince="Kabul">Hindu Kush Mts.</location>
: vicinity of
<collectingRegion id="49DFF8359267B01986C2819BFC76CE80" box="[933,1001,908,933]" country="Afghanistan" name="Kabul" pageId="5" pageNumber="580">Kabul</collectingRegion>
(
<typeStatus id="54A088759267B0198690819BFBB6CE81" box="[1015,1065,908,932]" pageId="5" pageNumber="580">type</typeStatus>
locality) and recently discovered localities in province
<collectingRegion id="49DFF8359267B01984F481A7FE6FCEED" box="[403,496,944,968]" country="Afghanistan" name="Bamyan" pageId="5" pageNumber="580">Bamyan</collectingRegion>
in
<collectingCountry id="F30C76479267B019877C81B8FD38CEED" box="[539,679,943,968]" name="Afghanistan" pageId="5" pageNumber="580">Afghanistan</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B733C8A9267B01987D181B8FCACC910" ID-GBIF-Occurrence="1434656562" country="Iran" county="Subspecies" elevation="3000" location="Shir Kuh Mts." municipality="Iranian" pageId="5" pageNumber="580" specimenCount="1" stateProvince="Yazd" typeStatus="holotype">
<collectingCounty id="62C54E5B9267B01987D181B8FCACCEED" box="[694,819,943,968]" pageId="5" pageNumber="580">Subspecies</collectingCounty>
<emphasis id="B96FEAC59267B019865B81A7FC4DCEED" box="[828,978,944,968]" italics="true" pageId="5" pageNumber="580">athamantides</emphasis>
is known from
<collectingMunicipality id="6BC0ACAD9267B01981F481B8FB7DCEED" box="[1171,1250,943,968]" pageId="5" pageNumber="580">Iranian</collectingMunicipality>
provinces
<collectingRegion id="49DFF8359267B019800381B8FA03CEED" box="[1380,1436,943,968]" country="Iran" name="Yazd" pageId="5" pageNumber="580">Yazd</collectingRegion>
(
<location id="8EC4600C9267B01985F981C3FEDECEC9" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B01985F981C3FEDECEC9" box="[158,321,980,1005]" country="Iran" county="Subspecies" municipality="Iranian" name="Shir Kuh Mts." pageId="5" pageNumber="580" stateProvince="Yazd">Shir Kuh Mts.</location>
) and
<collectingRegion id="49DFF8359267B019841881C2FE46CEC9" box="[383,473,981,1004]" country="Iran" name="Kerman" pageId="5" pageNumber="580">Kerman</collectingRegion>
(
<collectingRegion id="49DFF8359267B019848F81C3FDDBCEC9" box="[488,580,980,1004]" country="Iran" name="Kerman" pageId="5" pageNumber="580">Kerman</collectingRegion>
Mts.).
<location id="8EC4600C9267B01987F281C3FD52CEC8" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B01987F281C3FD52CEC8" box="[661,717,980,1005]" country="Iran" county="Subspecies" municipality="Iranian" name="Thus" pageId="5" pageNumber="580" stateProvince="Yazd">Thus</location>
<taxonomicName id="4C1B4D549267B01987B381C3FCD2CEC9" box="[724,845,980,1004]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B01987B381C3FCD2CEC9" box="[724,845,980,1004]" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
can be treated as an endemic of the
<location id="8EC4600C9267B019818381C3FAACCEC8" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B019818381C3FAACCEC8" box="[1252,1331,980,1005]" country="Iran" county="Subspecies" municipality="Iranian" name="Iranian" pageId="5" pageNumber="580" stateProvince="Yazd">Iranian</location>
plateau.
<location id="8EC4600C9267B01985F081EFFF69C935" LSID="urn:lsid:plazi:treatment:03B287C19266B01985F0850FFB12CACA:8EC4600C9267B01985F081EFFF69C935" box="[151,246,1016,1040]" country="Iran" county="Subspecies" municipality="Iranian" name="Imagoes" pageId="5" pageNumber="580" stateProvince="Yazd">Imagoes</location>
prefer dry slopes of mountains at altitudes up to
<elevation id="0036D1E49267B019865281EFFC10C92A" box="[821,911,1016,1040]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="580" unit="m" value="3000.0">
<quantity id="4CE39B329267B019865281EFFC10C92A" box="[821,911,1016,1040]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="580" unit="m" value="3000.0">3000 m</quantity>
</elevation>
. Flight period is from the end of April to the beginning of July, depending on a locality and an elevation.
</materialsCitation>
</paragraph>
<paragraph id="8BA436D79267B01985A08628FC1EC9E0" blockId="5.[151,1436,151,1221]" pageId="5" pageNumber="580">
All known
<taxonomicName id="4C1B4D549267B01984218657FE48C97D" box="[326,471,1088,1112]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEAC59267B01984218657FE48C97D" box="[326,471,1088,1112]" italics="true" pageId="5" pageNumber="580">Athamanthia</emphasis>
</taxonomicName>
are monophagous species feeding on the genus
<taxonomicName id="4C1B4D549267B01986908657FAB7C97D" box="[1015,1320,1087,1112]" class="Magnoliopsida" family="Polygonaceae" genus="Atraphaxis" kingdom="Plantae" order="Polygonales" pageId="5" pageNumber="580" phylum="Tracheophyta" rank="genus">
<emphasis id="B96FEAC59267B01986908657FBEDC97D" box="[1015,1138,1088,1112]" italics="true" pageId="5" pageNumber="580">Atraphaxis</emphasis>
(Polygonaceae)
</taxonomicName>
(
<bibRefCitation id="EF8A4B269267B01980518628FF4EC959" author="Zhdanko" pageId="5" pageNumber="580" refString="Zhdanko, A. B. (2000) Genus Athamanthia. In: Tuzov, V. K., Bogdanov, P. V., Churkin, S. V., Dantchenko, A. V., Devyatkin, A. L., Murzin, V. S., Samodurov, G. D. &amp; Zhdanko, A. B. (Eds.), Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Lycaenidae). Vol. 2. Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae. Pensoft, Sofia-Moscow, pp. 131 - 135." type="book" year="2000">Zhdanko 2000</bibRefCitation>
). The most probable hostplant of
<taxonomicName id="4C1B4D549267B019873D8673FD49C959" box="[602,726,1124,1148]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B019873D8673FD49C959" box="[602,726,1124,1148]" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
is
<taxonomicName id="4C1B4D549267B019879B8673FC4AC959" box="[764,981,1124,1148]" class="Magnoliopsida" family="Polygonaceae" genus="Atraphaxis" kingdom="Plantae" order="Polygonales" pageId="5" pageNumber="580" phylum="Tracheophyta" rank="species" species="spinosa">
<emphasis id="B96FEAC59267B019879B8673FC4AC959" box="[764,981,1124,1148]" italics="true" pageId="5" pageNumber="580">Atraphaxis spinosa</emphasis>
</taxonomicName>
, which is known as the hostplant of
<taxonomicName id="4C1B4D549267B01980E38673FE49C985" authority="Zhdanko 2000" authorityName="Zhdanko" authorityYear="2000" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="phoenicura">
<emphasis id="B96FEAC59267B01980E38673FE88C985" italics="true" pageId="5" pageNumber="580">A. phoenicura</emphasis>
(
<bibRefCitation id="EF8A4B269267B01984428690FE50C985" author="Zhdanko" box="[293,463,1159,1184]" pageId="5" pageNumber="580" refString="Zhdanko, A. B. (2000) Genus Athamanthia. In: Tuzov, V. K., Bogdanov, P. V., Churkin, S. V., Dantchenko, A. V., Devyatkin, A. L., Murzin, V. S., Samodurov, G. D. &amp; Zhdanko, A. B. (Eds.), Guide to the butterflies of Russia and adjacent territories (Lepidoptera, Lycaenidae). Vol. 2. Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae. Pensoft, Sofia-Moscow, pp. 131 - 135." type="book" year="2000">Zhdanko 2000</bibRefCitation>
)
</taxonomicName>
. According to
<bibRefCitation id="EF8A4B269267B019871B8690FBBFC985" author="Rechinger" box="[636,1056,1159,1184]" pageId="5" pageNumber="580" refString="Rechinger, K. H. &amp; Schiman-Czeika, H. (1968) Atraphaxis. In: Rechinger, K. H. (Ed.), Flora Iranica. Vol. 56. Polygonaceae. Akademische Druck- und Verlagsanstalt, Graz, pp. 30 - 35." type="book" year="1968">Rechinger &amp; Schiman-Czeika (1968)</bibRefCitation>
, it has a wide range in the Iranian Plateau from NE
<collectingCountry id="F30C76479267B019843D86BAFE18C9E1" box="[346,391,1197,1220]" name="Iran" pageId="5" pageNumber="580">Iran</collectingCountry>
to Pakistani
<collectingRegion id="49DFF8359267B019877F86BBFD02C9E0" box="[536,669,1196,1221]" country="Pakistan" name="Balochistan" pageId="5" pageNumber="580">Balochistan</collectingRegion>
across
<collectingCountry id="F30C76479267B019879686BBFCE2C9E0" box="[753,893,1196,1221]" name="Afghanistan" pageId="5" pageNumber="580">Afghanistan</collectingCountry>
.
</paragraph>
<paragraph id="8BA436D79267B01985F085CEFB12CACA" blockId="5.[151,1165,2009,2031]" box="[151,1165,2009,2031]" pageId="5" pageNumber="580">
<emphasis id="B96FEAC59267B01985F085CEFE9DCACA" bold="true" box="[151,258,2009,2031]" pageId="5" pageNumber="580">PLATE 3.</emphasis>
Distribution of
<taxonomicName id="4C1B4D549267B01984C285CDFD8DCACA" box="[421,530,2009,2031]" class="Insecta" family="Lycaenidae" genus="Athamanthia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="balucha">
<emphasis id="B96FEAC59267B01984C285CDFD8DCACA" box="[421,530,2009,2031]" italics="true" pageId="5" pageNumber="580">A. balucha</emphasis>
</taxonomicName>
. Squares indicate on the type localities of the taxa in question.
</paragraph>
</subSubSection>
</treatment>
</document>