663 lines
97 KiB
XML
663 lines
97 KiB
XML
<document id="ADC1C46901410C732177242CF75289CB" ID-CLB-Dataset="7166" ID-DOI="10.1093/jmammal/gyab023" ID-GBIF-Dataset="d1c8718a-3b16-45c7-867d-c8ba4e9ff482" ID-ISSN="1545-1542" ID-Zenodo-Dep="5613298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619530923730" checkinUser="felipe" docAuthor="Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C. & Heaney, Lawrence R." docDate="2021" docId="03A41512FFFE482BFEB42CFD44FA0244" docLanguage="en" docName="jMammalogy.102.2.1-22.pdf.imf" docOrigin="Journal of Mammalogy 102 (2)" docStyle="DocumentStyle:8488A0FD6434D222CC79ECAD85E66EFE.2:JourMammal.2015-.journal_article" docStyleId="8488A0FD6434D222CC79ECAD85E66EFE" docStyleName="JourMammal.2015-.journal_article" docStyleVersion="2" docTitle="Crateromys ballik Ochoa & Mijares & Piper & Reyes & Heaney 2021, new species" docType="treatment" docUuid="49891D03-4EC3-4CFB-BE8B-37385B11E3B1" docUuidSource="ZooBank" docVersion="7" lastPageNumber="15" masterDocId="FF9D6D6AFFF24825FF8C2A7447020664" masterDocTitle="Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="13" updateTime="1698937702425" updateUser="ExternalLinkService">
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<mods:title id="5374AD8C443712B5CC69ED623E331863">Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini)</mods:title>
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<mods:name id="4BD9ED5D7BECA41EF49720F0FD780FF6" type="personal">
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<mods:roleTerm id="81F1AE596DB97F060B8CEF48FB0F3492">Author</mods:roleTerm>
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<mods:namePart id="7FA1699ECFFB6B1ECB595344539A5FF8">Ochoa, Janine</mods:namePart>
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<mods:affiliation id="B334F5A68FCF6D4CFD698D075A9F03AA">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:nameIdentifier id="288B6F750368D2EF213569B199751B3C" type="email">jochoa@up.edu.ph</mods:nameIdentifier>
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<mods:namePart id="B45B0C53AD3DFBAAF3F67AB072334ECD">Mijares, Armand S. B.</mods:namePart>
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<mods:affiliation id="F07443759C048123168A61B04D226FBF">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="45109C73ECE0211680B4D905A549064B">Piper, Philip J.</mods:namePart>
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<mods:affiliation id="71DD2A3176DA66BF6DFB66B3F3292E7A">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="B40411CAD036826DED457DED6B34487B">Reyes, Marian C.</mods:namePart>
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<mods:affiliation id="A2EE02DF5A1A6A75A4069B6BD9C0F5FE">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="6C0DA3190D6C7517F4F742308CAE9429">Heaney, Lawrence R.</mods:namePart>
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<mods:affiliation id="015FC4308860201B14E5B663B51E10ED">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:title id="7DCD4BC03F67F319242A9F76CED2CD57">Journal of Mammalogy</mods:title>
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<mods:part id="EDD7A64704D4B59D013E7AD51EE852C0">
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<mods:date id="728ECBB466881187FC2C81B1D0FCFC96">2021</mods:date>
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<mods:number id="185E0D10A0C40943675F78C4D2C4D2F0">2021-04-23</mods:number>
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<mods:number id="5EC4A2CE9BCD1D10C2F82114F1AE130B">102</mods:number>
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<treatment id="03A41512FFFE482BFEB42CFD44FA0244" ID-DOI="http://doi.org/10.5281/zenodo.4723440" ID-GBIF-Taxon="181653060" ID-Zenodo-Dep="4723440" LSID="urn:lsid:zoobank.org:act:49891D03-4EC3-4CFB-BE8B-37385B11E3B1" httpUri="http://treatment.plazi.org/id/03A41512FFFE482BFEB42CFD44FA0244" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="C317F78FFFFE4829FEB42CFD458700C4" box="[312,645,1673,1696]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BB2A404FFFE4829FEB42CFD458700C4" blockId="12.[144,814,1673,1984]" box="[312,645,1673,1696]" pageId="12" pageNumber="13">
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<heading id="D0FA1368FFFE4829FEB42CFD458700C4" box="[312,645,1673,1696]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="4">
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<taxonomicName id="4C0DDF87FFFE4829FEB42CFD46FA00C4" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[312,504,1673,1696]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ballik" status="sp. nov.">
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<emphasis id="B9797816FFFE4829FEB42CFD46FA00C4" box="[312,504,1673,1696]" italics="true" pageId="12" pageNumber="13">Crateromys ballik</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24AC56DFFFE4829FD892CFD458700C4" box="[517,645,1673,1696]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C317F78FFFFE4829FF712CDD45650084" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BB2A404FFFE4829FF712CDD45C200A4" blockId="12.[144,814,1673,1984]" box="[253,704,1705,1728]" pageId="12" pageNumber="13">
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<figureCitation id="1336B881FFFE4829FF712CDD465A00A4" box="[253,344,1705,1728]" captionStart="Fig" captionStartId="9.[101,136,1865,1886]" captionTargetBox="[106,767,201,1850]" captionTargetId="figure-521@9.[105,767,200,1851]" captionTargetPageId="9" captionText="Fig. 4.—Occlusal view of maxillary molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-15997), left side; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus(FMNH193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-J3-10389), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), right side (mirrored); (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="12" pageNumber="13">Figs. 4D</figureCitation>
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,
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||
<figureCitation id="1336B881FFFE4829FEEA2CDD468400A4" box="[358,390,1705,1728]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="12" pageNumber="13">5D</figureCitation>
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,
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<figureCitation id="1336B881FFFE4829FE182CDD46A300A4" box="[404,417,1705,1728]" captionStart="Fig" captionStartId="13.[101,136,1272,1293]" captionTargetBox="[228,1362,200,1259]" captionTargetId="figure-463@13.[228,1362,200,1259]" captionTargetPageId="13" captionText="Fig. 8.—Holotype of Crateromys ballik n. sp. (PNM II-1977-I8-402), right dentary: (A) labial view of mandible (mirrored); (B) lingual view of mandible." figureDoi="http://doi.org/10.5281/zenodo.5613323" httpUri="https://zenodo.org/record/5613323/files/figure.png" pageId="12" pageNumber="13">8</figureCitation>
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,
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||
<figureCitation id="1336B881FFFE4829FE232CDD46BE00A4" box="[431,444,1705,1728]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="12" pageNumber="13">9</figureCitation>
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, and 10;
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||
<tableCitation id="C68F91BFFFFE4829FD932CDD457500A4" box="[543,631,1705,1728]" captionStart="Table 1" captionStartId="3.[125,181,1354,1375]" captionTargetPageId="3" captionText="Table 1.—Reported radiometric ages for provenance and study sites of fossil cloud rat specimens. Holotypes are underlined. *Minimum uranium series ages for Callao Layer 14. **Depth of specimen is above the last recorded occurrence of pottery; specimen is not directly associated with dated sample. 14C calibration was done using OxCal 4.3 (Bronk Ramsey 2017) and ranges shown are at 95.4% probability. Numbers in the last three columns are specimen numbers; see lists of holotypes and paratypes in text." httpUri="http://table.plazi.org/id/DF72F48CFFF14826FFF12F3E449803D3" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF14826FFF12F3E449803D3">Tables 1</tableCitation>
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and
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||
<tableCitation id="C68F91BFFFFE4829FD202CDD45BB00A4" box="[684,697,1705,1728]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF54822FFF0284C4442040F">3</tableCitation>
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;
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</paragraph>
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<paragraph id="8BB2A404FFFE4829FEDA2CBD45650084" blockId="12.[144,814,1673,1984]" box="[342,615,1737,1760]" pageId="12" pageNumber="13">Supplementary Data SD2</paragraph>
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</subSubSection>
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<subSubSection id="C317F78FFFFE4829FF272C9D437B0506" pageId="12" pageNumber="13" type="materials_examined">
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<materialsCitation id="3B65AE59FFFE4829FF272C9D43CF0766" ID-GBIF-Occurrence="3091612304" collectionCode="PNM" country="Philippines" county="Luzon Island" elevation="165" latitude="17.6645" location="Dalan Serkot Cave" longLatPrecision="14" longitude="121.82" municipality="Penablanca Municipality" pageId="12" pageNumber="13" specimenCode="PNM II-1977-I8-402" specimenCount="1" stateProvince="Cagayan Province" typeStatus="holotype">
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<paragraph id="8BB2A404FFFE4829FF272C9D45420104" blockId="12.[144,814,1673,1984]" pageId="12" pageNumber="13">
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<emphasis id="B9797816FFFE4829FF272C9D46280164" box="[171,298,1769,1792]" italics="true" pageId="12" pageNumber="13">
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<typeStatus id="54B61AA6FFFE4829FF272C9D460E0164" box="[171,268,1769,1792]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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.—
|
||
</emphasis>
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||
<specimenCode id="DBAB0C7FFFFE4829FEA62C9D450C0164" box="[298,526,1769,1792]" pageId="12" pageNumber="13">PNM II-1977-I8-402</specimenCode>
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right dentary with m1–3; incisor and all posterior processes missing (
|
||
<figureCitation id="1336B881FFFE4829FDE72D7D441C0144" box="[619,798,1801,1824]" captionStart="Fig" captionStartId="13.[101,136,1272,1293]" captionTargetBox="[228,1362,200,1259]" captionTargetId="figure-463@13.[228,1362,200,1259]" captionTargetPageId="13" captionText="Fig. 8.—Holotype of Crateromys ballik n. sp. (PNM II-1977-I8-402), right dentary: (A) labial view of mandible (mirrored); (B) lingual view of mandible." figureDoi="http://doi.org/10.5281/zenodo.5613323" httpUri="https://zenodo.org/record/5613323/files/figure.png" pageId="12" pageNumber="13">Figs. 8A and 8B</figureCitation>
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||
). The m1–3 are moderately worn (
|
||
<figureCitation id="1336B881FFFE4829FE772D5D454D0124" box="[507,591,1833,1856]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="12" pageNumber="13">Fig. 5D</figureCitation>
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||
), indicating that the animal was an adult at the time of death.
|
||
</paragraph>
|
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<paragraph id="8BB2A404FFFE4829FF272D1D43CF0766" blockId="12.[144,814,1673,1984]" lastBlockId="12.[861,1531,201,1986]" pageId="12" pageNumber="13">
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<emphasis id="B9797816FFFE4829FF272D1D465701E4" box="[171,341,1897,1920]" italics="true" pageId="12" pageNumber="13">Type locality.—</emphasis>
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<location id="8ED2F2DFFFFE4829FED92D1D452901E4" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FED92D1D452901E4" box="[341,555,1897,1920]" country="Philippines" county="Luzon Island" latitude="17.6645" longLatPrecision="14" longitude="121.82" municipality="Penablanca Municipality" name="Dalan Serkot Cave" pageId="12" pageNumber="13" stateProvince="Cagayan Province">Dalan Serkot Cave</location>
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,
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<location id="8ED2F2DFFFFE4829FDB72D1D442B01E4" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FDB72D1D442B01E4" box="[571,809,1897,1920]" country="Philippines" county="Luzon Island" latitude="17.6645" longLatPrecision="14" longitude="121.82" municipality="Penablanca Municipality" name="Barangay San Roque" pageId="12" pageNumber="13" stateProvince="Cagayan Province">Barangay San Roque</location>
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,
|
||
<collectingMunicipality id="6BD63E7EFFFE4829FF1C2DFD46A101C4" box="[144,419,1929,1952]" pageId="12" pageNumber="13">Peñablanca Municipality</collectingMunicipality>
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,
|
||
<collectingRegion id="49C96AE6FFFE4829FE392DFD458301C4" box="[437,641,1929,1952]" country="Philippines" name="Cagayan" pageId="12" pageNumber="13">Cagayan Province</collectingRegion>
|
||
,
|
||
<collectingRegion id="49C96AE6FFFE4829FD1F2DFD442B01C4" box="[659,809,1929,1952]" pageId="12" pageNumber="13">Luzon Island</collectingRegion>
|
||
,
|
||
<collectingCountry id="F31AE494FFFE4829FF1C2DDD460A01A4" box="[144,264,1961,1984]" name="Philippines" pageId="12" pageNumber="13">Philippines</collectingCountry>
|
||
(
|
||
<figureCitation id="1336B881FFFE4829FEA62DDD467901A4" box="[298,379,1961,1984]" captionStart="Fig" captionStartId="2.[144,179,2005,2026]" captionTargetBox="[228,1445,202,1991]" captionTargetId="figure-13@2.[226,1448,200,1992]" captionTargetPageId="2" captionText="Fig. 1.—Map of the Callao Cave Complex, showing location of cave sites: Callao (1), Eme (2), Dalan Serkot (3), Minori (4), and Musang (5). Inset shows the location within Luzon Island." figureDoi="http://doi.org/10.5281/zenodo.5613302" httpUri="https://zenodo.org/record/5613302/files/figure.png" pageId="12" pageNumber="13">Fig. 1</figureCitation>
|
||
),
|
||
<quantity id="4CF509E1FFFE4829FE282DDD46F801A4" box="[420,506,1961,1984]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="13" unit="m" value="165.0">
|
||
<elevation id="00204337FFFE4829FE282DDD46F801A4" box="[420,506,1961,1984]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="12" pageNumber="13" unit="m" value="165.0">165 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE39C2C3FFFE4829FD1F2DDD442B01A4" box="[659,809,1959,1984]" degrees="17" direction="north" minutes="39" orientation="latitude" pageId="12" pageNumber="13" precision="1" seconds="52.2" value="17.6645">
|
||
17°39
|
||
<emphasis id="B9797816FFFE4829FD5F2DD345D801A4" box="[723,730,1959,1984]" italics="true" pageId="12" pageNumber="13">′</emphasis>
|
||
52.2
|
||
<emphasis id="B9797816FFFE4829FC852DD3441601A4" box="[777,788,1959,1984]" italics="true" pageId="12" pageNumber="13">″</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE39C2C3FFFE4829FCD12ABF44E80686" box="[861,1002,201,226]" degrees="121" direction="east" minutes="49" orientation="longitude" pageId="12" pageNumber="13" precision="15" seconds="12" value="121.82">
|
||
121°49
|
||
<emphasis id="B9797816FFFE4829FC272ABD44B00686" box="[939,946,201,226]" italics="true" pageId="12" pageNumber="13">′</emphasis>
|
||
12
|
||
<emphasis id="B9797816FFFE4829FC402ABD44D50686" box="[972,983,201,226]" italics="true" pageId="12" pageNumber="13">″</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
, Square 2, Layer 2 (aceramic layer, spit 24, approximately
|
||
<quantity id="4CF509E1FFFE4829FC692A9F43200766" box="[997,1058,235,258]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="13" unit="m" value="1.2">
|
||
<elevation id="00204337FFFE4829FC692A9F43200766" box="[997,1058,235,258]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="13" unit="m" value="1.2">1.2 m</elevation>
|
||
</quantity>
|
||
below surface).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BB2A404FFFE4829FCF42B7F42DF0466" blockId="12.[861,1531,201,1986]" pageId="12" pageNumber="13">
|
||
<materialsCitation id="3B65AE59FFFE4829FCF42B7F438B0706" ID-GBIF-Occurrence="3091612311" collectingDate="1977-02" collectionCode="PNM" collectorName="Supplementary Data SD" county="Eme Cave" location="Callao Cave" municipality="Minori Cave" pageId="12" pageNumber="13" specimenCode="PNM II-1977-J3-10389" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B9797816FFFE4829FCF42B7F43020746" box="[888,1024,267,290]" italics="true" pageId="12" pageNumber="13">
|
||
<typeStatus id="54B61AA6FFFE4829FCF42B7F44E30746" box="[888,993,267,290]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
(
|
||
<emphasis id="B9797816FFFE4829FB852B7F43140746" box="[1033,1046,267,290]" italics="true" pageId="12" pageNumber="13">n</emphasis>
|
||
= 6; Supplementary Data SD2)
|
||
<location id="8ED2F2DFFFFE4829FA392B7F44950726" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FA392B7F44950726" county="Eme Cave" municipality="Minori Cave" name="Callao Cave" pageId="12" pageNumber="13">Callao Cave</location>
|
||
: partial maxilla with left and right M1–3 (
|
||
<specimenCode id="DBAB0C7FFFFE4829FA332B5F431F0706" pageId="12" pageNumber="13">PNM II-1977-J3-10389</specimenCode>
|
||
;
|
||
<figureCitation id="1336B881FFFE4829FBA72B3F43780706" box="[1067,1146,331,354]" captionStart="Fig" captionStartId="15.[101,136,1031,1052]" captionTargetBox="[107,1483,200,1018]" captionTargetId="figure-570@15.[107,1483,200,1018]" captionTargetPageId="15" captionText="Fig. 10.—Crateromys ballik n. sp. partial maxillary (PNM II-1977-J3-10389) with palate and all molar teeth, showing the incisive foramina (if), base of zygomatic arch (za), and palatal grooves (pg)." figureDoi="http://doi.org/10.5281/zenodo.5613329" httpUri="https://zenodo.org/record/5613329/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFE4829FB182B3F426607E6" ID-GBIF-Occurrence="3091612316" collectingDate="1977-02" collectionCode="PNM" collectorName="Supplementary Data SD" county="Eme Cave" location="Callao Cave" municipality="Minori Cave" pageId="12" pageNumber="13" specimenCode="PNM II-1977-J3-11075/2" specimenCount="1" typeStatus="paratype">
|
||
partial right dentary with incisor fragment and m1–2 (
|
||
<specimenCode id="DBAB0C7FFFFE4829FBB32B1F425507E6" box="[1087,1367,363,386]" pageId="12" pageNumber="13">PNM II-1977-J3-11075/2</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFE4829FAE02B1F42F807C6" ID-GBIF-Occurrence="3091612315" collectingDate="1977-02" collectionCode="PNM" collectorName="Supplementary Data SD" county="Eme Cave" location="Minori Cave" municipality="Minori Cave" pageId="12" pageNumber="13" specimenCode="PNM II-1977-J11A-17018" specimenCount="1" typeStatus="paratype">
|
||
<location id="8ED2F2DFFFFE4829FAE02B1F42F507E6" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FAE02B1F42F507E6" box="[1388,1527,363,386]" county="Eme Cave" municipality="Minori Cave" name="Minori Cave" pageId="12" pageNumber="13">Minori Cave</location>
|
||
: partial left dentary with m1–3 (
|
||
<specimenCode id="DBAB0C7FFFFE4829FB4A2BFF42ED07C6" box="[1222,1519,395,418]" pageId="12" pageNumber="13">PNM II-1977-J11A-17018</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFE4829FCD12BDF42F807A6" ID-GBIF-Occurrence="3091612310" box="[861,1530,427,450]" collectingDate="1977-02" collectionCode="PNM" collectorName="Supplementary Data SD" county="Eme Cave" location="Callao Cave" municipality="Minori Cave" pageId="12" pageNumber="13" specimenCode="PNM II-1977-J11A-16692" specimenCount="1" typeStatus="paratype">
|
||
partial left dentary with m1–2 (
|
||
<specimenCode id="DBAB0C7FFFFE4829FB4A2BDF42ED07A6" box="[1222,1519,427,450]" pageId="12" pageNumber="13">PNM II-1977-J11A-16692</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFE4829FCD12BBF42C00786" ID-GBIF-Occurrence="3091612306" box="[861,1474,459,482]" collectingDate="1977-02" collectionCode="PNM" collectorName="Supplementary Data SD" county="Eme Cave" location="Callao Cave" municipality="Minori Cave" pageId="12" pageNumber="13" specimenCode="PNM II-1977-J11A-896" specimenCount="1" typeStatus="paratype">
|
||
partial left dentary with m2–3 (
|
||
<specimenCode id="DBAB0C7FFFFE4829FB212BBF42B50786" box="[1197,1463,459,482]" pageId="12" pageNumber="13">PNM II-1977-J11A-896</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFE4829FA462BBF42DB0466" ID-GBIF-Occurrence="3091612318" collectingDate="1977-02" collectionCode="PNM" collectorName="Supplementary Data SD" county="Eme Cave" location="Eme Cave" municipality="Minori Cave" pageId="12" pageNumber="13" specimenCode="PNM II-1977-G-292" specimenCount="1" typeStatus="paratype">
|
||
<location id="8ED2F2DFFFFE4829FA462BBF44950466" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FA462BBF44950466" county="Eme Cave" municipality="Minori Cave" name="Eme Cave" pageId="12" pageNumber="13">Eme Cave</location>
|
||
: partial left dentary with m1–3 (
|
||
<specimenCode id="DBAB0C7FFFFE4829FB622B9F42D30466" box="[1262,1489,491,514]" pageId="12" pageNumber="13">PNM II-1977-G-292</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB2A404FFFE4829FCF4287F437B0506" blockId="12.[861,1531,201,1986]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9797816FFFE4829FCF4287F43560446" box="[888,1108,523,546]" italics="true" pageId="12" pageNumber="13">Age of specimens.—</emphasis>
|
||
The
|
||
<typeStatus id="54B61AA6FFFE4829FB0B287F43E60446" box="[1159,1252,523,546]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
comes from an aceramic layer (Layer 2) of
|
||
<location id="8ED2F2DFFFFE4829FBAC285F43E90426" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FBAC285F43E90426" box="[1056,1259,555,578]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Dalan Serkot Cave" pageId="12" pageNumber="13">Dalan Serkot Cave</location>
|
||
, dated to 7,254–6,995 cal BP (
|
||
<tableCitation id="C68F91BFFFFE4829FC00283F44DA0406" box="[908,984,587,610]" captionStart="Table 1" captionStartId="3.[125,181,1354,1375]" captionTargetPageId="3" captionText="Table 1.—Reported radiometric ages for provenance and study sites of fossil cloud rat specimens. Holotypes are underlined. *Minimum uranium series ages for Callao Layer 14. **Depth of specimen is above the last recorded occurrence of pottery; specimen is not directly associated with dated sample. 14C calibration was done using OxCal 4.3 (Bronk Ramsey 2017) and ranges shown are at 95.4% probability. Numbers in the last three columns are specimen numbers; see lists of holotypes and paratypes in text." httpUri="http://table.plazi.org/id/DF72F48CFFF14826FFF12F3E449803D3" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF14826FFF12F3E449803D3">Table 1</tableCitation>
|
||
); specimen 10389 derives from the
|
||
<location id="8ED2F2DFFFFE4829FAC1283F42F80406" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FAC1283F42F80406" box="[1357,1530,587,610]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Late Pleistocene" pageId="12" pageNumber="13">Late Pleistocene</location>
|
||
levels of
|
||
<location id="8ED2F2DFFFFE4829FC31281F433D04E6" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FC31281F433D04E6" box="[957,1087,619,642]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Callao Cave" pageId="12" pageNumber="13">Callao Cave</location>
|
||
at least ca. 66,700 BP; specimen 11075/2 comes from
|
||
<location id="8ED2F2DFFFFE4829FC6828FF432004C6" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FC6828FF432004C6" box="[996,1058,651,674]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Layer" pageId="12" pageNumber="13">Layer</location>
|
||
11 of
|
||
<location id="8ED2F2DFFFFE4829FBE528FF43AD04C6" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FBE528FF43AD04C6" box="[1129,1199,651,674]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Callao" pageId="12" pageNumber="13">Callao</location>
|
||
which is older than ca. 30,000 cal BP and less than 52,600 BP; specimen 292 derives from an aceramic layer in
|
||
<location id="8ED2F2DFFFFE4829FB9028BF43880486" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FB9028BF43880486" box="[1052,1162,715,738]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Eme Cave" pageId="12" pageNumber="13">Eme Cave</location>
|
||
dated to 3,985–3,702 cal BP; specimens 17018 and 16692 come from aceramic
|
||
<location id="8ED2F2DFFFFE4829FADD289F42B50566" LSID="urn:lsid:plazi:treatment:03A41512FFFE482BFEB42CFD44FA0244:8ED2F2DFFFFE4829FADD289F42B50566" box="[1361,1463,747,770]" county="Minori" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Holocene" pageId="12" pageNumber="13">Holocene</location>
|
||
levels of
|
||
<collectingMunicipality id="6BD63E7EFFFE4829FCF0297F43060546" box="[892,1028,779,802]" pageId="12" pageNumber="13">Minori Cave</collectingMunicipality>
|
||
(prior to the introduction of pottery on Luzon at 4,500–4,000 cal BP); specimen 896 derives from aceramicbearing level from
|
||
<collectingCounty id="62D3DC88FFFE4829FBA4293F43740506" box="[1064,1142,843,866]" pageId="12" pageNumber="13">Minori</collectingCounty>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFFE4829FCF4291F43F30586" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="8BB2A404FFFE4829FCF4291F43F30586" blockId="12.[861,1531,201,1986]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9797816FFFE4829FCF4291F430505E6" box="[888,1031,875,898]" italics="true" pageId="12" pageNumber="13">Etymology.—</emphasis>
|
||
The new species name comes from the Dupaningan Agta language of NE Luzon, an adjective meaning “small” (see
|
||
<bibRefCitation id="EF9CD9F5FFFE4829FC6A29DF435605A6" author="Reid" box="[998,1108,939,962]" firstAuthor="Reid" journalOrPublisher="Oceanic Linguistics" pageId="12" pageNumber="13" pagination="37 - 72" part="33" refId="ref18837" refString="REID, L. A. 1994. Possible non-Austronesian lexical elements in Philippine Negrito languages. Oceanic Linguistics 33: 37 - 72." title="Possible non-Austronesian lexical elements in Philippine Negrito languages" type="journal article" year="1994">Reid 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9CD9F5FFFE4829FBEE29DF420105A6" author="Robinson" box="[1122,1283,939,962]" firstAuthor="Robinson" journalOrPublisher="Canberra, Australian Capital Territory, Australia" pageId="12" pageNumber="13" refId="ref19117" refString="ROBINSON, L. C. 2011. Dupaningan Agta: grammar, vocabulary, and texts. Pacific Linguistics. Canberra, Australian Capital Territory, Australia." title="Dupaningan Agta: grammar, vocabulary, and texts. Pacific Linguistics" type="book" year="2011">Robinson 2011</bibRefCitation>
|
||
); we use it as an adjective in the nominative singular neuter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFFE4829FCF4299F43620226" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="8BB2A404FFFE4829FCF4299F43620226" blockId="12.[861,1531,201,1986]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9797816FFFE4829FCF4299F431B0266" box="[888,1049,1003,1026]" italics="true" pageId="12" pageNumber="13">Distribution.—</emphasis>
|
||
Known from four cave sites in the Callao Cave Complex, Peñablanca Municipality,
|
||
<collectingRegion id="49C96AE6FFFE4829FABF2E7F42F80246" box="[1331,1530,1035,1058]" country="Philippines" name="Cagayan" pageId="12" pageNumber="13">Cagayan Province</collectingRegion>
|
||
(
|
||
<figureCitation id="1336B881FFFE4829FCEA2E5F44A60226" box="[870,932,1067,1090]" captionStart="Fig" captionStartId="2.[144,179,2005,2026]" captionTargetBox="[228,1445,202,1991]" captionTargetId="figure-13@2.[226,1448,200,1992]" captionTargetPageId="2" captionText="Fig. 1.—Map of the Callao Cave Complex, showing location of cave sites: Callao (1), Eme (2), Dalan Serkot (3), Minori (4), and Musang (5). Inset shows the location within Luzon Island." figureDoi="http://doi.org/10.5281/zenodo.5613302" httpUri="https://zenodo.org/record/5613302/files/figure.png" pageId="12" pageNumber="13">Fig. 1</figureCitation>
|
||
), as noted above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFFE4829FCF42E3F432D00C6" pageId="12" pageNumber="13" type="diagnosis">
|
||
<paragraph id="8BB2A404FFFE4829FCF42E3F432D00C6" blockId="12.[861,1531,201,1986]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9797816FFFE4829FCF42E3F43040206" box="[888,1030,1099,1122]" italics="true" pageId="12" pageNumber="13">Diagnosis.—</emphasis>
|
||
A relatively small cloud rat of the genus
|
||
<taxonomicName id="4C0DDF87FFFE4829FCD12E1F44DF02E6" authorityName="Thomas" authorityYear="1895" box="[861,989,1131,1154]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFE4829FCD12E1F44DF02E6" box="[861,989,1131,1154]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
||
</taxonomicName>
|
||
as diagnosed and defined by Musser and Gordon (1981),
|
||
<bibRefCitation id="EF9CD9F5FFFE4829FC382EFF439202C6" author="Musser" box="[948,1168,1163,1186]" etAl="et al." firstAuthor="Musser" journalOrPublisher="American Museum Novitates" pageId="12" pageNumber="13" pagination="1 - 25" part="2821" refId="ref18546" refString="MUSSER, G. G., L. R. HEANEY, AND D. S. RABOR. 1985. Philippine rats: description of a new species of Crateromys from Dinagat Island. American Museum Novitates 2821: 1 - 25." title="Philippine rats: description of a new species of Crateromys from Dinagat Island" type="journal article" year="1985">Musser et al. (1985)</bibRefCitation>
|
||
, and Musser and Heaney (1992), distinctly smaller and less robust in comparison to the only living congener on Luzon,
|
||
<taxonomicName id="4C0DDF87FFFE4829FB192EBF42560286" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1173,1364,1227,1250]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFE4829FB192EBF42560286" box="[1173,1364,1227,1250]" italics="true" pageId="12" pageNumber="13">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C68F91BFFFFE4829FAE42EBF42BE0286" box="[1384,1468,1227,1250]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF54822FFF0284C4442040F">Table 3</tableCitation>
|
||
;
|
||
<figureCitation id="1336B881FFFE4829FA432EBF44680366" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
|
||
). Incisive foramina (if) extend nearly (about
|
||
<quantity id="4CF509E1FFFE4829FAFA2E9F42CF0366" box="[1398,1485,1259,1282]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="0.5">0.5 mm</quantity>
|
||
anterior) to the anterior margin of the M1 alveolus (
|
||
<figureCitation id="1336B881FFFE4829FA162F7F42E80346" box="[1434,1514,1291,1314]" captionStart="Fig" captionStartId="15.[101,136,1031,1052]" captionTargetBox="[107,1483,200,1018]" captionTargetId="figure-570@15.[107,1483,200,1018]" captionTargetPageId="15" captionText="Fig. 10.—Crateromys ballik n. sp. partial maxillary (PNM II-1977-J3-10389) with palate and all molar teeth, showing the incisive foramina (if), base of zygomatic arch (za), and palatal grooves (pg)." figureDoi="http://doi.org/10.5281/zenodo.5613329" httpUri="https://zenodo.org/record/5613329/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
). Palate is narrow, and palatal grooves end about parallel to t1 on M1 (
|
||
<figureCitation id="1336B881FFFE4829FC6C2F3F43310306" box="[992,1075,1355,1378]" captionStart="Fig" captionStartId="15.[101,136,1031,1052]" captionTargetBox="[107,1483,200,1018]" captionTargetId="figure-570@15.[107,1483,200,1018]" captionTargetPageId="15" captionText="Fig. 10.—Crateromys ballik n. sp. partial maxillary (PNM II-1977-J3-10389) with palate and all molar teeth, showing the incisive foramina (if), base of zygomatic arch (za), and palatal grooves (pg)." figureDoi="http://doi.org/10.5281/zenodo.5613329" httpUri="https://zenodo.org/record/5613329/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
). Dentary is moderately robust relative to molar dimensions (
|
||
<figureCitation id="1336B881FFFE4829FBD52F1F43B203E6" box="[1113,1200,1387,1410]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="12" pageNumber="13">Fig. 9A</figureCitation>
|
||
). Lower incisors proportionately narrow,
|
||
<quantity id="4CF509E1FFFE4829FC712FFF435703C6" box="[1021,1109,1419,1442]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
|
||
at midpoint. Upper toothrow length is
|
||
<quantity id="4CF509E1FFFE4829FCF72FDF44F003A6" box="[891,1010,1451,1474]" metricMagnitude="-2" metricUnit="m" metricValue="1.145" pageId="12" pageNumber="13" unit="mm" value="11.45">11.45 mm</quantity>
|
||
; lower toothrow length ranges from
|
||
<quantity id="4CF509E1FFFE4829FA102FDF44D20386" metricMagnitude="-2" metricUnit="m" metricValue="1.1565" metricValueMax="1.291" metricValueMin="1.022" pageId="12" pageNumber="13" unit="mm" value="11.565" valueMax="12.91" valueMin="10.22">10.22 to 12.91 mm</quantity>
|
||
. M1 length is
|
||
<quantity id="4CF509E1FFFE4829FBE12FBF43D00386" box="[1133,1234,1483,1506]" metricMagnitude="-3" metricUnit="m" metricValue="4.17" pageId="12" pageNumber="13" unit="mm" value="4.17">4.17 mm</quantity>
|
||
; M1 width is
|
||
<quantity id="4CF509E1FFFE4829FAE52FBF42CC0386" box="[1385,1486,1483,1506]" metricMagnitude="-3" metricUnit="m" metricValue="3.27" pageId="12" pageNumber="13" unit="mm" value="3.27">3.27 mm</quantity>
|
||
. m1 length ranges from
|
||
<quantity id="4CF509E1FFFE4829FBB12F9F43F30066" box="[1085,1265,1515,1538]" metricMagnitude="-3" metricUnit="m" metricValue="4.805" metricValueMax="5.41" metricValueMin="4.2" pageId="12" pageNumber="13" unit="mm" value="4.805" valueMax="5.41" valueMin="4.2">4.2 to 5.41 mm</quantity>
|
||
; m1 width ranges from
|
||
<quantity id="4CF509E1FFFE4829FCD12C7F43180046" box="[861,1050,1547,1570]" metricMagnitude="-3" metricUnit="m" metricValue="3.065" metricValueMax="3.34" metricValueMin="2.79" pageId="12" pageNumber="13" unit="mm" value="3.065" valueMax="3.34" valueMin="2.79">2.79 to 3.34 mm</quantity>
|
||
(
|
||
<tableCitation id="C68F91BFFFFE4829FBA12C7F43830046" box="[1069,1153,1547,1570]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF54822FFF0284C4442040F">Table 3</tableCitation>
|
||
). Labial cusps (t3 and t6) of M1 and M2 are less prominent relative to t2 and t5. M1 and M
|
||
<quantity id="4CF509E1FFFE4829FA622C5F44670006" metricMagnitude="3" metricUnit="kg" metricValue="2.0" pageId="12" pageNumber="13" unit="t" value="2.0">2 t</quantity>
|
||
6 is directly fused with t5, notches are absent between t5 and t6. M1 anteroconid is not connected to the anterolabial– anterolingual loph.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFFE482BFCF42CDF44FA0244" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="8BB2A404FFFE4828FCF42CDF45010389" blockId="12.[861,1531,201,1986]" lastBlockId="13.[101,772,1366,1997]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
<emphasis id="B9797816FFFE4829FCF42CDF43E400A6" box="[888,1254,1707,1730]" italics="true" pageId="12" pageNumber="13">Description and comparisons.—</emphasis>
|
||
Overall,
|
||
<taxonomicName id="4C0DDF87FFFE4829FAC22CDF42B900A6" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1358,1467,1707,1730]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFE4829FAC22CDF42B900A6" box="[1358,1467,1707,1730]" italics="true" pageId="12" pageNumber="13">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
is a small member of the genus
|
||
<taxonomicName id="4C0DDF87FFFE4829FB222CBF42280086" authorityName="Thomas" authorityYear="1895" box="[1198,1322,1739,1762]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFE4829FB222CBF42280086" box="[1198,1322,1739,1762]" italics="true" pageId="12" pageNumber="13">Crateromys</emphasis>
|
||
</taxonomicName>
|
||
, smaller than
|
||
<taxonomicName id="4C0DDF87FFFE4829FA572CBF44E90166" baseAuthorityName="Meyer" baseAuthorityYear="1895" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFE4829FA572CBF44E90166" italics="true" pageId="12" pageNumber="13">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFE4829FBAE2C9F439E0166" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1058,1180,1771,1794]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFE4829FBAE2C9F439E0166" box="[1058,1180,1771,1794]" italics="true" pageId="12" pageNumber="13">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
, similar in size but a bit smaller on average than
|
||
<taxonomicName id="4C0DDF87FFFE4829FB802D7F43780146" authorityName="Musser & Gordon" authorityYear="1981" box="[1036,1146,1803,1826]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFE4829FB802D7F43780146" box="[1036,1146,1803,1826]" italics="true" pageId="12" pageNumber="13">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
, and smaller than
|
||
<taxonomicName id="4C0DDF87FFFE4829FAB22D7F42A40146" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1342,1446,1803,1826]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFFE4829FAB22D7F42A40146" box="[1342,1446,1803,1826]" italics="true" pageId="12" pageNumber="13">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C68F91BFFFFE4829FA3A2D7F44680126" captionStart="Table 2" captionStartId="6.[124,180,651,672]" captionTargetPageId="6" captionText="Table 2.—Measurements (in mm) of maxillary and mandibular teeth of Carpomys and Musseromys." httpUri="http://table.plazi.org/id/DF72F48CFFF44823FFF028FF433D04C4" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF44823FFF028FF433D04C4">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="C68F91BFFFFE4829FC2F2D5F44B20126" box="[931,944,1835,1858]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="12" pageNumber="13" tableUuid="DF72F48CFFF54822FFF0284C4442040F">3</tableCitation>
|
||
;
|
||
<figureCitation id="1336B881FFFE4829FC4D2D5F43030126" box="[961,1025,1835,1858]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
|
||
). The dentary of
|
||
<taxonomicName id="4C0DDF87FFFE4829FB482D5F42280126" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1220,1322,1835,1858]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFE4829FB482D5F42280126" box="[1220,1322,1835,1858]" italics="true" pageId="12" pageNumber="13">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
is less robust relative to its molar dimensions compared to
|
||
<taxonomicName id="4C0DDF87FFFE4829FA942D3F42CF0106" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1304,1485,1867,1890]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFE4829FA942D3F42CF0106" box="[1304,1485,1867,1890]" italics="true" pageId="12" pageNumber="13">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFE4829FCD12D1F44DB01E6" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[861,985,1899,1922]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFE4829FCD12D1F44DB01E6" box="[861,985,1899,1922]" italics="true" pageId="12" pageNumber="13">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1336B881FFFE4829FC602D1F434201E6" box="[1004,1088,1899,1922]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="12" pageNumber="13">Fig. 9A</figureCitation>
|
||
), and less robust in absolute terms than
|
||
<taxonomicName id="4C0DDF87FFFE4829FCD12DFF44C901C6" authorityName="Musser & Gordon" authorityYear="1981" box="[861,971,1931,1954]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFE4829FCD12DFF44C901C6" box="[861,971,1931,1954]" italics="true" pageId="12" pageNumber="13">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
. The mental foramen of the dentary is situated at or slightly above the apex of the superior and inferior masseteric ridges on
|
||
<taxonomicName id="4C0DDF87FFFF4828FF5C2F22463A0309" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[208,312,1366,1389]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFF4828FF5C2F2247EE0309" box="[208,236,1366,1389]" italics="true" pageId="13" pageNumber="14">Cr</emphasis>
|
||
.
|
||
<emphasis id="B9797816FFFF4828FF772F22463A0309" box="[251,312,1366,1389]" italics="true" pageId="13" pageNumber="14">ballik</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1336B881FFFF4828FEC42F2246980309" box="[328,410,1366,1389]" captionStart="Fig" captionStartId="13.[101,136,1272,1293]" captionTargetBox="[228,1362,200,1259]" captionTargetId="figure-463@13.[228,1362,200,1259]" captionTargetPageId="13" captionText="Fig. 8.—Holotype of Crateromys ballik n. sp. (PNM II-1977-I8-402), right dentary: (A) labial view of mandible (mirrored); (B) lingual view of mandible." figureDoi="http://doi.org/10.5281/zenodo.5613323" httpUri="https://zenodo.org/record/5613323/files/figure.png" pageId="13" pageNumber="14">Fig. 8A</figureCitation>
|
||
), whereas it is placed high above the apex in
|
||
<taxonomicName id="4C0DDF87FFFF4828FF532F02464903E9" authorityName="Musser & Gordon" authorityYear="1981" box="[223,331,1398,1421]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFF4828FF532F02464903E9" box="[223,331,1398,1421]" italics="true" pageId="13" pageNumber="14">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
, situated on the dorsal margin of the dentary and anterior to m1, and is much lower on
|
||
<taxonomicName id="4C0DDF87FFFF4828FDC22FE2440003C9" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[590,770,1430,1453]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFF4828FDC22FE2440003C9" box="[590,770,1430,1453]" italics="true" pageId="13" pageNumber="14">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFF4828FF182FC2460C03A9" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[148,270,1462,1485]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFF4828FF182FC2460C03A9" box="[148,270,1462,1485]" italics="true" pageId="13" pageNumber="14">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
(see also Musser and Gordon 1981; Gonzales and Kennedy 1996;
|
||
<bibRefCitation id="EF9CD9F5FFFF4828FEB02FA246F10389" author="Reyes" box="[316,499,1494,1517]" etAl="et al." firstAuthor="Reyes" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="13" pageNumber="14" pagination="84 - 97" part="130" refId="ref18862" refString="REYES, M. C., T. INGICCO, P. J. PIPER, N. AmANO, AND A. F. PAWLIK. 2017. First fossil evidence of the extinct Philippine cloud rat Crateromys paulus (Muridae: Murinae: Phloeomyini) from Ilin Island, Mindoro, and insights into its Holocene abundance. Proceedings of the Biological Society of Washington 130: 84 - 97." title="First fossil evidence of the extinct Philippine cloud rat Crateromys paulus (Muridae: Murinae: Phloeomyini) from Ilin Island, Mindoro, and insights into its Holocene abundance" type="journal article" year="2017">Reyes et al. 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF72F48CFFFF4828FFE92E8C47C4034E" ID-DOI="http://doi.org/10.5281/zenodo.5613323" ID-Zenodo-Dep="5613323" httpUri="https://zenodo.org/record/5613323/files/figure.png" pageId="13" pageNumber="14" startId="13.[101,136,1272,1293]" targetBox="[228,1362,200,1259]" targetPageId="13">
|
||
<paragraph id="8BB2A404FFFF4828FFE92E8C47C4034E" blockId="13.[101,1488,1272,1322]" pageId="13" pageNumber="14">
|
||
<emphasis id="B9797816FFFF4828FFE92E8C47A70369" bold="true" box="[101,165,1272,1293]" pageId="13" pageNumber="14">Fig. 8.</emphasis>
|
||
—Holotype of
|
||
<taxonomicName id="4C0DDF87FFFF4828FEBB2E8C46E60369" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[311,484,1272,1293]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ballik" status="sp. nov.">
|
||
<emphasis id="B9797816FFFF4828FEBB2E8C46E60369" box="[311,484,1272,1293]" italics="true" pageId="13" pageNumber="14">Crateromys ballik</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFFF4828FE662E8C451D0369" box="[490,543,1272,1293]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM II-1977-I8-402), right dentary: (A) labial view of mandible (mirrored); (B) lingual view of mandible.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2A404FFFF4828FF0C2F82424400E9" blockId="13.[101,772,1366,1997]" lastBlockId="13.[819,1489,1366,1997]" pageId="13" pageNumber="14">
|
||
The lower incisor, as represented by Callao Cave
|
||
<typeStatus id="54B61AA6FFFF4828FD2B2F8244010069" box="[679,771,1526,1549]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
specimen 11072/2, is robust, orange-yellow on the anterior surface. The tip is missing, but at a place estimated to have been the midpoint, it is
|
||
<quantity id="4CF509E1FFFF4828FEEA2C2246BA0009" box="[358,440,1622,1645]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="14" unit="mm" value="1.8">1.8 mm</quantity>
|
||
wide, and from the broken tip to the anteriormost projection of bone along the medial edge is
|
||
<quantity id="4CF509E1FFFF4828FFF32CE247D400C9" box="[127,214,1686,1709]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="14" unit="mm" value="3.7">3.7 mm</quantity>
|
||
. The dorsal wear surface stops ca.
|
||
<quantity id="4CF509E1FFFF4828FDDF2CE2459000C9" box="[595,658,1686,1709]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="mm" value="1.0">1 mm</quantity>
|
||
above the beginning of the dentary bone. It appears that the tip would have extended about to the level of the occlusal surface of the mandibular molars. The lower incisor of
|
||
<taxonomicName id="4C0DDF87FFFF4828FDAC2C8245D70169" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[544,725,1782,1805]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFF4828FDAC2C8245D70169" box="[544,725,1782,1805]" italics="true" pageId="13" pageNumber="14">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
is a slightly paler version of orange-yellow on the anterior surface. At its midpoint, the incisor averages about
|
||
<quantity id="4CF509E1FFFF4828FDCE2D4245970129" box="[578,661,1846,1869]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="14" unit="mm" value="2.2">2.2 mm</quantity>
|
||
wide and about
|
||
<quantity id="4CF509E1FFFF4828FF2A2D2246300109" box="[166,306,1878,1901]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="10.7" metricValueMin="8.7" pageId="13" pageNumber="14" unit="mm" value="9.7" valueMax="10.7" valueMin="8.7">8.7–10.7 mm</quantity>
|
||
long (
|
||
<emphasis id="B9797816FFFF4828FEFA2D2246810109" box="[374,387,1878,1901]" italics="true" pageId="13" pageNumber="14">n</emphasis>
|
||
= 3) from the tip to the anteriormost projection of bone along the medial edge. The dorsal wear surface ends ca.
|
||
<quantity id="4CF509E1FFFF4828FF7B2DE2464B01C9" box="[247,329,1942,1965]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="14" unit="mm" value="2.9">2.9 mm</quantity>
|
||
above the beginning of the dentary bone along the medial edge, and the tip extends to about the level slightly below the occlusal plane of the molars. The lower incisor of
|
||
<taxonomicName id="4C0DDF87FFFF4828FC012F02430403E9" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[909,1030,1398,1421]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFF4828FC012F02430403E9" box="[909,1030,1398,1421]" italics="true" pageId="13" pageNumber="14">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
is similar in color to
|
||
<taxonomicName id="4C0DDF87FFFF4828FB7E2F0242AA03E9" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1266,1448,1398,1421]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFF4828FB7E2F0242AA03E9" box="[1266,1448,1398,1421]" italics="true" pageId="13" pageNumber="14">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
. At its midpoint, the incisor averages about
|
||
<quantity id="4CF509E1FFFF4828FB502FE2422C03C9" box="[1244,1326,1430,1453]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="14" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide, which is wider than the generally larger
|
||
<taxonomicName id="4C0DDF87FFFF4828FB062FC2424303A9" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1162,1345,1462,1485]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFF4828FB062FC2424303A9" box="[1162,1345,1462,1485]" italics="true" pageId="13" pageNumber="14">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
, but we suggest caution in interpreting this as the
|
||
<specimenCount id="9D0B6F8DFFFF4828FB402FA242680389" box="[1228,1386,1494,1517]" pageId="13" pageNumber="14" type="generic">two specimens</specimenCount>
|
||
available to us were raised in captivity.
|
||
<figureCitation id="1336B881FFFF4828FBF52F8243D60069" box="[1145,1236,1526,1549]" captionStart="Fig" captionStartId="9.[101,136,1865,1886]" captionTargetBox="[106,767,201,1850]" captionTargetId="figure-521@9.[105,767,200,1851]" captionTargetPageId="9" captionText="Fig. 4.—Occlusal view of maxillary molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-15997), left side; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus(FMNH193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-J3-10389), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), right side (mirrored); (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="13" pageNumber="14">
|
||
Figure
|
||
<quantity id="4CF509E1FFFF4828FB4B2F8243D60069" box="[1223,1236,1526,1549]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="13" pageNumber="14" unit="in" value="4.0">4</quantity>
|
||
</figureCitation>
|
||
in Musser and Gordon (1981) shows the lower incisor of
|
||
<taxonomicName id="4C0DDF87FFFF4828FB122C6242090049" authorityName="Musser & Gordon" authorityYear="1981" box="[1182,1291,1558,1581]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFF4828FB122C6242090049" box="[1182,1291,1558,1581]" italics="true" pageId="13" pageNumber="14">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
as having a dorsal wear surface that stops about
|
||
<quantity id="4CF509E1FFFF4828FB0A2C4243C50029" box="[1158,1223,1590,1613]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="2.0">2 mm</quantity>
|
||
above the anteriormost projection of bone along the medial edge, with the tip ending somewhat below the occlusal plane of the molars.
|
||
</paragraph>
|
||
<paragraph id="8BB2A404FFFF482BFCC12CE246210106" blockId="13.[819,1489,1366,1997]" lastBlockId="14.[144,814,1675,1986]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
On our single maxillary of
|
||
<taxonomicName id="4C0DDF87FFFF4828FB0B2CE243F200C9" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1159,1264,1686,1709]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFF4828FB0B2CE243F200C9" box="[1159,1264,1686,1709]" italics="true" pageId="13" pageNumber="14">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
(Callao Cave specimen 10389;
|
||
<figureCitation id="1336B881FFFF4828FC4B2CC2431500A9" box="[967,1047,1718,1741]" captionStart="Fig" captionStartId="15.[101,136,1031,1052]" captionTargetBox="[107,1483,200,1018]" captionTargetId="figure-570@15.[107,1483,200,1018]" captionTargetPageId="15" captionText="Fig. 10.—Crateromys ballik n. sp. partial maxillary (PNM II-1977-J3-10389) with palate and all molar teeth, showing the incisive foramina (if), base of zygomatic arch (za), and palatal grooves (pg)." figureDoi="http://doi.org/10.5281/zenodo.5613329" httpUri="https://zenodo.org/record/5613329/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
|
||
), the incisive foramina (if) extend posteriorly to about
|
||
<quantity id="4CF509E1FFFF4828FC792CA2434E0089" box="[1013,1100,1750,1773]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="0.5">0.5 mm</quantity>
|
||
anterior to the anterior margin of the M1 alveolus. In two
|
||
<taxonomicName id="4C0DDF87FFFF4828FBCB2C8243C60169" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1095,1220,1782,1805]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFF4828FBCB2C8243C60169" box="[1095,1220,1782,1805]" italics="true" pageId="13" pageNumber="14">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
this distance is
|
||
<quantity id="4CF509E1FFFF4828FAF72C8242CD0169" box="[1403,1487,1782,1805]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="14" unit="mm" value="1.8">1.8 mm</quantity>
|
||
and
|
||
<quantity id="4CF509E1FFFF4828FCEE2D6244B80149" box="[866,954,1814,1837]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="14" unit="mm" value="3.3">3.3 mm</quantity>
|
||
, and in six
|
||
<taxonomicName id="4C0DDF87FFFF4828FBB12D6243F50149" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1085,1271,1814,1837]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFF4828FBB12D6243F50149" box="[1085,1271,1814,1837]" italics="true" pageId="13" pageNumber="14">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
, the range is
|
||
<quantity id="4CF509E1FFFF4828FA1C2D62448E0129" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="3.3" metricValueMin="1.6" pageId="13" pageNumber="14" unit="mm" value="2.45" valueMax="3.3" valueMin="1.6">1.6 to 3.3 mm</quantity>
|
||
, averaging
|
||
<quantity id="4CF509E1FFFF4828FB822D4243650129" box="[1038,1127,1846,1869]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="14" unit="mm" value="2.7">2.7 mm</quantity>
|
||
. In
|
||
<taxonomicName id="4C0DDF87FFFF4828FB192D4243FF0129" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1173,1277,1846,1869]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFF4828FB192D4243FF0129" box="[1173,1277,1846,1869]" italics="true" pageId="13" pageNumber="14">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
10389, the palatal bridge is narrow and longitudinally scored by deep medial palatal grooves. This trait is also seen in
|
||
<taxonomicName id="4C0DDF87FFFF4828FB662D0242A001E9" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1258,1442,1910,1933]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFF4828FB662D0242A001E9" box="[1258,1442,1910,1933]" italics="true" pageId="13" pageNumber="14">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and appears to be a shared character of Luzon
|
||
<taxonomicName id="4C0DDF87FFFF4828FB762DE2427B01C9" authorityName="Thomas" authorityYear="1895" box="[1274,1401,1942,1965]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFF4828FB762DE2427B01C9" box="[1274,1401,1942,1965]" italics="true" pageId="13" pageNumber="14">Crateromys</emphasis>
|
||
</taxonomicName>
|
||
, but the grooves in
|
||
<taxonomicName id="4C0DDF87FFFF4828FC202DC2431001A9" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[940,1042,1974,1997]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFF4828FC202DC2431001A9" box="[940,1042,1974,1997]" italics="true" pageId="13" pageNumber="14">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
10389 end about parallel to t1 (
|
||
<figureCitation id="1336B881FFFF4828FAFE2DC242C201A9" box="[1394,1472,1974,1997]" captionStart="Fig" captionStartId="15.[101,136,1031,1052]" captionTargetBox="[107,1483,200,1018]" captionTargetId="figure-570@15.[107,1483,200,1018]" captionTargetPageId="15" captionText="Fig. 10.—Crateromys ballik n. sp. partial maxillary (PNM II-1977-J3-10389) with palate and all molar teeth, showing the incisive foramina (if), base of zygomatic arch (za), and palatal grooves (pg)." figureDoi="http://doi.org/10.5281/zenodo.5613329" httpUri="https://zenodo.org/record/5613329/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C0DDF87FFFC482BFE9E2CFF46CF00C6" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[274,461,1675,1698]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFC482BFE9E2CFF46CF00C6" box="[274,461,1675,1698]" italics="true" pageId="14" pageNumber="15">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
, the grooves flare laterally and become shallow anterior to M1. On other
|
||
<taxonomicName id="4C0DDF87FFFC482BFDFA2CDF47BF0086" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9797816FFFC482BFDFA2CDF45F700A6" box="[630,757,1707,1730]" italics="true" pageId="14" pageNumber="15">Crateromys</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, the palatal bridge is relatively wider and the medial palatal grooves (pg) are shallower and less conspicuous (
|
||
<bibRefCitation id="EF9CD9F5FFFC482BFD522C9F46050146" author="Musser" etAl="et al." firstAuthor="Musser" journalOrPublisher="American Museum Novitates" pageId="14" pageNumber="15" pagination="1 - 25" part="2821" refId="ref18546" refString="MUSSER, G. G., L. R. HEANEY, AND D. S. RABOR. 1985. Philippine rats: description of a new species of Crateromys from Dinagat Island. American Museum Novitates 2821: 1 - 25." title="Philippine rats: description of a new species of Crateromys from Dinagat Island" type="journal article" year="1985">Musser et al. 1985</bibRefCitation>
|
||
; Gonzales and Kennedy 1996). The anterior root of the zygomatic arch (za) appears to originate lateral to t3 on M1 (
|
||
<figureCitation id="1336B881FFFC482BFF4A2D3F46110106" box="[198,275,1867,1890]" captionStart="Fig" captionStartId="15.[101,136,1031,1052]" captionTargetBox="[107,1483,200,1018]" captionTargetId="figure-570@15.[107,1483,200,1018]" captionTargetPageId="15" captionText="Fig. 10.—Crateromys ballik n. sp. partial maxillary (PNM II-1977-J3-10389) with palate and all molar teeth, showing the incisive foramina (if), base of zygomatic arch (za), and palatal grooves (pg)." figureDoi="http://doi.org/10.5281/zenodo.5613329" httpUri="https://zenodo.org/record/5613329/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF72F48CFFFC482BFF1C2C67457C0021" ID-DOI="http://doi.org/10.5281/zenodo.5613327" ID-Zenodo-Dep="5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="14" pageNumber="15" startId="14.[144,179,1555,1576]" targetBox="[150,807,201,1541]" targetPageId="14">
|
||
<paragraph id="8BB2A404FFFC482BFF1C2C67457C0021" blockId="14.[144,813,1555,1605]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9797816FFFC482BFF1C2C6747D2004C" bold="true" box="[144,208,1555,1576]" pageId="14" pageNumber="15">Fig. 9.</emphasis>
|
||
—Bivariate plots of
|
||
<taxonomicName id="4C0DDF87FFFC482BFE1A2C674504004C" authorityName="Thomas" authorityYear="1895" box="[406,518,1555,1576]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFC482BFE1A2C674504004C" box="[406,518,1555,1576]" italics="true" pageId="14" pageNumber="15">Crateromys</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFC482BFDBB2C674588004C" authorityName="Thomas" authorityYear="1895" box="[567,650,1555,1576]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFC482BFDBB2C674588004C" box="[567,650,1555,1576]" italics="true" pageId="14" pageNumber="15">Batomys</emphasis>
|
||
</taxonomicName>
|
||
dental and mandibular measurements (in mm). Data are in Table 3.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2A404FFFC482BFF272D1F44C504E4" blockId="14.[144,814,1675,1986]" lastBlockId="14.[861,1531,201,1056]" pageId="14" pageNumber="15">
|
||
The molar morphology of
|
||
<taxonomicName id="4C0DDF87FFFC482BFE5F2D1F453801E6" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[467,570,1899,1922]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFE5F2D1F453801E6" box="[467,570,1899,1922]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0DDF87FFFC482BFD5C2D1F47DD01C6" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9797816FFFC482BFD5C2D1F442F01E6" box="[720,813,1899,1922]" italics="true" pageId="14" pageNumber="15">Batomys</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
on Luzon based on the traits listed above for the genus
|
||
<taxonomicName id="4C0DDF87FFFC482BFF1C2DDF460D01A6" authorityName="Thomas" authorityYear="1895" box="[144,271,1963,1986]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFC482BFF1C2DDF460D01A6" box="[144,271,1963,1986]" italics="true" pageId="14" pageNumber="15">Crateromys</emphasis>
|
||
</taxonomicName>
|
||
. In comparison to the fossil
|
||
<taxonomicName id="4C0DDF87FFFC482BFDD12DDF45B801A6" authorityName="Thomas" authorityYear="1895" box="[605,698,1963,1986]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFC482BFDD12DDF45B801A6" box="[605,698,1963,1986]" italics="true" pageId="14" pageNumber="15">Batomys</emphasis>
|
||
</taxonomicName>
|
||
described below,
|
||
<taxonomicName id="4C0DDF87FFFC482BFC262ABD430C0684" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[938,1038,201,224]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFC262ABD430C0684" box="[938,1038,201,224]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
is significantly larger, has higher crowns, and has a more robust dentary (
|
||
<figureCitation id="1336B881FFFC482BFB152A9D43E50764" box="[1177,1255,233,256]" captionStart="Fig" captionStartId="13.[101,136,1272,1293]" captionTargetBox="[228,1362,200,1259]" captionTargetId="figure-463@13.[228,1362,200,1259]" captionTargetPageId="13" captionText="Fig. 8.—Holotype of Crateromys ballik n. sp. (PNM II-1977-I8-402), right dentary: (A) labial view of mandible (mirrored); (B) lingual view of mandible." figureDoi="http://doi.org/10.5281/zenodo.5613323" httpUri="https://zenodo.org/record/5613323/files/figure.png" pageId="14" pageNumber="15">Figs. 8</figureCitation>
|
||
and
|
||
<figureCitation id="1336B881FFFC482BFAA82A9D42460764" box="[1316,1348,233,256]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="14" pageNumber="15">9A</figureCitation>
|
||
). Overall molar morphology follows that shown in
|
||
<figureCitation id="1336B881FFFC482BFB542B7D42880744" box="[1240,1418,265,288]" captionStart="Fig" captionStartId="4.[861,896,1502,1523]" captionTargetBox="[867,1527,202,1488]" captionTargetId="figure-650@4.[865,1527,200,1489]" captionTargetPageId="4" captionText="Fig. 2.—Maxillary and mandibular left molar toothrows of Carpomys dakal n. sp. (A and B, respectively; PNM II-1977-J11A-15997 and PNM II-1976-M-101) and Crateromys schadenbergi (C and D, respectively; FMNH 62295), showing the location of cusps and conids referred to in the text. pc = posterior cingulum." figureDoi="http://doi.org/10.5281/zenodo.5613304" httpUri="https://zenodo.org/record/5613304/files/figure.png" pageId="14" pageNumber="15">Figs. 2C and 2D</figureCitation>
|
||
, except as noted below. Measurements of individual teeth are provided in
|
||
<tableCitation id="C68F91BFFFFC482BFCF42B3D44C50704" box="[888,967,329,352]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="14" pageNumber="15" tableUuid="DF72F48CFFF54822FFF0284C4442040F">Table 3</tableCitation>
|
||
. Among Luzon cloud rats,
|
||
<taxonomicName id="4C0DDF87FFFC482BFB632B3D42510704" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1263,1363,329,352]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFB632B3D42510704" box="[1263,1363,329,352]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
is intermediate in size between
|
||
<taxonomicName id="4C0DDF87FFFC482BFB9D2B1D43C907E4" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1041,1227,361,384]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFC482BFB9D2B1D43C907E4" box="[1041,1227,361,384]" italics="true" pageId="14" pageNumber="15">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFC482BFA882B1D42B407E4" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1284,1462,361,384]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="cagayanensis" status="sp. nov.">
|
||
<emphasis id="B9797816FFFC482BFA882B1D42B407E4" box="[1284,1462,361,384]" italics="true" pageId="14" pageNumber="15">B. cagayanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFFC482BFA332B1D42F907E4" box="[1471,1531,361,384]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1336B881FFFC482BFCEA2BFD44A707C4" box="[870,933,393,416]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
|
||
). In comparison to
|
||
<taxonomicName id="4C0DDF87FFFC482BFBF62BFD423007C4" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1146,1330,393,416]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFC482BFBF62BFD423007C4" box="[1146,1330,393,416]" italics="true" pageId="14" pageNumber="15">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
, crown length and width of
|
||
<taxonomicName id="4C0DDF87FFFC482BFC312BDD432307A4" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[957,1057,425,448]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFC312BDD432307A4" box="[957,1057,425,448]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
are smaller (
|
||
<tableCitation id="C68F91BFFFFC482BFB202BDD43F807A4" box="[1196,1274,425,448]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="14" pageNumber="15" tableUuid="DF72F48CFFF54822FFF0284C4442040F">Table 3</tableCitation>
|
||
;
|
||
<figureCitation id="1336B881FFFC482BFA842BDD424407A4" box="[1288,1350,425,448]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
|
||
). Apart from absolute size of the teeth and the dentary,
|
||
<taxonomicName id="4C0DDF87FFFC482BFA802BBD42730784" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1292,1393,457,480]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFA802BBD42730784" box="[1292,1393,457,480]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0DDF87FFFC482BFCD12B9D431A0464" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[861,1048,489,512]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFC482BFCD12B9D431A0464" box="[861,1048,489,512]" italics="true" pageId="14" pageNumber="15">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
in its m1 configuration: the anteroconid is not connected to the second loph in
|
||
<taxonomicName id="4C0DDF87FFFC482BFA8C287D426B0444" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1280,1385,521,544]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFA8C287D426B0444" box="[1280,1385,521,544]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
including in specimens with heavily worn teeth (
|
||
<figureCitation id="1336B881FFFC482BFB7D285D42470424" box="[1265,1349,553,576]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="14" pageNumber="15">Fig. 5D</figureCitation>
|
||
; Supplementary Data SD2A–D), whereas this is connected in
|
||
<taxonomicName id="4C0DDF87FFFC482BFACF283D42F80404" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1347,1530,585,608]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFC482BFACF283D42F80404" box="[1347,1530,585,608]" italics="true" pageId="14" pageNumber="15">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1336B881FFFC482BFCEA281D44B504E4" box="[870,951,617,640]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="14" pageNumber="15">Fig. 5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB2A404FFFC482BFCF428FD44FA0244" blockId="14.[861,1531,201,1056]" pageId="14" pageNumber="15">
|
||
In comparison to
|
||
<taxonomicName id="4C0DDF87FFFC482BFBB828FD43AF04C4" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1076,1197,649,672]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFC482BFBB828FD43AF04C4" box="[1076,1197,649,672]" italics="true" pageId="14" pageNumber="15">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
, crown length and width of
|
||
<taxonomicName id="4C0DDF87FFFC482BFA5728FD449804A4" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFA5728FD449804A4" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
also are smaller (
|
||
<figureCitation id="1336B881FFFC482BFBEE28DD43A104A4" box="[1122,1187,681,704]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
|
||
;
|
||
<tableCitation id="C68F91BFFFFC482BFB3828DD420704A4" box="[1204,1285,681,704]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="14" pageNumber="15" tableUuid="DF72F48CFFF54822FFF0284C4442040F">Table 3</tableCitation>
|
||
). Instead, tooth measurements of
|
||
<taxonomicName id="4C0DDF87FFFC482BFC6F28BD43450484" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[995,1095,713,736]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFC6F28BD43450484" box="[995,1095,713,736]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
overlap with the range of measurements published for fossil
|
||
<taxonomicName id="4C0DDF87FFFC482BFBB9289D43A60564" authorityName="Musser & Gordon" authorityYear="1981" box="[1077,1188,745,768]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFC482BFBB9289D43A60564" box="[1077,1188,745,768]" italics="true" pageId="14" pageNumber="15">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1336B881FFFC482BFB39289D43F60564" box="[1205,1268,745,768]" captionStart="Fig" captionStartId="14.[144,179,1555,1576]" captionTargetBox="[150,807,201,1541]" captionTargetId="figure-687@14.[147,809,200,1542]" captionTargetPageId="14" captionText="Fig. 9.—Bivariate plots of Crateromys and Batomys dental and mandibular measurements (in mm). Data are in Table 3." figureDoi="http://doi.org/10.5281/zenodo.5613327" httpUri="https://zenodo.org/record/5613327/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF9CD9F5FFFC482BFA88289D42BC0564" author="Reyes" box="[1284,1470,745,768]" etAl="et al." firstAuthor="Reyes" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="14" pageNumber="15" pagination="84 - 97" part="130" refId="ref18862" refString="REYES, M. C., T. INGICCO, P. J. PIPER, N. AmANO, AND A. F. PAWLIK. 2017. First fossil evidence of the extinct Philippine cloud rat Crateromys paulus (Muridae: Murinae: Phloeomyini) from Ilin Island, Mindoro, and insights into its Holocene abundance. Proceedings of the Biological Society of Washington 130: 84 - 97." title="First fossil evidence of the extinct Philippine cloud rat Crateromys paulus (Muridae: Murinae: Phloeomyini) from Ilin Island, Mindoro, and insights into its Holocene abundance" type="journal article" year="2017">Reyes et al. 2017</bibRefCitation>
|
||
). M1 and M2 of
|
||
<taxonomicName id="4C0DDF87FFFC482BFC52297D434A0544" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[990,1096,777,800]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFC52297D44F80544" box="[990,1018,777,800]" italics="true" pageId="14" pageNumber="15">Cr</emphasis>
|
||
.
|
||
<emphasis id="B9797816FFFC482BFB87297D434A0544" box="[1035,1096,777,800]" italics="true" pageId="14" pageNumber="15">ballik</emphasis>
|
||
</taxonomicName>
|
||
are differentiated from
|
||
<taxonomicName id="4C0DDF87FFFC482BFAD8297D42C70544" authorityName="Musser & Gordon" authorityYear="1981" box="[1364,1477,777,800]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFC482BFAD8297D42C70544" box="[1364,1477,777,800]" italics="true" pageId="14" pageNumber="15">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
(see figure
|
||
<quantity id="4CF509E1FFFC482BFC2E295D44C90524" box="[930,971,809,832]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="14" pageNumber="15" unit="in" value="6.0">6 in</quantity>
|
||
Musser and Gordon 1981) and
|
||
<taxonomicName id="4C0DDF87FFFC482BFA91295D42970524" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1309,1429,809,832]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFC482BFA91295D42970524" box="[1309,1429,809,832]" italics="true" pageId="14" pageNumber="15">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1336B881FFFC482BFA29295D42F00524" box="[1445,1522,809,832]" captionStart="Fig" captionStartId="9.[101,136,1865,1886]" captionTargetBox="[106,767,201,1850]" captionTargetId="figure-521@9.[105,767,200,1851]" captionTargetPageId="9" captionText="Fig. 4.—Occlusal view of maxillary molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-15997), left side; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus(FMNH193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-J3-10389), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), right side (mirrored); (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="14" pageNumber="15">Fig. 4F</figureCitation>
|
||
) by less prominent labial cusps (t3 and t6). The t
|
||
<quantity id="4CF509E1FFFC482BFADD293D427B0504" box="[1361,1401,841,864]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="14" pageNumber="15" unit="in" value="6.0">6 in</quantity>
|
||
M1 and M2 of
|
||
<taxonomicName id="4C0DDF87FFFC482BFCF7291D44E805E4" authorityName="Musser & Gordon" authorityYear="1981" box="[891,1002,873,896]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFC482BFCF7291D44E805E4" box="[891,1002,873,896]" italics="true" pageId="14" pageNumber="15">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFC482BFBAC291D439B05E4" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1056,1177,873,896]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFC482BFBAC291D439B05E4" box="[1056,1177,873,896]" italics="true" pageId="14" pageNumber="15">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
is separated by a relatively deep notch from t5, and this notch is absent in
|
||
<taxonomicName id="4C0DDF87FFFC482BFA9829FD427505C4" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1300,1399,905,928]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFA9829FD427505C4" box="[1300,1399,905,928]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
. On the m3, the hypoconid and entoconid loph is chevronate on
|
||
<taxonomicName id="4C0DDF87FFFC482BFA1929DD42F905A4" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1429,1531,937,960]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ballik">
|
||
<emphasis id="B9797816FFFC482BFA1929DD42F905A4" box="[1429,1531,937,960]" italics="true" pageId="14" pageNumber="15">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFFC482BFC1C29BD43480584" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[912,1098,969,992]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFC482BFC1C29BD43480584" box="[912,1098,969,992]" italics="true" pageId="14" pageNumber="15">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
(i.e., both of the Luzon
|
||
<taxonomicName id="4C0DDF87FFFC482BFAE329BD42E90584" authorityName="Thomas" authorityYear="1895" box="[1391,1515,969,992]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFFC482BFAE329BD42E90584" box="[1391,1515,969,992]" italics="true" pageId="14" pageNumber="15">Crateromys</emphasis>
|
||
</taxonomicName>
|
||
), whereas it is fused in a nearly transverse loph in
|
||
<taxonomicName id="4C0DDF87FFFC482BFAED299D42CC0264" authorityName="Musser & Gordon" authorityYear="1981" box="[1377,1486,1001,1024]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="paulus">
|
||
<emphasis id="B9797816FFFC482BFAED299D42CC0264" box="[1377,1486,1001,1024]" italics="true" pageId="14" pageNumber="15">Cr. paulus</emphasis>
|
||
</taxonomicName>
|
||
and in
|
||
<taxonomicName id="4C0DDF87FFFC482BFCF52E7D44F30244" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[889,1009,1033,1056]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFC482BFCF52E7D44F30244" box="[889,1009,1033,1056]" italics="true" pageId="14" pageNumber="15">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |