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<document id="8AD24BB88012361E3DAC67D3A8A23E5B" ID-CLB-Dataset="55622" ID-DOI="10.5852/ejt.2022.819.1773" ID-GBIF-Dataset="ad96f711-d952-4f41-8241-5646ea1751aa" ID-ISSN="2118-9773" ID-Zenodo-Dep="6533014" ID-ZooBank="6A5B371D-2A28-43C1-A6C3-12C8E7016515" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652116231013" checkinUser="felipe" docAuthor="Żyła, Dagmara, Tokareva, Alexandra &amp; Koszela, Katarzyna" docDate="2022" docId="03B287A6FFA9FFF2FDC1FB75FB7BF967" docLanguage="en" docName="ejt-819_1-22.pdf" docOrigin="European Journal of Taxonomy 819 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.819.1773" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Acrostilicus hospes Hubbard 1896" docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FF8BFFDEFFAEFFFBFF8BFFBAFFEFFFB6" masterDocTitle="Phylogenetic position of genera Acrostilicus Hubbard and Pachystilicus Casey (Staphylinidae, Paederinae) and their redescription" masterLastPageNumber="22" masterPageNumber="1" pageNumber="8" updateTime="1699330068693" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4F7FD25E5A2B4A014B80C22412B13F45">Phylogenetic position of genera Acrostilicus Hubbard and Pachystilicus Casey (Staphylinidae, Paederinae) and their redescription</mods:title>
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<mods:namePart id="98B21ECDA287D34B7D5A386DAC1F6432">Żyła, Dagmara</mods:namePart>
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<mods:affiliation id="ED2CEBF97705B5F7D231CC69A3ADF81A">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warsaw, Poland. Department of Invertebrate Zoology and Parasitology, University of Gdańsk, 80 - 308, Gdańsk, Poland. Department of Ecology, Evolution and Organismal Biology, Iowa State University, Ames, IA 50011, USA.</mods:affiliation>
<mods:nameIdentifier id="165FD463751A34B6F0770D6B469E3DDE" type="email">zyladagmara@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2D20095EFF2A0E36605F35D1F08FC0EB">Tokareva, Alexandra</mods:namePart>
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<mods:affiliation id="B8316D3C8FFA6122F7D67A38CCABA6B8">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warsaw, Poland. Department of Invertebrate Zoology and Parasitology, University of Gdańsk, 80 - 308, Gdańsk, Poland. Department of Ecology, Evolution and Organismal Biology, Iowa State University, Ames, IA 50011, USA.</mods:affiliation>
<mods:nameIdentifier id="00EAA173E62CF9B8004BAEF01F91CE17" type="email">alexan4tok@gmail.com</mods:nameIdentifier>
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<mods:namePart id="90938F0407BCBFD1640A0B3F2DC7DCB5">Koszela, Katarzyna</mods:namePart>
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<mods:affiliation id="05D0AFCA2FBA7CEEE9441BFA2C021920">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warsaw, Poland. Department of Invertebrate Zoology and Parasitology, University of Gdańsk, 80 - 308, Gdańsk, Poland. Department of Ecology, Evolution and Organismal Biology, Iowa State University, Ames, IA 50011, USA.</mods:affiliation>
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<treatment id="03B287A6FFA9FFF2FDC1FB75FB7BF967" ID-DOI="http://doi.org/10.5281/zenodo.6533964" ID-GBIF-Taxon="195278030" ID-Zenodo-Dep="6533964" LSID="urn:lsid:plazi:treatment:03B287A6FFA9FFF2FDC1FB75FB7BF967" httpUri="http://treatment.plazi.org/id/03B287A6FFA9FFF2FDC1FB75FB7BF967" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
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<paragraph id="8BA436B0FFA9FFFCFDC1FB75FC05FB5F" blockId="7.[586,1002,1231,1292]" box="[586,1002,1231,1258]" pageId="7" pageNumber="8">
<heading id="D0EC81DCFFA9FFFCFDC1FB75FC05FB5F" box="[586,1002,1231,1258]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C1B4D33FFA9FFFCFDC1FB75FC05FB5F" ID-CoL="9QPT" authority="Hubbard, 1896" authorityName="Hubbard" authorityYear="1896" box="[586,1002,1231,1258]" class="Insecta" family="Staphylinidae" genus="Acrostilicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hospes">
<emphasis id="B96FEAA2FFA9FFFCFDC1FB75FCDDFB5C" bold="true" box="[586,818,1231,1258]" italics="true" pageId="7" pageNumber="8">Acrostilicus hospes</emphasis>
<bibRefCitation id="EF8A4B41FFA9FFFCFCB1FB75FC05FB5F" author="Hubbard H. G." box="[826,1002,1231,1257]" pageId="7" pageNumber="8" pagination="299 - 302" refId="ref12173" refString="Hubbard H. G. 1896. Additional notes on the insect guests of the Florida land tortoise. Proceedings of the Entomological Society of Washington 3: 299 - 302." type="journal article" year="1896">Hubbard, 1896</bibRefCitation>
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<figureCitation id="13202A35FFA9FFFCFD7CFB48FCD3FABA" box="[759,828,1266,1292]" captionStart="Fig" captionStartId="8.[189,232,1916,1942]" captionTargetBox="[311,1251,1092,1878]" captionTargetId="figure-270@8.[311,1277,1092,1878]" captionTargetPageId="8" captionText="Fig. 2. Acrostilicus hospes Hubbard, 1896, habitus photograph and drawings of the apical abdominal sternite and genital structures. A. Habitus, lectotype, ♂ (NMNH). B. Male sternite VIII. C. Aedeagus, parameral view. D. Aedeagus, lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6533022" httpUri="https://zenodo.org/record/6533022/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
</paragraph>
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<paragraph id="8BA436B0FFA9FFFCFF36FA8EFC75FAF9" blockId="7.[189,922,1332,1359]" box="[189,922,1332,1359]" pageId="7" pageNumber="8">
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<treatmentCitation id="0ABA10A1FFA9FFFCFF36FA8EFD7DFAF8" author="Hubbard H. G." box="[189,658,1332,1358]" page="229" pageId="7" pageNumber="8" year="1896">
<taxonomicName id="4C1B4D33FFA9FFFCFF36FA8EFD7DFAF8" ID-CoL="9QPT" authority="Hubbard, 1896: 229" authorityName="Hubbard" authorityPageNumber="229" authorityYear="1896" box="[189,658,1332,1358]" class="Insecta" family="Staphylinidae" genus="Acrostilicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hospes">
<emphasis id="B96FEAA2FFA9FFFCFF36FA8EFE70FAF8" box="[189,415,1332,1358]" italics="true" pageId="7" pageNumber="8">Acrostilicus hospes</emphasis>
<bibRefCitation id="EF8A4B41FFA9FFFCFE2CFA8EFD7DFAF8" author="Hubbard H. G." box="[423,658,1332,1358]" pageId="7" pageNumber="8" pagination="299 - 302" refId="ref12173" refString="Hubbard H. G. 1896. Additional notes on the insect guests of the Florida land tortoise. Proceedings of the Entomological Society of Washington 3: 299 - 302." type="journal article" year="1896">Hubbard, 1896: 229</bibRefCitation>
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(original description).
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<taxonomicName id="4C1B4D33FFA9FFFCFF36FACFFE70FA39" ID-CoL="9QPT" authorityName="Hubbard" authorityYear="1896" box="[189,415,1397,1423]" class="Insecta" family="Staphylinidae" genus="Acrostilicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hospes">Acrostilicus hospes</taxonomicName>
</emphasis>
<treatmentCitation id="0ABA10A1FFA9FFFCFE30FACFFD21FA39" author="Blackwelder R. E." box="[443,718,1397,1423]" page="107" pageId="7" pageNumber="8" year="1939">
<bibRefCitation id="EF8A4B41FFA9FFFCFE30FACFFD21FA39" author="Blackwelder R. E." box="[443,718,1397,1423]" pageId="7" pageNumber="8" pagination="93 - 125" refId="ref11472" refString="Blackwelder R. E. 1939. A generic revision of the staphylinid beetles of the tribe Paederini. Proceedings of the United States National Museum 87 (3069): 93 - 125. https: // doi. org / 10.5479 / si. 00963801.87 - 3069.93" type="journal article" year="1939">Blackwelder 1939: 107</bibRefCitation>
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, 117; 1952: 37 (notes). —
<treatmentCitation id="0ABA10A1FFA9FFFCFB8CFACCFAA2FA39" author="Moore I. &amp; Legner E. F." box="[1031,1357,1397,1424]" page="101" pageId="7" pageNumber="8" year="1979">
<bibRefCitation id="EF8A4B41FFA9FFFCFB8CFACCFAA2FA39" author="Moore I. &amp; Legner E. F." box="[1031,1357,1397,1424]" pageId="7" pageNumber="8" refId="ref12946" refString="Moore I. &amp; Legner E. F. 1979. An Illustrated Guide to the Genera of the Staphylinidae of America North of Mexico exclusive of the Aleocharinae (Coleoptera). Division of Agricultural Sciences, California University. Available from http: // www. faculty. ucr. edu / ~ legneref / eflpub / 185. htm [accessed 11 Apr. 2022]." type="url" year="1979">Moore &amp; Legner 1979: 101</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ABA10A1FFA9FFFCFF61FA22FE5AFA04" author="Frania H. E." box="[234,437,1432,1458]" page="368" pageId="7" pageNumber="8" year="1986">
<bibRefCitation id="EF8A4B41FFA9FFFCFF61FA22FE5AFA04" author="Frania H. E." box="[234,437,1432,1458]" pageId="7" pageNumber="8" pagination="467 - 480" refId="ref12048" refString="Frania H. E. 1986. Status of Eustilicus Sharp, Trochoderus Sharp, Deroderus Sharp, and Stilocharis Sharp (Coleoptera: Staphylinidae: Paederinae: Paederini) with implications for classification of the Medonina and Stilicina. Canadian Journal of Zoology 64 (2): 467 - 480. https: // doi. org / 10.1139 / z 86 - 070" type="journal article" year="1986">Frania 1986: 368</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ABA10A1FFA9FFFCFE69FA23FD14FA04" author="Newton A. F. &amp; Thayer M. K. &amp; Ashe J. S. &amp; Chandler D. S." box="[482,763,1432,1459]" page="387" pageId="7" pageNumber="8" year="2001">
<bibRefCitation id="EF8A4B41FFA9FFFCFE69FA23FD14FA04" author="Newton A. F. &amp; Thayer M. K. &amp; Ashe J. S. &amp; Chandler D. S." box="[482,763,1432,1459]" pageId="7" pageNumber="8" pagination="1" refId="ref13134" refString="Newton A. F., Thayer M. K., Ashe J. S. &amp; Chandler D. S. 2001. Staphylinidae Latreille, 1802. In: Arnett Jr J. H. &amp; Thomas R. H. (eds) American Beetles. V. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press LLC, Boca Raton, FL, USA. https: // doi. org / 10.1201 / 9781482274325" type="book chapter" year="2001">
Newton
<emphasis id="B96FEAA2FFA9FFFCFDCEFA23FD6FFA04" box="[581,640,1432,1458]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2001: 387
</bibRefCitation>
</treatmentCitation>
(comment on taxonomic status). —
<treatmentCitation id="0ABA10A1FFA9FFFCFB28FA22FE8FFA60" author="Brunke A. J. &amp; Buffam J." page="154" pageId="7" pageNumber="8" year="2018">
<bibRefCitation id="EF8A4B41FFA9FFFCFB28FA22FE8FFA60" author="Brunke A. J. &amp; Buffam J." pageId="7" pageNumber="8" pagination="145 - 159" refId="ref11758" refString="Brunke A. J. &amp; Buffam J. 2018. A review of Nearctic rove beetles (Staphylinidae) specialized on the burrows and nests of Vertebrates. In: Biology of Rove Beetles (Staphylinidae). Life History, Evolution, Ecology and Distribution: 145 - 159. Springer, Cham. https: // doi. org / 10.1007 / 978 - 3 - 319 - 70257 - 5 _ 8" type="book chapter" year="2018">Brunke &amp; Buffam 2018: 154</bibRefCitation>
</treatmentCitation>
(note on biology). —
<treatmentCitation id="0ABA10A1FFA9FFFCFDEDFA01FC2BFA60" author="Brunke A. J. &amp; Schnepp K." box="[614,964,1467,1494]" page="888" pageId="7" pageNumber="8" year="2021">
<bibRefCitation id="EF8A4B41FFA9FFFCFDEDFA01FC2BFA60" author="Brunke A. J. &amp; Schnepp K." box="[614,964,1467,1494]" pageId="7" pageNumber="8" pagination="883 - 894" refId="ref11834" refString="Brunke A. J. &amp; Schnepp K. 2021. Taxonomic changes in Nearctic Paederinae, new records and a redescription of enigmatic genus Acrostilicus Hubbard (Coleoptera: Staphylinidae). Coleopterists Bulletin 75 (4): 883 - 894. https: // doi. org / 10.1649 / 0010 - 065 X- 75.4.883" type="journal article" year="2021">Brunke &amp; Schnepp 2021: 888</bibRefCitation>
</treatmentCitation>
(redescription).
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</paragraph>
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<heading id="D0EC81DCFFA9FFFCFF36FA47FED9F9A1" bold="true" box="[189,310,1533,1559]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B96FEAA2FFA9FFFCFF36FA47FED9F9A1" bold="true" box="[189,310,1533,1559]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436B0FFA9FFFCFF36F991FCD9F93A" blockId="7.[189,1399,1579,1676]" pageId="7" pageNumber="8">
The species and genus can be differentiated from all other
<taxonomicName id="4C1B4D33FFA9FFFCFC0DF991FC07F9F3" authorityName="Casey" authorityYear="1905" box="[902,1000,1579,1605]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="16" phylum="Arthropoda" rank="subTribe" subTribe="Stilicina">Stilicina</taxonomicName>
based on the combination of the following characters: the presence of a single median tooth on the labrum, mandibles symmetrical, posterior margin of head rounded, and legs elongated.
</paragraph>
</subSubSection>
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<emphasis id="B96FEAA2FFA9FFFCFF36F909FE83F97B" bold="true" box="[189,364,1715,1741]" pageId="7" pageNumber="8">Type material</emphasis>
</heading>
</paragraph>
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<materialsCitation id="3B733CEDFFA9FFFCFF61F958FD64F8F5" ID-GBIF-Occurrence="3764006330" collectionCode="NMNH" collectorName="CollHubbard &amp; Schwarz" country="United States of America" county="Cotype No" location="Clearwater" municipality="Clearwater" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B96FEAA2FFA9FFFCFF61F958FE89F94A" bold="true" box="[234,358,1762,1788]" pageId="7" pageNumber="8">
<typeStatus id="54A08812FFA9FFFCFF61F958FE89F94A" box="[234,358,1762,1788]" pageId="7" pageNumber="8" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated here)
<collectingCountry id="F30C7620FFA9FFFCFF36F8BFFF17F8A9" box="[189,248,1797,1823]" name="United States of America" pageId="7" pageNumber="8">USA</collectingCountry>
<specimenCount id="9D1DFD39FFA9FFFCFE98F8BCFEC3F896" box="[275,300,1798,1824]" pageId="7" pageNumber="8" type="male"></specimenCount>
; “
<collectingMunicipality id="6BC0ACCAFFA9FFFCFECFF8BFFE25F8A9" box="[324,458,1797,1823]" pageId="7" pageNumber="8">Clearwater</collectingMunicipality>
,
<locationDeviation id="9A162F92FFA9FFFCFE5CF8BFFDD6F8A9" box="[471,569,1797,1823]" pageId="7" pageNumber="8">27.6 Fla</locationDeviation>
/
<collectorName id="26EE5366FFA9FFFCFDDFF8BFFD02F8A9" box="[596,749,1797,1823]" pageId="7" pageNumber="8">CollHubbard</collectorName>
and
<collectorName id="26EE5366FFA9FFFCFCA1F8BFFC60F8A9" box="[810,911,1797,1823]" pageId="7" pageNumber="8">Schwarz</collectorName>
/ Gopher / Cotype No 22508 U.S.N.M /
<taxonomicName id="4C1B4D33FFA9FFFCFF36F892FDFBF8F4" authority="Hubbard" authorityName="Hubbard" authorityYear="1896" box="[189,532,1832,1858]" class="Insecta" family="Staphylinidae" genus="Acrostilicus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hospes">
<emphasis id="B96FEAA2FFA9FFFCFF36F892FE70F8F4" box="[189,415,1832,1858]" italics="true" pageId="7" pageNumber="8">Acrostilicus hospes</emphasis>
Hubbard
</taxonomicName>
”;
<collectionCode id="ED0AAE75FFA9FFFCFDA1F893FD67F8F5" box="[554,648,1833,1859]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8" type="Museum">NMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA436B0FFA9FFFCFF61F8D0FBACF87D" blockId="7.[189,1398,1898,1995]" pageId="7" pageNumber="8">
<materialsCitation id="3B733CEDFFA9FFFCFF61F8D0FBD0F87D" ID-GBIF-Occurrence="3764006302" collectionCode="NMNH" collectorName="Hubbard &amp; Schwarz" country="United States of America" county="Cotype No" location="Clearwater" municipality="Clearwater" pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<emphasis id="B96FEAA2FFA9FFFCFF61F8D0FE4DF832" bold="true" box="[234,418,1898,1924]" pageId="7" pageNumber="8">
<typeStatus id="54A08812FFA9FFFCFF61F8D0FE4DF832" box="[234,418,1898,1924]" pageId="7" pageNumber="8" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
<collectingCountry id="F30C7620FFA9FFFCFF36F837FF17F811" box="[189,248,1933,1959]" name="United States of America" pageId="7" pageNumber="8">USA</collectingCountry>
<specimenCount id="9D1DFD39FFA9FFFCFE9BF837FEBBF811" box="[272,340,1933,1959]" pageId="7" pageNumber="8" type="female">2 ♀♀</specimenCount>
; “
<collectingMunicipality id="6BC0ACCAFFA9FFFCFEE6F837FE1CF811" box="[365,499,1933,1959]" pageId="7" pageNumber="8">Clearwater</collectingMunicipality>
,
<locationDeviation id="9A162F92FFA9FFFCFE74F837FDB0F811" box="[511,607,1933,1959]" pageId="7" pageNumber="8">27.6 Fla</locationDeviation>
/ Coll
<collectorName id="26EE5366FFA9FFFCFD39F837FCF6F811" box="[690,793,1933,1959]" pageId="7" pageNumber="8">Hubbard</collectorName>
and
<collectorName id="26EE5366FFA9FFFCFCD8F837FC57F811" box="[851,952,1933,1959]" pageId="7" pageNumber="8">Schwarz</collectorName>
/ Cotype No 22508 U.S.N.M; Funiak 4.7 Fla / Coll Hubbard and Schwarz / Cotype No 22508 U.S.N.M.”;
<collectionCode id="ED0AAE75FFA9FFFCFC6AF80BFBD0F87D" box="[993,1087,1969,1995]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="8" type="Museum">NMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C301653BFFA6FFF2FF36FEB2FC16F9D4" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BA436B0FFA6FFF3FF36FEB2FE84FE94" blockId="8.[189,363,264,290]" box="[189,363,264,290]" pageId="8" pageNumber="9">
<heading id="D0EC81DCFFA6FFF3FF36FEB2FE84FE94" bold="true" box="[189,363,264,290]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B96FEAA2FFA6FFF3FF36FEB2FE84FE94" bold="true" box="[189,363,264,290]" pageId="8" pageNumber="9">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436B0FFA6FFF3FF36FE82FC51FEE4" blockId="8.[189,958,311,338]" box="[189,958,311,338]" pageId="8" pageNumber="9">
MEASUREMENTS. Medium sized (whole body length:
<quantity id="4CE39B55FFA6FFF3FC94FE8DFC5EFEE4" box="[799,945,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.7" metricValueMin="4.4" pageId="8" pageNumber="9" unit="mm" value="4.55" valueMax="4.7" valueMin="4.4">4.44.7 mm</quantity>
);
</paragraph>
<paragraph id="8BA436B0FFA6FFF3FF36FEC3FD42FE00" blockId="8.[189,1399,376,438]" pageId="8" pageNumber="9">HABITUS. Slender, elongated, with long legs. Integument shining, with coarse, umbilicate sculpture. Head and pronotum differently punctured.</paragraph>
<paragraph id="8BA436B0FFA6FFF3FF36FE67FC4AFE41" blockId="8.[189,933,477,504]" box="[189,933,477,504]" pageId="8" pageNumber="9">
COLOURATION. Pale brown/sandy with darker head tip (
<figureCitation id="13202A35FFA6FFF3FCB2FE67FC77FE41" box="[825,920,477,503]" captionStart="Fig" captionStartId="8.[189,232,1916,1942]" captionTargetBox="[311,1251,1092,1878]" captionTargetId="figure-270@8.[311,1277,1092,1878]" captionTargetPageId="8" captionText="Fig. 2. Acrostilicus hospes Hubbard, 1896, habitus photograph and drawings of the apical abdominal sternite and genital structures. A. Habitus, lectotype, ♂ (NMNH). B. Male sternite VIII. C. Aedeagus, parameral view. D. Aedeagus, lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6533022" httpUri="https://zenodo.org/record/6533022/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="8BA436B0FFA6FFF2FF36FDA5FBD5FE66" blockId="8.[187,1401,542,1019]" lastBlockId="9.[187,1399,265,465]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
HEAD. Orbicular (HL:
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; HW:
<quantity id="4CE39B55FFA6FFF3FDF4FDA4FCF2FD8F" box="[639,797,542,569]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="8" pageNumber="9" unit="mm" value="0.875" valueMax="0.9" valueMin="0.85">0.850.9 mm</quantity>
), widest in middle, wider and longer than pronotum, temples rounded and more than ½ of head length, posterior margin rounded, not emarginate, disk rugose, integument smooth and shining. Eyes large, about ¼ of head length, ½ of temple length, shifted anteriorly, slightly protruding, with setae between ommatidia. Antenna 11-segmented; antennomeres 1 and 2 with sparse pubescence; antennomeres 3 and 11 more densely pubescent, with tomentose pubescence. Antennomere 1 elongate, widest at tip; antennomeres 26 elongate; antennomere 7 weakly elongate; antennomeres 810 gradually becoming transverse; antennomere 11 elongate, 1.5 × as long as antennomere 10. Clypeal margin straight. Labrum large, transverse, twice as wide as long or wider, expanded, covering mandibles when closed, weakly sclerotised, with single short median tooth; six long, pale setae on anterior margin, evenly distributed from one edge to another, and multiple shorter setae, organised into two rows. Mandible without prostheca, with three teeth on both right and left mandible (largest one closest to base). Maxillary palpus 4-segmented; maxillary palpomere 1 short; maxillary palpomere 2 longer than wide, slightly expanded towards apex, with only few setae; maxillary palpomere 3 longer than palpomere 2 and as wide as it, slightly expanded towards apex with denser pubescence, vase-like; maxillary palpomere 4 small, acicular, glossy, and thin, equal to or shorter than width of palpomere 3. Labial palpus 3-segmented; labial palpomere 1 longer than wide; labial palpomere 2 more than twice as long as wide, wider than labial palpomere 1; labial palpomere 3 thin, much shorter and slightly narrower than labial palpomere 2. Mentum transverse, rectangular. Submentum with pair of setae on each side. Ligula entire, not bilobed, dorsal plate without setae. Gular sutures fully fused, not reaching posterior margin of head. Neck narrow, less than 1/5 of head width.
</paragraph>
<caption id="DF646638FFA6FFF3FF36F8C6FC48F86B" ID-DOI="http://doi.org/10.5281/zenodo.6533022" ID-Zenodo-Dep="6533022" httpUri="https://zenodo.org/record/6533022/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1916,1942]" targetBox="[311,1251,1092,1878]" targetPageId="8">
<paragraph id="8BA436B0FFA6FFF3FF36F8C6FC48F86B" blockId="8.[188,1399,1916,2013]" pageId="8" pageNumber="9">
<emphasis id="B96FEAA2FFA6FFF3FF36F8C6FEE2F820" bold="true" box="[189,269,1916,1942]" pageId="8" pageNumber="9">Fig. 2.</emphasis>
<taxonomicName id="4C1B4D33FFA6FFF3FE9DF8C6FD55F820" authority="Hubbard, 1896" authorityName="Hubbard" authorityYear="1896" box="[278,698,1916,1942]" class="Insecta" family="Staphylinidae" genus="Acrostilicus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hospes">
<emphasis id="B96FEAA2FFA6FFF3FE9DF8C6FE14F820" box="[278,507,1916,1942]" italics="true" pageId="8" pageNumber="9">Acrostilicus hospes</emphasis>
<bibRefCitation id="EF8A4B41FFA6FFF3FD8DF8C6FD55F820" author="Hubbard H. G." box="[518,698,1916,1942]" pageId="8" pageNumber="9" pagination="299 - 302" refId="ref12173" refString="Hubbard H. G. 1896. Additional notes on the insect guests of the Florida land tortoise. Proceedings of the Entomological Society of Washington 3: 299 - 302." type="journal article" year="1896">Hubbard, 1896</bibRefCitation>
</taxonomicName>
, habitus photograph and drawings of the apical abdominal sternite and genital structures.
<emphasis id="B96FEAA2FFA6FFF3FDA2F825FDD2F80F" bold="true" box="[553,573,1951,1977]" pageId="8" pageNumber="9">A</emphasis>
. Habitus, lectotype, ♂ (NMNH).
<emphasis id="B96FEAA2FFA6FFF3FC45F81AFC0DF80C" bold="true" box="[974,994,1952,1978]" pageId="8" pageNumber="9">B</emphasis>
. Male sternite VIII.
<emphasis id="B96FEAA2FFA6FFF3FB53F825FB03F80F" bold="true" box="[1240,1260,1951,1977]" pageId="8" pageNumber="9">C</emphasis>
. Aedeagus, parameral view.
<emphasis id="B96FEAA2FFA6FFF3FEF4F879FE7BF86B" bold="true" box="[383,404,1987,2013]" pageId="8" pageNumber="9">D</emphasis>
. Aedeagus, lateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA436B0FFA7FFF2FF36FE42FBE5FA98" blockId="9.[186,1401,503,1327]" pageId="9" pageNumber="10">
THORAX. Pronotum longer than wide (PL:
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; PW:
<quantity id="4CE39B55FFA7FFF2FCFDFE4DFBE3FDA4" box="[886,1036,503,530]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="9" pageNumber="10" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
), with anterior angles obtuse, narrowed in front, widest in middle, disc shiny, coarsely and sparsely punctured, punctures sparser than on head, evenly distributed. Basisternum of prosternum long, with longitudinal carina, prolonged behind coxa and slightly expanded laterally but not connected to hypomeron, without macrosetae or microsculpture, but surface wrinkled. Furcasternum of prosternum longer than ½ of basisternum length, triangular, acute, with sharp longitudinal carina and transversal carina. Hypomeron not delimited from pronotal disk by carina. Furcasternum of mesosternum with longitudinal carina, short, reaching ⅓ of distance between coxae, triangular. Elytra quadrate, longer and wider than pronotum (EL:
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; EW:
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), without epipleural ridge, without row of setae on edge of posterior margin and without stiff upright bristles; humeral angle indistinct, rounded; surface shiny, covered with setae not organised in distinctive rows. Scutellum without ridges, moderate, impunctate. Hind wing fully developed, MP3 vein absent, veins MP4 and CuA fused with each other. Trochantins moderate sized, oval. Coxae large, exerted. Mesocoxa contiguous, ridge below coxal rests absent. Tibiae without spines or long bristles on outer edge. Protibia with two fully developed, longitudinally placed, comb-like rows of setae, and three associated macrosetae. Tarsi 5-segmented, with one pair of empodial setae on each tarsus, equal or slightly shorter than claws (not shorter than half). Protarsus with tarsomeres 14 not inflated, narrower or equal to meso- and metatarsomeres, with dense pale adhesive setae on ventral side. Protarsomere 1 shorter than protarsomere 2, protarsomere 3 equal to protarsomere 2, but longer than protarsomere 4, protarsomere 4 not bilobed, protarsomere 5 longest, equal to protarsomeres 12 combined. Mesotarsus with mesotarsomere 1 longer than mesotarsomere 2, mesotarsomere 3 shorter than mesotarsomere 2, but longer than mesotarsomere 4, mesotarsomere 4 not bilobed, mesotarsomere 5 equal to mesotarsomere 1. Metatarsi with metatarsomere 1 twice as long as metatarsomere 2 and longer than metatarsomere 5, metatarsomeres 24 decreasing in length, metatarsomere 4 similar as metatarsomere 3 or shortest, metatarsomere 5 equal to 3, shorter than metatarsomeres 24 combined.
</paragraph>
<paragraph id="8BA436B0FFA7FFF2FF36FAECFC16F9D4" blockId="9.[188,1399,1365,1634]" pageId="9" pageNumber="10">
ABDOMEN. Finely sparsely punctate, widest at tergite V. Tergites IIIVI shallowly impressed at base, with fringe of setae on posterior margin. Tergites IIIVII with pair of paratergites on each side. Tergite VIII with posterior margin rounded. Sternite III without keel between coxae. Sternite VII with straight apical margin. Female: posterior margin of sternite VIII straight. Male: sternite VIII with moderately shallow and broad median emargination of posterior margin; sides of emargination on sternum VIII rounded (
<figureCitation id="13202A35FFA7FFF2FF4DF9B9FECCF9AB" box="[198,291,1539,1565]" captionStart="Fig" captionStartId="8.[189,232,1916,1942]" captionTargetBox="[311,1251,1092,1878]" captionTargetId="figure-270@8.[311,1277,1092,1878]" captionTargetPageId="8" captionText="Fig. 2. Acrostilicus hospes Hubbard, 1896, habitus photograph and drawings of the apical abdominal sternite and genital structures. A. Habitus, lectotype, ♂ (NMNH). B. Male sternite VIII. C. Aedeagus, parameral view. D. Aedeagus, lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6533022" httpUri="https://zenodo.org/record/6533022/files/figure.png" pageId="9" pageNumber="10">Fig. 2B</figureCitation>
). Aedeagus with parameres reduced and fused to median lobe; ventral process slightly shorter than uneverted internal sac; in parameral view with apex of ventral process rounded, sides sinuate; dorsal plate rather large and relatively weakly sclerotised (
<figureCitation id="13202A35FFA7FFF2FCE0F9F2FC02F9D4" box="[875,1005,1608,1634]" captionStart="Fig" captionStartId="8.[189,232,1916,1942]" captionTargetBox="[311,1251,1092,1878]" captionTargetId="figure-270@8.[311,1277,1092,1878]" captionTargetPageId="8" captionText="Fig. 2. Acrostilicus hospes Hubbard, 1896, habitus photograph and drawings of the apical abdominal sternite and genital structures. A. Habitus, lectotype, ♂ (NMNH). B. Male sternite VIII. C. Aedeagus, parameral view. D. Aedeagus, lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6533022" httpUri="https://zenodo.org/record/6533022/files/figure.png" pageId="9" pageNumber="10">Fig. 2CD</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C301653BFFA7FFF2FF36F933FB7BF967" box="[189,1172,1673,1745]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BA436B0FFA7FFF2FF36F933FEB9F915" blockId="9.[189,342,1673,1699]" box="[189,342,1673,1699]" pageId="9" pageNumber="10">
<heading id="D0EC81DCFFA7FFF2FF36F933FEB9F915" bold="true" box="[189,342,1673,1699]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B96FEAA2FFA7FFF2FF36F933FEB9F915" bold="true" box="[189,342,1673,1699]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA436B0FFA7FFF2FF36F90DFB7BF967" blockId="9.[189,1172,1719,1745]" box="[189,1172,1719,1745]" pageId="9" pageNumber="10">
The species has only been recorded from
<collectingRegion id="49DFF852FFA7FFF2FD2EF90DFD15F967" box="[677,762,1719,1745]" country="United States of America" name="Florida" pageId="9" pageNumber="10">Florida</collectingRegion>
(
<collectingCountry id="F30C7620FFA7FFF2FC82F90DFCA8F967" box="[777,839,1719,1745]" name="United States of America" pageId="9" pageNumber="10">USA</collectingCountry>
) (
<bibRefCitation id="EF8A4B41FFA7FFF2FCD5F90DFB68F967" author="Brunke A. J. &amp; Schnepp K." box="[862,1159,1719,1745]" pageId="9" pageNumber="10" pagination="883 - 894" refId="ref11834" refString="Brunke A. J. &amp; Schnepp K. 2021. Taxonomic changes in Nearctic Paederinae, new records and a redescription of enigmatic genus Acrostilicus Hubbard (Coleoptera: Staphylinidae). Coleopterists Bulletin 75 (4): 883 - 894. https: // doi. org / 10.1649 / 0010 - 065 X- 75.4.883" type="journal article" year="2021">Brunke &amp; Schnepp 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>