174 lines
20 KiB
XML
174 lines
20 KiB
XML
<document id="AC726797F4C4C36179A1D943D4C48EC4" ID="10.11646/zootaxa.4202.1.1" ID-DOI="10.11646/zootaxa.4202.1.1" ID-GBIF-Dataset="726c8af4-e8a8-46cb-ab0b-0ef22e432dae" ID-ISSN="1175-5326" ID-Zenodo-Dep="192220" ID-ZooBank="E89680AA-E413-4110-965E-F4084D45D4C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480664280883" checkinUser="plazi" docAuthor="Nabozhenko, Maxim & Keskin, Bekir" docDate="2016" docId="03ADFF28F84B496BFF1EF9F03DF8FE2A" docLanguage="en" docName="zootaxa.4202.1.1.pdf" docOrigin="Zootaxa 4202 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Odocnemis kakunini Nabozhenko & Keskin, 2016, sp. n." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="FF948750F8534971FF89FFC63C0AFFDF" masterDocTitle="Revision of the genus Odocnemis Allard, 1876 (Coleoptera: Tenebrionidae: Helopini) from Turkey, the Caucasus and Iran with observations on feeding habits" masterLastPageNumber="97" masterPageNumber="1" pageNumber="25" updateTime="1698395560961" updateUser="plazi">
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<mods:mods id="FE4B072C7F9817C9F40D442C622D49D7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F95CF9C563F4499722CABCDABD199E0B">
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<mods:title id="EC4A3102DD787F9F67E9313C0E7ADCA2">Revision of the genus Odocnemis Allard, 1876 (Coleoptera: Tenebrionidae: Helopini) from Turkey, the Caucasus and Iran with observations on feeding habits</mods:title>
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<mods:name id="ED95DEB9C7B3C211FE13C89B139CFCD2" type="personal">
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</mods:role>
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<mods:namePart id="5EA9A1EB455D373CE53ED35DE363E0F2">Nabozhenko, Maxim</mods:namePart>
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</mods:name>
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<mods:roleTerm id="40876387EDE3D857187A6EADE8E15E99">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4B82120C3D192EF557D8A4C5A2E8DBD5">Keskin, Bekir</mods:namePart>
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<mods:typeOfResource id="C48FB53D57114552A497A35E5D035F1C">text</mods:typeOfResource>
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<mods:title id="95E6D5377EB2EE6220DD42E40C9E7379">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0E05679B4575E253D0DE2022D7A002F4">
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<mods:date id="AE03BCE3994B6E86161A248CE1FFD8C7">2016</mods:date>
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<mods:detail id="254C2500F23FE9C0839202A2C6C563B0" type="volume">
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<mods:number id="CE872818F2459F55F3975E9293B8B4A5">4202</mods:number>
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</mods:detail>
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<mods:detail id="7608ED0FDF0CA5CEA163A3DACE587033" type="issue">
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<mods:number id="8AFD34F6C141417E861C52FE129817C2">1</mods:number>
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<mods:classification id="8E25835E4F0F33AE9C083BED800569E7">journal article</mods:classification>
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<mods:identifier id="957C8A7C59A7A9AD07879920C13340AA" type="DOI">10.11646/zootaxa.4202.1.1</mods:identifier>
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<mods:identifier id="DDE88734BFFB75163437C4D172577966" type="GBIF-Dataset">726c8af4-e8a8-46cb-ab0b-0ef22e432dae</mods:identifier>
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<mods:identifier id="5E2E661931A9A89A4CD102B0D9FD0CB2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F79B5F8C0C71236284F32C323759BA3A" type="Zenodo-Dep">192220</mods:identifier>
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<mods:identifier id="A914147F326E99AC3F7D5DC2CC36B771" type="ZooBank">E89680AA-E413-4110-965E-F4084D45D4C3</mods:identifier>
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<treatment id="03ADFF28F84B496BFF1EF9F03DF8FE2A" ID-DOI="http://doi.org/10.5281/zenodo.5681098" ID-GBIF-Taxon="124940374" ID-Zenodo-Dep="5681098" LSID="urn:lsid:plazi:treatment:03ADFF28F84B496BFF1EF9F03DF8FE2A" httpUri="http://treatment.plazi.org/id/03ADFF28F84B496BFF1EF9F03DF8FE2A" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
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<subSubSection id="C31E1DB5F84B4969FF1EF9F03D44F9AE" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BBB4E3EF84B4969FF1EF9F03DD6F98F" blockId="24.[151,476,1590,1650]" box="[151,476,1590,1616]" pageId="24" pageNumber="25">
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<heading id="D0F3F952F84B4969FF1EF9F03DD6F98F" bold="true" box="[151,476,1590,1616]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
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<emphasis id="B970922CF84B4969FF1EF9F03DD6F98F" bold="true" box="[151,476,1590,1616]" pageId="24" pageNumber="25">
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<taxonomicName id="4C0435BDF84B4969FF1EF9F03D98F98F" box="[151,402,1590,1616]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kakunini" status="sp. nov.">
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<emphasis id="B970922CF84B4969FF1EF9F03D98F98F" bold="true" box="[151,402,1590,1616]" italics="true" pageId="24" pageNumber="25">Odocnemis kakunini</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2432F57F84B4969FE13F9F03DD6F98F" box="[410,476,1590,1616]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB4E3EF84B4969FF1EF99F3D44F9AE" blockId="24.[151,476,1590,1650]" box="[151,334,1625,1650]" pageId="24" pageNumber="25">
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<figureCitation id="133F52BBF84B4969FF1EF99F3CF9F9AD" box="[151,243,1625,1650]" captionStart="FIGURE 12. O" captionStartId="25.[151,250,1961,1983]" captionTargetBox="[194,1394,811,1930]" captionTargetId="figure@25.[189,1398,805,1940]" captionTargetPageId="25" captionText="FIGURE 12. O. kakunini sp. n., male. A) Aedeagus, ventral view. B) Aedeagus, lateral view. C) Gastral spicula. D) Inner sternite VIII. E) Protibia. G) Mesotibia. H) Metatibia." httpUri="https://zenodo.org/record/192232/files/figure.png" pageId="24" pageNumber="25">Figs. 12</figureCitation>
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,
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<figureCitation id="133F52BBF84B4969FF76F99C3D10F9AD" box="[255,282,1626,1650]" captionStart="FIGURE 43. A" captionStartId="76.[151,250,1529,1551]" captionTargetBox="[156,1428,194,1505]" captionTargetId="figure@76.[150,1436,193,1508]" captionTargetPageId="76" captionText="FIGURE 43. A) O. punctata (lectotype of O. pseudoclara). B) O. punctata (Turkey—Iraq border, Çukurca). C), D) O. amanosica sp. n. E), F) O. cordiformis sp. n. G), H). O. kakunini sp. n. A), B), D), F), H) Females. C), E), G) Males." httpUri="https://zenodo.org/record/192262/files/figure.png" pageId="24" pageNumber="25">43</figureCitation>
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G, H
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C31E1DB5F84B4969FF1EF9673928F901" pageId="24" pageNumber="25" type="materials_examined">
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<paragraph id="8BBB4E3EF84B4969FF1EF9673928F901" blockId="24.[151,1437,1697,2010]" pageId="24" pageNumber="25">
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<materialsCitation id="3B6C4463F84B4969FF1EF9673914F901" ID-GBIF-Occurrence="1413110263" collectingDate="2015-05-20" collectingDateMax="2015-05-22" collectingDateMin="2015-05-20" collectionCode="ZIN" collectorName="D. G. Kasatkin & S. O. Kakunin" country="Iran" latitude="34.98795" location="Shamshir" longLatPrecision="1" longitude="46.430794" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kermanshah" typeStatus="holotype">
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<emphasis id="B970922CF84B4969FF1EF9673D4DF965" bold="true" box="[151,327,1697,1722]" pageId="24" pageNumber="25">
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<typeStatus id="54BFF09CF84B4969FF1EF9673CD8F965" box="[151,210,1697,1722]" pageId="24" pageNumber="25">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54BFF09CF84B4969FED9F9673DB2F965" box="[336,440,1697,1722]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
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(
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<specimenCount id="9D0285B7F84B4969FE43F9643DEAF965" box="[458,480,1698,1722]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
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) (
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<collectionCode id="ED15D6FBF84B4969FE73F9643E26F965" box="[506,556,1698,1722]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="24" pageNumber="25">ZIN</collectionCode>
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) and
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<typeStatus id="54BFF09CF84B4969FDE7F9643EC4F965" box="[622,718,1698,1722]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
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(
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<specimenCount id="9D0285B7F84B4969FD6BF9643EFEF965" box="[738,756,1698,1722]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
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) (private collection of Sergey Kakunin):
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<collectingCountry id="F3130EAEF84B4969FB58F964390BF966" box="[1233,1281,1698,1721]" name="Iran" pageId="24" pageNumber="25">Iran</collectingCountry>
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,
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<collectingRegion id="49C080DCF84B4969FA87F9673991F965" box="[1294,1435,1697,1722]" country="Iran" name="Kermanshah" pageId="24" pageNumber="25">Kermanshah</collectingRegion>
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Prov., near
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<location id="8EDB18E5F84B4969FE9FF9033D8AF901" LSID="urn:lsid:plazi:treatment:03ADFF28F84B496BFF1EF9F03DF8FE2A:8EDB18E5F84B4969FE9FF9033D8AF901" box="[278,384,1733,1758]" country="Iran" latitude="34.98795" longLatPrecision="1" longitude="46.430794" name="Shamshir" pageId="24" pageNumber="25" stateProvince="Kermanshah">Shamshir</location>
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vill.,
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<geoCoordinate id="EE3028F9F84B4969FE48F9033E5CF902" box="[449,598,1733,1758]" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="34.98795">34.987951 N</geoCoordinate>
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,
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<geoCoordinate id="EE3028F9F84B4969FDE8F9033EF9F902" box="[609,755,1733,1758]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="46.430794">46.430794 E</geoCoordinate>
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,
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<date id="FFBA68FEF84B4969FD76F9033F93F901" box="[767,921,1733,1758]" pageId="24" pageNumber="25" value="2015-05-20" valueMax="2015-05-22" valueMin="2015-05-20">
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<collectingDate id="EFFE9116F84B4969FD76F9033F93F901" box="[767,921,1733,1758]" pageId="24" pageNumber="25" value="2015-05-20" valueMax="2015-05-22" valueMin="2015-05-20">20–22.v.2015</collectingDate>
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</date>
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, leg.
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<collectorName id="26F12BE8F84B4969FC5CF900387CF901" box="[981,1142,1733,1758]" pageId="24" pageNumber="25">D.G. Kasatkin</collectorName>
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,
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<collectorName id="26F12BE8F84B4969FB08F9003914F901" box="[1153,1310,1733,1758]" pageId="24" pageNumber="25">S.O. Kakunin</collectorName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E1DB5F84B496BFF4EF92F3EC5FEC2" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BBB4E3EF84B4968FF4EF92F3D53FD30" blockId="24.[151,1437,1697,2010]" lastBlockId="25.[151,1437,151,752]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<emphasis id="B970922CF84B4969FF4EF92F3DABF8DD" bold="true" box="[199,417,1769,1794]" pageId="24" pageNumber="25">Description, male.</emphasis>
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Body length
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<quantity id="4CFCE3DBF84B4969FDB3F92C3E81F8DE" box="[570,651,1770,1794]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="mm" value="12.0">12 mm</quantity>
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, body width 4,3 mm.
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<emphasis id="B970922CF84B4969FC0BF92F3FB7F8DD" bold="true" box="[898,957,1769,1794]" italics="true" pageId="24" pageNumber="25">Body</emphasis>
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slender, dull, brown.
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<emphasis id="B970922CF84B4969FB3AF92F38FBF8DD" bold="true" box="[1203,1265,1769,1794]" italics="true" pageId="24" pageNumber="25">Head</emphasis>
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widest at level of the eyes. Eyes large, convex. Ratio of head width at eyes to distance between eyes: 1.8. Genae convex with strongly rounded margin, outer margin of head between gena and clypeus without emargination. Transverse depression between frons and clypeus very deep, surface of clypeus is below the surface of frons. Punctation of head irregularly coarse and dense; punctation near eyes more coarse and dense (puncture diameter 1.5–2 × as wide as distance between punctures). Antennae long (reaching 1/3 of elytral length), with 4 apical antennomeres extending beyond base of pronotum; antennomer 3 1.8 × as long as antennomere 4.
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<emphasis id="B970922CF84B4969FBDDF80438C3F806" bold="true" box="[1108,1225,1986,2009]" italics="true" pageId="24" pageNumber="25">Pronotum</emphasis>
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weakly transverse (1.14 × as wide as long), widest at middle, 1.3 × as wide as head. Lateral margins weakly regularly rounded, weakly and shortly sinuate in base. Anterior margin stright; base weakly widely rounded. Anterior angles right, rounded on apex; posterior angles weakly obtuse, distinct on apex. All margins with narrow bead, base more widely beaded. Disc weakly convex with slightly depressed middle line. Punctation of disc fine, not dense; puncture diameters 3–4 × as short as distance between punctures. Hypomera not flattened on outer margins, with fine dense winkles.
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<emphasis id="B970922CF84A4968FEF2FE8A3DC9FEBA" bold="true" box="[379,451,332,357]" italics="true" pageId="25" pageNumber="26">Elytra</emphasis>
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elongate (1.8 × as long as wide), widest behind middle, 1.27 × as wide and 2.65 × as long as pronotum, 1.7 × as wide as head. Elytral base little wider than base of pronotum. Strial punctures elongate, merged in interrupted deep furrows. Elytral intervals weakly convex, with fine and sparse punctation and distinct tubercles apically and laterally. Epipleural carinae completely visible dorsally, epipleura reaching sutural angles of elytra, not flattened (but thickened) on apex, narrow inner carinae of epipleura not reaching elytral apex. Lateral margins of elytra not sinuate or weakly sinuate on apex.
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<emphasis id="B970922CF84A4968FCBBFE393FB7FDC7" bold="true" box="[818,957,511,536]" italics="true" pageId="25" pageNumber="26">Ventral side</emphasis>
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of body without pubescence. Abdominal ventrites 1–4 with regular, moderately dense and not coarse punctation; ventrite 1 without hair brush; ventrite 5 with finer and denser punctation and shallow round depression, without pubescence, not beaded on apex.
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<emphasis id="B970922CF84A4968FAECFD8E3991FD80" bold="true" box="[1381,1435,584,607]" italics="true" pageId="25" pageNumber="26">Legs</emphasis>
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long, slender. Femora smooth and shining on inner side, not pubescent. Protibiae straight, regularly widened to apex, with regularly located along the entire length small 7 teeth; mesotibiae weakly curved and sharply widened on apex, without teeth or granules on inner side; metatibiae straight without teeth or granules. Trochanters with single long setae.
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</paragraph>
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<caption id="DF7B1EB6F84A4968FF1EF86F3EBDF801" httpUri="https://zenodo.org/record/192232/files/figure.png" pageId="25" pageNumber="26" targetBox="[194,1394,811,1930]" targetPageId="25">
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||
<paragraph id="8BBB4E3EF84A4968FF1EF86F3EBDF801" blockId="25.[151,1435,1961,2014]" pageId="25" pageNumber="26">
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<emphasis id="B970922CF84A4968FF1EF86F3D28F861" bold="true" box="[151,290,1961,1983]" pageId="25" pageNumber="26">FIGURE 12.</emphasis>
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<taxonomicName id="4C0435BDF84A4968FEA2F86C3DA9F860" box="[299,419,1961,1983]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kakunini" status="sp. nov.">
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<emphasis id="B970922CF84A4968FEA2F86C3DA9F860" box="[299,419,1961,1983]" italics="true" pageId="25" pageNumber="26">O. kakunini</emphasis>
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</taxonomicName>
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<emphasis id="B970922CF84A4968FE25F86C3DE2F860" bold="true" box="[428,488,1962,1983]" pageId="25" pageNumber="26">
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<taxonomicNameLabel id="A2432F57F84A4968FE25F86C3DE2F860" box="[428,488,1962,1983]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, male. A) Aedeagus, ventral view. B) Aedeagus, lateral view. C) Gastral spicula. D) Inner sternite VIII. E) Protibia. G) Mesotibia. H) Metatibia.
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</paragraph>
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</caption>
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<paragraph id="8BBB4E3EF849496BFF4EFF513EC5FEC2" blockId="26.[151,1436,151,501]" pageId="26" pageNumber="27">
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<emphasis id="B970922CF849496BFF4EFF513DB2FF6F" bold="true" box="[199,440,151,176]" pageId="26" pageNumber="27">Description, female.</emphasis>
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Body length
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<quantity id="4CFCE3DBF849496BFDDEFF5E3EA3FF70" box="[599,681,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="26" pageNumber="27" unit="mm" value="11.0">11 mm</quantity>
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, body width
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<quantity id="4CFCE3DBF849496BFCCBFF5E3F94FF70" box="[834,926,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="26" pageNumber="27" unit="mm" value="4.2">4.2 mm</quantity>
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. Body more robust. Pronotum widest before middle. Antennae shorter, their 3 apical antennomeres extending beyond base of pronotum, reaching 1/5 of elytral length. Elytra more convex, dorsal elytral carina not completely visible dorsally. Depression of abdominal ventrite 5 wide and occupies half of surface of the ventrite.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C31E1DB5F849496BFF4EFEE13DAAFEBA" pageId="26" pageNumber="27" type="etymology">
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<paragraph id="8BBB4E3EF849496BFF4EFEE13DAAFEBA" blockId="26.[151,1436,151,501]" pageId="26" pageNumber="27">
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<emphasis id="B970922CF849496BFF4EFEE13D44FE9F" bold="true" box="[199,334,295,320]" pageId="26" pageNumber="27">Etymology.</emphasis>
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Named in honour of Sergey Kakunin (Russian Entomological Society,
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<collectingRegion id="49C080DCF849496BFB1CFEE13904FE9F" box="[1173,1294,295,320]" country="Russia" name="Krasnodar" pageId="26" pageNumber="27">Krasnodar</collectingRegion>
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), one of the collector of the species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E1DB5F849496BFF4EFEA93DF8FE2A" pageId="26" pageNumber="27" type="diagnosis">
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<paragraph id="8BBB4E3EF849496BFF4EFEA93DF8FE2A" blockId="26.[151,1436,151,501]" pageId="26" pageNumber="27">
|
||
<emphasis id="B970922CF849496BFF4EFEA93D4BFE57" bold="true" box="[199,321,367,392]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
|
||
The species is similar to
|
||
<taxonomicName id="4C0435BDF849496BFDF5FEB63F2CFE57" box="[636,806,367,392]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov.">
|
||
<emphasis id="B970922CF849496BFDF5FEB63F2CFE57" box="[636,806,367,392]" italics="true" pageId="26" pageNumber="27">O. cordiformis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B970922CF849496BFCB8FEA93F7CFE57" bold="true" box="[817,886,367,392]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A2432F57F849496BFCB8FEA93F7CFE57" box="[817,886,367,392]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from which it differs by the brown body, not cordiform pronotum, the structure of aedeagus, absence of teeth on mesotibiae.
|
||
<taxonomicName id="4C0435BDF849496BFBBFFE5338B6FE72" box="[1078,1212,404,429]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="kakunini" status="sp. nov.">
|
||
<emphasis id="B970922CF849496BFBBFFE5338B6FE72" box="[1078,1212,404,429]" italics="true" pageId="26" pageNumber="27">O. kakunini</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B970922CF849496BFB4FFE523903FE72" bold="true" box="[1222,1289,404,429]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="A2432F57F849496BFB4FFE523903FE72" box="[1222,1289,404,429]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from other species of the
|
||
<taxonomicName id="4C0435BDF849496BFE0DFE713DFDFE0F" box="[388,503,439,464]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="praelonga">
|
||
<emphasis id="B970922CF849496BFE0DFE713DFDFE0F" box="[388,503,439,464]" italics="true" pageId="26" pageNumber="27">praelonga</emphasis>
|
||
</taxonomicName>
|
||
species-group by protibiae regularly widened to apex with small teeth regularly located along the entire length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |